Helicopsyche woldai sp.n.

(Figs 1­8)

Material examined: Male holotype: Panama, Province of Panama, Barro Colorado Island, Snyder­Molino Trail, marker 3, 27.v­1.ix.1987, H. Wolda, light trap (NMNH).

Paratypes: Same data as holotype, except: 1 male: 2.ii.1977 (UCD). 1 male: 25.x.1977 (UCD). 2 males: 22.xi.1977 (NRM). 1 male: 6.xii.1977 (UCD). Same date: 2 males (NRM). 1 male: 20.xii.1977 (UCD). 2 males: 6.ix.1978 (UCD). 1 male: 8.xi.1978 (NRM). 7 males: 27.v­1.ix.1987 (NMNH). 15 males: 2.ix­29.xii.1987 (NMNH). 1 male: 14­ 20.ix.1988 (NMNH). 11 males: 12.x.1988 ­ 3.i.1989 (NRM). 2 males: 23.viii­12.ix.1989 (NMNH). 3 males: 1.xi­19.xii.1989 (NMNH). 1 male: 6­12.vi.1990 (NMNH). 7 males: 25.vii­2.x.1990 (NMNH). 4 males: 14.xi­11.xii.1990 (NMNH).

Etymology: woldai, named after Henk Wolda, the collector of all material in this work. The name is to be treated as a noun in genitive case.

Diagnosis. H. woldai sp.n. can be separated from other Helicopsychidae in lateral view by the slender, parallelogram­shaped gonocoxite that has a strongly produced, fingerlike posterior corner.

Description

Head: Antenna scape as long as eye diameter. Maxillary palps with distal joint slightly shorter than proximal joint. Proximal joint about as long as scape. Cephalic warts large, rounded, convex. Postantennal warts large, slightly club­shaped.

Thorax: Pronotum with small lateral setal warts tangent with median warts. Mesonotal setal warts small, oblong, about 2x longer than wide.

Legs: Fore leg anterior spur about 2x longer than the posterior spur. Thoracic setal warts, and leg spurs as in H. fridae sp.n.

Wings (Fig. 1): Fore wing 2.6­3.3 (N=36 paratypes, mean=3.0), 3.0 mm (holotype); venation as in H. fridae, except Dc one third the wing length; crossvein R—M as long as the basal part of R5; M1 about 1.2 x longer than stalk M1+2; Crossvein M—Cu1 meets Cu1 at a distance before bifurcation of Cu1 equal to half the length of Cu1a. Hind wing 2.0­2.4 (N=36 paratypes, mean= 2.2 mm), 2.4 mm (holotype), with 21 hamuli; venation as in H. fridae sp.n., except M divides slightly after bifurcation of Rs; crossvein Cu1—M forms short stump on Cu1a.

VIth sternal process (Figs 2, 3) oriented posteroventrad, sub straight; apically pointed, about 0.55 x the segment length, reaches segment VII (Fig. 2); in ventral view (Fig. 3), slightly narrowing mid way and widened toward apex; sub apical lamellae absent; numerous spines cover ventral face of apex (Fig. 3).

Genitalia (Figs 4­8): Segment IX, lateral view (Fig. 4), anterior lobe located dorsally on segment, produced anteriorly, anterodorsal margin slightly concave; anteroventral margin shallowly concave; in dorsal view (Fig. 5), with inner margin widely hyperboloid; in ventral view (Fig. 6) with bi­lobed central process on posterior margin; lateral apodeme (Fig. 4) curving along its length, meets anterior margin below anterior lobe apex; tergal transverse apodeme absent; long sternal transverse apodeme weakly developed along posterior margin. Segment X, lateral view (Fig. 4), tapering along its length, slightly curving ventrally, apex truncate; ventral margin slightly concave; dorsal margin gently convex; basal, dorsal incision absent; in dorsal view (Fig. 5), narrowing along its length, apical notch small, V­shaped; with about eleven long megasetae in dorsolateral group starting at midway. Superior appendage (Fig. 4), strongly club­shaped, oriented posterolaterad. Gonocoxite, lateral view (Fig. 4), generally rectangular; dorsal margin undulate; apex unusually long and slender, curved mesad along its length; at midway slightly broader than height of central part of Xth tergum (Fig. 4); anterodorsal margin smooth; posteroventral margin sub straight; widely incised toward basimesal lobe; with well developed row of processes at posteroventral margin. Basimesal lobe, lateral view, well developed, produced posteriorly, slightly triangular, apex pointed, with about three stout megasetae at median margin (Fig. 6); in ventral view (Fig. 4), narrowly cone­shaped, with smooth margins; median margin slightly undulate, without marginal setae. Basal plate long (Fig. 4), with irregular dorsal and sub straight ventral margin; in ventral view (Fig. 6), triangular, apex pointed. Phallus (Figs 7, 8): very thick, at central part strongly bent ventrad; posteroventral part well sclerotized; basal one fourth slightly thicker than midlength; phallic basis present; endotheca strongly produced into irregular dorsal lobe (Fig. 7), and single posterior lobe (Fig. 8); sperm channel uniformly wide inside phallus, sharply narrowing and bent ventrally at anterior opening.

Affinities: Only H. grenadensis has long Xth tergal setae, a cone­shaped basimesal lobe, and a serial of processes on the posteroventral margin of the gonocoxite, but H. woldai sp.n. has a more rectangular gonocoxite and a longer basimesal lobe. H. minima Siebold (Puerto Rico) also has a rectangular gonocoxite, but lacks the basimesal lobe and long Xth tergal setae.