Achlidon puntarenas (Hobbs, 1991)

(Figs. 6, 7, 77)

Pseudothelphusa puntarenas Hobbs, 1991: 295, fig. 1.— Rodríguez 1992: 184.— Hobbs 1994: 101.— Suárez 2005: 156.— Villalobos Hiriart & Álvarez 2008: 294 (in list).— Villalobos & Alvarez 2010: 472. Pseudothelphusa punctarenas [sic]— Ng et al. 2008: 176 (in list).

Material examined. COSTA RICA. Puntarenas Province, Pacific drainage: 1 male, holotype, USNM 250555, small stream in Emus Cave, 8.i.1989, F. Grady leg.; 2 males, paratypes, USNM 250556, same data; 7 females, paratypes, USNM 250557, 1.i.1989, F. Grady leg.; 1 male, 3 juv. males, 1 female, 5 juv. females, UCR-MZ 3215- 0 1, village Río Claro, cascade Las Cavernitas, 8°41'48.48" N 83°02'34.34"W, 106 m, 27.i.2014, I.S. Wehrtmann, C. Magalhães & E. Souza-Carvalho leg.

Description of gonopod. Nearly straight, only slightly curved towards latero-cephalic side on caudo-mesial view, narrower at end of proximal third. Marginal suture on mesial surface, not twisted to other surfaces. Distal portion of caudo-marginal projection slightly bilobate in mesial view. Marginal process (distal lobe of caudomarginal projection, sensu Villalobos & Alvarez 2010) triangular, produced beyond the mesial crest of apex, directed cephalodistally; proximal lobe broadly rounded, border smooth. Mesial process strongly developed, reniform. Distocaudal ridge elongated, extending distally from the beginning of second third of stem length. Apex elongated, narrow; both mesial, lateral crests nearly straight. Apex cavity situated distally, with spermatic channel opening positioned on cephalic end; field of apical spines weakly developed situated on middle, cephalic end.

Holotype and type locality. Male, cw 18.0, cl 11.5 (chelipeds dearticulated), USNM 250555. Costa Rica, Puntarenas Province, small stream tributary of Claro River, in Emus Cave, near Río Claro village [08°40’N 83° 04’W] (Pacific drainage).

Distribution. Southern Costa Rica, Pacific drainage (Fig. 77).

Remarks. The G1 morphology of the present taxon clearly differs from that of the species of the genus Pseudothelphusa de Saussure, 1857. Villalobos Hiriart & Álvarez (2008) excluded P. puntarenas from the Pseudothelphusini Ortmann, 1897 and proposed that it should be placed in Achlidon . Such recommendation was based on the results of their phylogenetic analysis of the tribe Pseudothelphusini, which was subsequently published (Villalobos & Alvarez 2010) when they confirmed the new combination. Villalobos & Alvarez (2010) used somatic and sex characters to argue that four autapomorphies seen in the G1 of P. puntarenas supported such exclusion and that the apical position of the spermatic channel opening and the lack of torsion of the G1 would “indicate that the species belongs to the Hypolobocerini in the genus Achlidon ”.