Emesopsis Uhler 1893
Diagnosis. Recognized in Australia by the following combination of characters: head, thorax and S1 covered in dense wool-like pile; darkened areas of wings pitted with unpigmented dots, M and Cu separate at base of truncate discal cell; 2nd segment of rostrum distinctly swollen
Remarks. Emesopsis is probably the most morphologically diverse genus in the tribe. The genus has previously been diagnosed by the presence of a small subquadrate basal cell, however, this character is not present in one Australian species, Emesopsis infenestra, nor in an undescribed species from Lord Howe Island. Though these species lack a basal cell, they are otherwise clearly members of the genus. Furthermore, the basal cell has been found to be present in one wing and absent in the other within a specimen of E. fenestrella . The loss of the basal cell could result from the migration of the Cu and discal cell towards one another. This transformation of a large cell to small, and then obsolete subquadrate cell can be visualized by examining the wings of E. fenestrella, E. bunda and E. infenestra respectively.