2. Cloeon smaeleni Lestage, 1924

Cloeon smaeleni: Salles et al. 2014: figs 1-17; Gattolliat and Rabeantoandro 2002: figs 1-10.

Diagnoses.

Nymph. Following combination of characters: A) labrum on dorsal surface with stout, simple setae in anterior part, not forming an arc; B) both mandibles with long, stout setae between prostheca and mola; C) labial palp with enlarged, falcate segment III; C) maxillary palp with three segments; D) claw with long pointed elongation; with two rows of denticles; E) gills formed by two lamellae, upper lamella rounded and almost as developed as the lower one.

Distribution

(Fig. 5). Comoros (Grande Comore, Anjouan), Mayotte, La Réunion (Gattolliat 2004), Madagascar (Gattolliat and Rabeantoandro 2002), continental Afrotropics, Brazil (introduced, Salles et al. 2014 and citations therein).

Biological aspects.

In the Comoros, the specimens were collected in a crater lake on the island of Anjouan at an altitude of 900 m and in a cistern with collected rainwater on Grande Comore (altitude 100 m). In Mayotte, the specimens were collected at altitudes from sea level to 100 m.

Material examined.

Comoros • 66 nymphs; Anjouan, Lac Sacré, crater lake, slope of N'Tingiu Mtns ; 900 m; 12.03.1974; leg. F. Starmühlner; 2 on slides; GBIFCH00592748, GBIFCH00592749; 64 in alcohol; GBIFCH00515661, GBIFCH00515662, GBIFCH00515663; MZL, FAMU • 17 nymphs; Grande Comore, in village N of Moroni; 28.02.1974; leg. F. Starmühlner; in alcohol; GBIFCH00515664; MZL, FAMU . Mayotte • 1 nymph; Riv. Dembeni, Loc. Dembéni aval; 12°50'31"S, 45°10'22"E; 10 m; 17.05.2006; leg. N. Mary; on slide; GBIFCH00592751; MZL • 1 nymph; Bas. Ourovéni, Riv. Haoutoungou, Lac Karihani; 12°47'49"S, 45°07'20"E; 19.05.2006; leg. N. Mary; on slide; GBIFCH00592752; MZL • 3 nymphs; Bas. Riv. Djalimou, Djalimou aval; 12°57'14"S, 45°06'51"E; 15 m; 14.04.2009; leg. N. Mary; in alcohol; GBIFCH00515672; FAMU • 1 nymph; Bas. M’Tsangachéhi, Loc. M'Tsangachehi aval; 12°53'04"S, 45°07'55"E; 10 m; 14.04.2009; leg. N. Mary; in alcohol; GBIFCH00515676; MZL • 1 nymph; Bas. Riv. Bouyouni, Loc. Bouyouni intermediaire; 12°44'55"S, 45°08'36"E; 40 m; 16.04.2009; leg. N. Mary; in alcohol; GBIFCH00515678; MZL • 2 nymphs; Riv. Dembéni, Loc. Dembeniintermediaire; 12°51'07"S, 45°09'41"E; 95 m; 20.04.2009; leg. N. Mary; in alcohol; GBIFCH00515717; MZL • 1 nymph; Riv. Dapani aval, P1; 12°58'05"S, 45°07'39"E; 25 m; 19.08.2018; leg. N. Mary; in alcohol; MZL .