Key to the Chinese Species of Systolederus Bolivar

Modified from Zheng (2013)

1. Outer side and inner side of hind tibia without spine............................. Systolederus aspinus Deng, sp. nov.

- Outer side and inner side of hind tibia with several spines...................................................... 2

2. Pronotum short, not reaching, reaching or just surpassing the apex of hind femora.................................. 3

- Pronotum elongate, extending far beyond the apex of hind femora...............................................4

3. Anterior margin of vertex straight in dorsal view; hind wings reaching apex of hind femora................................................................................................... Systolederus nigritibia Zheng, 2005

- Anterior margin of vertex acute in dorsal view; hind wings reaching middle of hind tibia............................................................................................. Systolederus brachynotus Zheng & Ou, 2010

4. Upper margin of pronotum undulating or undulating before shoulders in profile.................................... 5

- Upper margin of pronotum straight in profile................................................................ 6

5. With a pair of abbreviated carinae between shoulders; upper margin of pronotum undulating before shoulders in profile; width of midfemur 2.3 times width of tegmina.................................... Systolederus guangxiensis Zheng, 1998

- Without abbreviated carinae between shoulders; upper margin of pronotum undulating in profile; width of midfemur equal to width of tegmina......................................................... Systolederus emeiensis Zheng, 1998

6. Antenna inserted between lower margins of eyes............................................................. 7

- Antenna inserted below lower margins of eyes.............................................................. 10

7. Head distinctly exerted above upper level of pronotum; posterior process of pronotum reaching apex of hind tibia............................................................................. Systolederus oculatus Zheng & Ou, 2012

- Head slightly exerted above upper level of pronotum; posterior process of pronotum reaching one third to middle of hind tibia.................................................................................................... 8

8. Length of first segment of hind tarsi longer than third, first and second pulvillus small, third pulvillus large......................................................................................... ‘ Systolederus choui Liang, 2012

- First and third segment of hind tarsi equal in length; three pulvilli equal in length...................................9

9. Posterior process of pronotum reaching one third of hind tibia; humeral angle nearly rounded; width of midfemur wider than width of tegmina........................................................ Systolederus fujianensis Zheng, 1993

- Posterior process of pronotum reaching behind middle of hind tibia; humeral angle obtuse; width of midfemur equal to width of tegmina............................................................ .. Systolederus longinota Zheng, 2005a

10. Humeral angle nearly rounded; apex of tegmina acute; width of midfemur equal to width of tegmina.................................................................................. Systolederus spicupennis Zheng & Jiang, 2003

- Humeral angle obtuse; apex of tegmina narrowly rounded or rounded; width of midfemur wider than or narrower than width of tegmina............................................................................................ 11

11. Posterior process of pronotum reaching apex of hind tibia............. Systolederus longipennis Zheng, Jiang &Liu, 2005

- Posterior process of pronotum reaching middle to two-thirds of hind tibia........................................12

12. Width of midfemur equal to width of tegmina; posterior process of pronotum reaching two- thirds of hind tibia; outer part lower side of hind femur yellow brown............................. Systolederus heishidingensi s Zheng & Xie, 2004

- Width of midfemur distinctly wider than width of tegmina; posterior process of pronotum reaching middle of hind tibia; outer part lower side of hind femur black...................................................................... 13

13. Apex of tegmina broadly rounded; lower margins of midfemora undulated.......... Systolederus orthonotus Zheng, 1998

- Apex of tegmina narrowly rounded; lower margins of midfemora straight........................................ 14

14. Lateral ocelli placed on middle of anterior margins of eyes; with a pair of abbreviated carinae between shoulders; width of midfemur equal to width of tegmina in male................................. Systolederus hunanensis Zheng, 2013

- Lateral ocelli placed on below middle of anterior margins of eyes; without abbreviated carinae between shoulders; width of midfemur wider than width of tegmina in male............................................................. 15

15. Width of longitudinal furrow of frontal ridge equal to width of first antennal segment; hind wings reaching apex of posterior process of pronotum; width of midfemur equal to width of tegmina in female.................................................................................................... Systolederus guizhouensis Deng & Zheng, 2013

- Width of longitudinal furrow of frontal ridge narrower than width of first antennal segment; hind wings surpassing apex of posterior process of pronotum; width of midfemur wider than width of tegmina in female..................................................................................... Systolederus guposhanensis Deng, Zheng & Wei, 2007