Onthophagus bidentatus Drapiez, 1819

Fig. 59A

Onthophagus bidentatus Drapiez, 1819: 134 (original description). Type locality: French Guiana. Name-bearing type: lectotype (MNHN), designated by Rossini et al. (2016), examined by FZVM.

Onthophagus bicornis Castelnau, 1840: 87 (original description). Type locality: French Guiana. Name-bearing type: lectotype (MNHN), designated by Rossini et al. (2016), examined by FZVM.

Onthophagus femoralis Kirsch, 1871: 362 (original description). Type locality: Colombia: Cundinamarca: Bogotá. Name-bearing type: unknown to us.

Onthophagus semichalcites d’Orbigny, 1902: 149 (original description). Type locality: unknown; the lectotype is mislabelled as though coming from Benin (Rossini et al. 2016). Name-bearing type: lectotype (MNHN), designated by Rossini et al. (2016), examined by FZVM.

Onthophagus bidentatus – Harold 1869d: 1026 (catalogue); 1880: 33 (redescription, distribution). — Gillet 1911b: 204 (catalogue). — Bruch 1911: 190 (cited for Argentina). — Gillet & Boucomont 1927: 204 (catalogue of species). — Boucomont 1932: 304, 321 (key, distribution). — Balthasar 1941: 352 (cited for Peru); 1951: 337 (cited for Peru). — Blackwelder 1944: 211 (list of species from Latin America). — Roze 1955: 45 (cited for Venezuela). — Vulcano & Pereira 1967: 564 (key). — Zunino & Halffter 1997: 161 (list). — Medina et al. 2001: 139 (checklist). — Pulido-Herrera & Zunino 2007: 97 (catalogue). — Ratcliffe et al. 2015: 195 (checklist for Peru). — Rossini et al. 2016: 496, 497 (comments, fig. 1a); 2018b: 9 (list of species of the hircus complex).

Onthophagus bicornis – Harold 1869d: 1026 (cited as synonym of Onthophagus bidentatus); 1880: 33 (cited as synonym of O. bidentatus). — Gillet 1911b: 204 (cited as synonym of O. bidentatus). — Gillet & Boucomont 1927: 204 (cited as synonym of O. bidentatus). — Blackwelder 1944: 211 (cited as synonym of O. bidentatus). — Pulido-Herrera & Zunino 2007: 97 (cited as synonym of O. bidentatus). — Rossini et al. 2016: 496 (cited as synonym of O. bidentatus).

Onthophagus femoralis – Harold 1880: 33 (cited as synonym of O. bidentatus). — Gillet 1911b: 204 (cited as synonym of O. bidentatus). — Gillet & Boucomont 1927: 204 (cited as synonym of O. bidentatus). — Blackwelder 1944: 211 (cited as synonym of O. bidentatus). — Pulido-Herrera & Zunino 2007: 97 (cited as synonym of O. bidentatus).

Onthophagus semichalcites – Gillet 1911b: 195 (cited for Nigeria). — Gillet & Boucomont 1927: 195 (cited for Nigeria). — Rossini et al. 2016: 496 (comments).

Onthophagus (Onthophagus) bidentatus – Vaz-de-Mello 2000: 194 (cited for Brazil). — Chamorro et al. 2018: 96 (cited for Ecuador). — Hielkema & Hielkema 2019: 90 (catalogue for the Guianas).

Material examined

VENEZUELA ‒ Aragua • 3 specs; Maracay, Choroni; 28 Jul. 1993; O. Hillert leg.; CEMT . ‒ Bolívar • 3 specs; Río Caura, Maripa; 20 Mar. 1995; Homburg leg.; CEMT • 2 specs; Anacoco; curso NM2006 leg.; CEMT . – Anzoátegui • 1 spec. n; Anaco; 8 Aug. 1992; Homburg leg.; CEMT . ‒ Táchira • 1 spec.; Libertador, San Joaquín de Navay; 7°47′59.99″ N, 71°40′24.24″ W; 655 m a.s.l.; T. Good et al. leg.; human faeces; CEMT .

Distribution

Panama, Colombia, Venezuela, Trinidad and Tobago, Guyana, Suriname, Brazil, Ecuador, Peru, and possibly Argentina (Rossini et al. 2016) .

Subregions of Venezuela

Coastal mainland, Plains, System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní of Northeast Guyana Shield, and Central Coast Mountain Range.

Literature records

Boucomont 1932: 321 (Venezuela: Distrito Capital: Caracas). — Balthasar 1941: 352 (Venezuela); 1951: 337 (Venezuela). — Roze 1955: 45 (Venezuela: Mérida, doubtful data). — Martínez 1959: 109 (Venezuela). — Blanco 1988: 46–47 (Venezuela: Táchira, doubtful data). — Havranek 1989: 61 (Venezuela: Táchira, doubtful data). — Pulido-Herrera & Zunino 2007: 97 (Venezuela).— Rossini et al. 2016: 496 (Venezuela). — Chamorro et al. 2019: 182 (Venezuela).