Dactylorhiza fuchsii (Druce) Soo

Dactylorhiza hebridensis (= Dactylorhiza hebridensis (Wilmott) Aver., ≡ Dactylorhiza fuchsii subsp. hebridensis (Wilmott) Soó).

Distribution in adjacent reg.

Russia (European Russia, Ural, Siberia), Kazakhstan (Altai).

Specimens examined and literature records.

Kokchetav: Akmola Region: Burabay District: Near Karas’e Lake, 4 Jul 1937, Shishkina s.n. (AA!); same loc., Zolotoborskiy forestry, 21 Jun 1972, Gorchakovskiy s.n.. (SVER 695767!); Borovskoy forest area, upper reaches of Imanayevsky Creek, 27 Jun 1974, Gorchakovskiy s.n.. (SVER 695769!); same loc., 16 Jun 1972 Gorchakovskiy s.n.. (SVER 695769!); Barmashinskiy experimental forestry, 1 Aug 1965, Tyulebergeneva s.n. (SVER 695763!); Zolotoborsky forestry, eastern shore of Shchuchye Lake, 3 km north of the Medvezhiy cordon, 17 Jun 1972, Gorchakovskiy s.n.. (SVER 695756!); Burabay forest area, near Akylbay cordon, 10 Jun 1913, Drobova 308 (LE!); Burabay forest area, near Dorofeyevka, 10 Jun 1913, Drobova 309 (LE!); “Burabay” State National Nature Park: Borovskoe forestry, 16 Jun 2019, Kubentayev and Alibekov s.n.. (NUR!); Barmashinskoe forestry, 7 Aug 2020, Kubentayev et al. s.n.. (NUR!); Temnoborskoe forestry, near Zhukey Lake, north-west coast, 11 Jun 2019, Kubentayev et al. s.n.. (NUR!); near Burabay, the headwaters of Imanayevsky Creek, 7 Jun 2011, Kupriyanov s.n.. (KUZ 00816!); near Makinka, 11 Jun 2011, Kupriyanov s.n.. (KUZ 01111!); Zerendi District: “Kokshetau” State National Nature Park: Zerendi forestry, near Zerenda, 28 May 2020, Kubentayev and Alibekov s.n.. (NUR!); Oramndybulakskoe forestry, near Karsak, 27 Jun 2020, Kubentayev and Alibekov s.n.. (NUR!); Zerendi forestry, near Krasniy cordon, 26 Jun 2019, Kubentayev et al. s.n.. (NUR!); Sandyktauskiy District: Sandyktau forestry, near "Chernichniy log", 16 Jun 1957, Gribanov s.n.. (AA!). North Kazakhstan Region: Aiyrtau District: Imantau Mount, 7 Aug 1965, Oleneva and Antoshenko s.n.. (695765 SEVR!); same loc., the valley of the stream flowing down from Imantau Mount, 30 May 1973, Gorchakovskiy s.n.. (SEVR 695760!); Imantau Mount, Bayan Gorge, 28 Aug 1981, Gorchakovskiy s.n.. (SEVR 627680!); “Kokshetau” State National Nature Park, near Syrymbet, 10 Aug 2020, Kubentayev et al. s.n.. (NUR!); near Lobanovo, 27 Jul 2019, Kubentayev et al. s.n.. (NUR!). Tobol-Ishim: Kostanay Region: Mendykara District: botanical nature monument "Planting of birch and pine forests near Borovskoye Lake", Jun 2009, Perezhogin (personal observation); Denisovskiy District: Ordzhonikidzevskoye forestry, near Denisovka (Pugachev 1994); Naurzum District: in the “Naurzum” State Nature Reserve, s.d., KSPI students (SVER 507474!). Karkaraly: Karaganda Region: Karkaraly Mountains, 27 Jun 1843, Schrenk s.n.. (AA!); same loc., 21 Jun 1890, Korzhinsky s.n.. (LE!); same loc., 1927, Melville s.n.. ( АА !), near Karkaraly, 3 Jul 1937, Dmitrieva s.n.. (AA!); same loc., same loc., 17 Jul 1987, Kupriyanov s.n.. (KG!); same loc., 18 Jun 2001, Ishmuratova s.n (KG!); the shore of Pashennoe Lake, 20 Jun 1914, s. collector 2071 (LE); 70 km southeast of Karkaraly, Kent Mountains, 19 Jul 1968, Rachkova 784 (LE!); Karkaraly Mountains, Alexandrov Klyuch, 8 Jul 1993, Mikhailov s.n.. (KG!); Karkaraly Mountains, Karkaralinka River valley, 5 Aug 1989, Denisova 692 (MW 0816794!). Irtysh: Pavlodar Region [without detailed locality] ( Kusnetsov and Pavlov 1958) .

Habitat and ecology.

Moist pine and birch forests, along the shores of forest streams and lakes, forest swamps.

Phenology.

Flowering in Jun-Jul; fruiting in Aug-Sep.

Conservation status.

It is included in the Red Book of Kazakhstan (category II). It is protected on the territories of the “Naurzum” State Nature Reserve; “Kokshetau” State National Nature Park, “Burabay” State National Nature Park, Karkaraly State National Nature Park; Botanical nature monument "Planting of birch and pine forests near Borovskoye Lake".

Notes.

During the revision of KUZ herbarium materials, we identified two specimens of D. fuchsii, that were previously incorrectly determined as Dactylorhiza russowii . Based on these incorrectly misidentified herbarium specimens, D. russowii was previously reported for the Kokchetav FR (near Burabay, the headwaters of Imanayevsky Creek; near Makinka) and in the overall flora of Kazakhstan (Kupriyanov 2020). As a result, there is currently no reliable information regarding the presence of D. russowii in Kazakhstan.