Tribe Holoneurini Enderlein, 1936

The tribe is characterized by synapomorphies characteristic of the valid tribe rank in Porricondylinae (Mamaev, 2001; Fedotova, Sidorenko, 2007): antennae with 10–11 flagellomeres; reduction of wing venation; absent or reduced parameres; tarsal claws narrow and curved near midlength; male and female flagellomeres with dense sensorial reticulation, and female flagellomeres with a long neck. Their wing venation is variable, with M 3 absent or slightly visible, Cu never forked, CuA 2 slightly visible distally or completely disappearing, m+rm usually slightly curved distally, and rs situated at an angle to R 5. The ovipositor is 2-segmented.

The larvae of some extant species of the Holoneurini have been associated with leaf litter and decaying wood (Mamaev, Krivosheina, 1965). Adult insects are usually captured in Malaise traps (Fedotova, Sidorenko, 2005, 2007).

The tribe includes 12 genera and 43 species: 9 extant genera with 38 species (number of species in parentheses): Cassidoides Mamaev, 1960 (8); Coccopsis Meijere, 1901 (8); Cognitepidosis Fedotova et Sidorenko, 2005 (1); Cryptoneurus Mamaev, 1964 (3); Dalaiella Mamaev, 1996 (1); Holoneurus Kieffer, 1894 (14), Melicepidosis Fedotova et Sidorenko, 2007 (1); Putepidosis Fedotova et Sidorenko, 2007 (1), Pyxicoccopsis Fedotova et Sidorenko, 2007 (1) and 3 extinct genera with 5 species: Gratomyia (1), Volnococcopsis (1) and Rovnoholoneurus gen. n. (3).