Subfamily Epigambriinae Handlirsch, 1939, stat. nov.

Epigambridae [sic] Handlirsch, 1913: 813 (nomen nudum); Zittel, 1921: 679 (nomen nudum); Zittel, 1924: 700 (nomen nudum); Labandeira, 2002: 232 (fig. 4).

Epigambriidae Handlirsch, 1939: 76; Bode 1953: 18; Brues et al., 1954: 782, 817; Carpenter, 1992: 355; Makarkin & Archibald, 2003: 176; Engel & Grimaldi, 2008: 8 (Table 3); Jepson et al., 2012: 40; Yang et al., 2012: 2.

Type genus. Epigambria Handlirsch, 1939 .

Diagnosis. May be distinguished from members of the polystoechotid genus-group of Ithonidae s.l. by a combination of the following character states: (1) a single gradate series between branches of RP in both fore- and hind wings [two series in genera of the polystoechotid genus-group (rarely: outer series distinct, proximal crossveins numerous, not arranged in inner series in forewing)]; (2) absence of crossveins between branches of MP and CuA in forewing [present in genera of the polystoechotid genus-group]; (3) several crossveins between RA and RP nearly perpendicular to these veins in hind wing [few and oblique in most members of the polystoechotid genus-group].

Included genera. Epigambria, Polyosmylus, and Rasnitsyneura, gen. nov., all from the Early Jurassic (early Toarcian) of Germany.

Remarks. Handlirsch (1913) and Zittel (1921, 1924) mentioned the name ‘Epigambridae’, but no genera were included to this family until 1939.