Brachycoleus genus

(Fig. 2, node 18).

The genus comprises ten species known from the Palearctic region (Hosseini & Mohammadi 2019, Aukema 2020) with four species reported from Iran. Brachycoleus is found to be monophyletic in both analyses with moderate support (IW= 62%, EW= 66%). This clade is supported by a unique synapomorphy (20-0) and four other morphological characters including: swollen frons (7-1), left paramere apophysis, subapically expanded (58-1), median process of posterior wall present/small (86-1) and dorsal margin of second valvula edentate or ventral margin moderately dentate (89-0). In the present study, the monophyly of Brachycoleus is studied for the first time.