Lepidophtalmus siriboia density

The mean burrow density, including both areas, was 31.7 ± 34.1 burrows.m −2 and there were significant differences between areas (F (1, 238) = 25.66; p <0.001) and between intertidal zones (F(1, 238) = 114.85; p <0.001), with interaction between factors (F(1, 238) = 7.76; p <0.001). In general, higher densities were recorded at Area 1 in LT (86.17 ± 41.3 burrows.m −2). Except for the HT zone (where no burrows were found in both areas), the intertidal zones of Area 1 have higher burrow densities than those of Area 1 (Figure 3). However, the two areas have similar patterns of distribution of burrow-density increases towards the sea (Figure 3).

CS: coarse sand; MS: medium sand; FS: fine sand; VFS: very fine sand; M.Sorted: moderately sorted.

The best-fitting regression model to explain the variation spatial of L. siriboia included the grain size and proportions of finer (silt + clay) (Table 2). A negative relationship was observed between the grain size and density of L. siriboia burrows. By contrast, the density of shrimp burrows increased with an increasing percentage of finer sediment (silt and clay).