3. Genus Dishkeya Stonis, 2020

(Figs 183–194)

Dishkeya Stonis, 2020: 125–127 .

Type species: Dishkeya gouaniae (Stonis & Diškus, 2007: 1286–1291) .

Diagnosis. In the male genitalia, species of Dishkeya are characterized by the modified socii usually forming a pseudognathos, a wide phallus with lateral processes and variously developed carinae, and absence of a juxta or anellus. In the female genitalia, the genus is characterized by the small ovipositor lobes and large lateral lobes. In contrast to majority of Tischeriidae, leaf mines of Dishkeya are linear on Rhamnaceae host plants. For major diagnostic characters of Dishkeya and their occurrence in other genera of Tischeriidae see Tabs 2, 3.

Molecular notes. According to the molecular analysis, Dishkeya is a separate clade, and always branches with Coptotrichoides; together these two genera are often close to the Coptotriche genus (Figs 62, 64).

Adult. Head: frontal tuft overlapping the frons, comprised of long, slender lamellar and piliform scales; collar comprised of slender lamellar scales. Forewing without a distinct pattern, yellowish ochre to pale grey, irregularly speckled with some darker scales (Stonis & Solis 2020). Hindwing slender, without androconia.

Male genitalia (Figs 183–194). Uncus with two long, very slender or wide lateral lobes. Socii spinose, modified, usually thickened medially and forming a pseudognathos (Figs 183–187). Tegumen long, without lateral frame-like thickening; diaphragm without spines. Valva slender to medium wide, usually with inner process or larger inner lobe; basal process of valva very short, except for D. ursipedella Diškus, Mey & Stonis (Stonis et al. 2022a) which is characterized by very long basal processess. Transtilla or anellus absent. Vinculum long, widely rounded or truncated distally. Phallus wide to very wide, occasionally constricted medially, with strongly developed lateral processes (sometimes also possessing spines) and variously developed, usually asymmetrical apical carinae (Figs 191–194); the recently described Colombian D. ursipedella (Stonis et al. 2022a) posseses particularly developed, five-claw carinae (Figs 190, 191).

Female genitalia. Ovipositor lobes small; the gap between ovipositor lobes relatively wide; second pair of ovipositor lobes almost equal in size to the main ovipositor lobes or divided. Lateral lobes very large, wide proximally. Anterior apophyses slightly shorter than posterior apophyses. Prela with three pairs of rod-like projections; caudal sclerite strongly developed, inverted V-shaped, with a large pointed lobe caudally.Antrum absent. Otherwise, female genitalia are unknown.

Bionomics. The genus is trophically associated with Rhamnaceae and known leaf mines are linear. However, there remains a need to discover and study the biology of additional species in this genus.

Species diversity and geographical distribution. Currently the genus is comprised of four species. It is predicted that this genus might have an almost continuous distribution along the West coast of the Americas (Stonis & Solis 2020); recently one new species was described from the Colombian Andes (Stonis et al. 2022a).