Emesopsis fenestrella n. sp.

Figs. 6, 7

Material examined. HOLOTYPE. AUSTRALIA: Queensland: 1 ɗ, Mt Hayward, 20.3°S 148.7°E, 350m, 20 Nov 1992, Monteith, Thompson & Janetzki, Pyrethrum. PARATYPES. AUSTRALIA: New South Wales: 1 Ψ, 1 ɗ, Gibraltar Range Nat. Park, 29.5o S 152.3o E, 30 Mar 1980 G.B. Monteith, Pyrethrum ex. Xanthorrhoea; Northern Territory: 1 ɗ, West Alligator Mouth, 12.2o S 132.3o E, 21 July 1979 (WA2), G. B. Monteith; Queensland: ɗ, Bulburin S.F., 9km E. Many Peaks, 24.5o S 151.4o E, 159 m, 600m, 17 Sept 1989, G.B. Monteith, Pyrethrum, Rainforest; 1 ɗ, Wallaman Falls, via Ingham, 18.6o S 145.8o E, 500m, 1 Oct 1980 G. Monteith, Pyrethrum, Rainforest; Ψ, Bulburn barracks, 24.5°S 151.5°E, 580m, 8 Oct 1999, G.B. Monteith, rainforest, Pyrethrum, trees. 7816; 1 Ψ, Lake Barrine, Atherton Tableland, 17.2o S 145.7o E, 736 m, 15 December 1961, D. McAlpine & R. Lossin.

Diagnosis. Recognized by the following combination of characters: M and Cu not fused basal of discal cell, small subquadrate subbasal cell present in at least one wing, R+M apical of discal cell weakly arched and faintly branched; eyes large; postocular region of head with tumid genal area.

Description. Body length 4.3 mm

Colour. As shown in Figure 7. Body mostly testaceous with brown markings. Lobes of head testaceous; AI pale stramineous with two narrow subapical brown annuli, A2-A4 testaceous with pale stramineous apical annulus; labium pale stramineous with testaceous markings, basal 2/3 of LI testaceous, LII basally testaceous becoming pale stramineous to apex, LIII pale stramineous basally becoming testaceous. Pronotum excluding appendages entirely testaceous. Forecoxae mostly pale stramineous with narrow testaceous basal annulus. Meso- and metacoxae testaceous. Forefemora with three brown annulations, one large basal, one small medial, and one large subapical, with medial and basal annulations converging on exterior surface. Meso- and metafemora mostly stramineous with three narrow annuli, one subapical, one medial, and one equidistant in between. Foretibia pale stramineous with testacous apical and vague smoky elongate subbasal annulation. Meso- and metatibia with brown subbasal narrow annulus, area preceding annulus pale stramineous, remainder of leg stramineous. Foretarsus mostly pale stramineous, pretarsus testaceous. Meso- and metatarsi stramineous. Forewings as in Figure 7. Abdomen mostly stramineous to testaceous.

Texture and Vestiture. Head dorsally covered in dense white wool-like pile interspersed with long upright setae; AI with short setae; labium with scattered short setae. Thorax covered dorsally and laterally with white woollike pile interspersed with long setae, lateral tumid areas of anterior lobe of pronotum with trident-shaped area bereft of setae. Coxae and forefemora with long setae. Foretibia covered in shorter spine-like setae. Meso and metafemora with long setae basally, remainder with short spine-like setae. Meso- and metatibia with spine-like setae, basally becoming more sparse. Abdomen blanketed with short, velutinous silken setae.

Structure. As in Figure 7. Postocular region slightly larger than anteocular region, latter elevated with depressed V-shaped area dorsally. Postocular region subglobose. Interocular groove deep, arching only slightly posteriorly. Labium strongly bent between LI and LII, LII strongly swollen, relative lengths 1:0.45:0.75. Pronotum constricted before midpoint; anterior lobe divided bilaterally by a deep pit in the disc, subglobose in dorsal view; posterior lobe almost two times longer than anterior lobe, disc with vague median longitudinal depression; posterior margin sinuate. Scutellum semicircular with posteriorly produced tubercle. Metanotum U-shaped with large spine. Forecoxae elongate, cylindrical. Forefemora elongate, subclylindrical, posteroventral and anteroventral series similar in structure. Foretarsus 2-segmented, second segment 4 times larger than first. Meso- and metacoxae globose. Meso- and metafemora thin, elongate, cylindrical. Meso- and metatibia thin, elongate, cylindrical. Meso- and metatarsi 3-segmented, segments approximately equal in length.

Distribution. E. fenestrella is known from the eastern coast of Australia, from the New South Wales – Queensland border to Cape York (Fig. 6).

Etymology. The specific epithet fenestrella (Latin for “small window”) refers to the exceptionally small size of the basal cell of the forewing.