Laeocathaica polytyla Moellendorff, 1899

Figs 2A, 3, 24A, B, 26, 43A, B, 45K, L, 51

Laeocathaica polytyla Möllendorff, 1899: 98, pl. 6, fig. 7; - Wiegmann 1900: 118, pl. 3, figs 105-107; - Gude 1902: 6; - Yen 1939: 150, pl. 15, fig. 41; - Schileyko 2004: 1686, fig. 2174B-D; - Chen and Zhang 2004: 331, fig. 322; - Páll-Gergely et al. 2022: 68, fig. 20A-C.

Laeocathaica (Laeocathaica) polytyla - Zilch 1968: 174; - Richardson 1983: 78.

Museum material.

SMF 9098, lectotype; Nanping, Sung-pan. SMF 9099, paratypes; six fms; Nanping, Sung-pan. Other SMF material see in Yen 1939 (150).

New material.

HBUMM00506, 13 fma, 1 dissected; Anchanghexiang [安昌河乡], Wenxian, Gansu Province, 1200 m, near point (33.066003°N, 104.460018°E); 1998-V-19, coll. Chen, D.-N. and Zhang, G.-Q. HBUMM00513, a subadult; Shuanghexiang [双河乡], Jiuzhaigouxian, Sichuan Province, 1100 m, near (33.179715°N, 104.264968°E); 1998-V-18, coll. Chen, D.-N. and Zhang, G.-Q. HBUMM00579, many fms; Jiuzhaigouxian, Sichuan Province; 1998-V-18, coll. Chen D. and Zhang, G.-Q. HBUMM05419, 1 fms; hill foot of Yuxushan, Wenxian, Gansu Province; slate and shrubs, dry; 2006-IX-27, coll. Wu, M., Liu, J.-M., Zheng, W. and Gao, L.-H. HBUMM06490, many fms; Yuxushan, Wenxian, Gansu Province, 1070 m a.s.l., 32.959861°N, 104.678222°E, shrubs and slate, 2011-VIII-07, coll. Wu, M., Xu, Q. and Budha, P. HBUMM06523, many fms and juvs; town of Wenxian, Gansu Province, 1024 m a.s.l., 32.941111°N, 104.668889°E; 2011-VIII-08, coll. Wu, M., Xu, Q. and Budha, P. HBUMM08454, 23 fms, east of the town of Jiuzhaigouxian, Sichuan Province, 2020-VIII, coll. Chen, Z.-G. CZG202008-w6, 1 subadult; Shangdezhen, Wenxian, Gansu Province, near point (32.907414°N, 104.76994°E); 2020-VIII, coll. Chen, Z.-G. CZG202008-w11, 2 fms, near point (32.94936°N, 104.693006°E), town of Wenxian, Gansu Province, 2020-VIII, coll. Chen, Z.-G. HBUMM05437, many fma; 1 fma dissected; Yuxushan, Wenxian, Gansu Province, 32.957259°N, 104.689152°E, shrubs and slate, 2006-IX-27, coll. Wu, M., Liu, J.-M., Zheng, W. and Gao, L.-H. ; DNA voucher HBUMM05411. HBUMM06753, 3 fma, 7 fms; Eastern bank of Baishuijiang, Lihuacun, Jiuzhaigouxian, Sichuan Province, 1425 m, 33.267222°N, 104.234722°E; 2011-VI-14, coll. Wu, M., Xu, Q. and Buhda, P.

Distribution.

Gansu: Wenxian. Sichuan: Jiuzhaigouxian (type locality).

Additional information of shell.

Tiny elongate granules on protoconch, which can be only observed in juvenile shells, are densely and radially arranged. The granules on protoconch are absent in adult shell because of erosion or weathering. On teleoconch the growth lines are indistinct. Spiral grooves are absent throughout.

General anatomy.

Eversible head wart prominent. Jaw arcuate, with three projecting ribs.

Anatomy of genital organs.

Penial sheath covering ~ 1/4 of penis. Penis proximately tubular and distally expanded. Inside penis, two pilasters fusing into one Y-shaped fork at proximal 1/5, two more pilasters of similar thickness also present. Fine pilasters on distal end of penis merging into ~ 8 thick folds. Vas deferens narrow throughout. Vagina between atrium and dart sac not elongated. Vagina between dart sac and insertion of bursa copulatrix duct ~ 1/2 length of dart sac. Dart sac ~ 1/2 length of penis. Love dart ~ 1 mm long, apically 2-bladed. Accessory sac small but externally distinguishable, internally solid, inserting into dart sac at distal 1/3, opening to dart sac chamber. Mucous glands four or five, each singly tubular or simply branched. Proximal accessory sac absent. Bursa copulatrix duct equally thick.

Remarks.

See Laeocathaica parapolytyla Wu, sp. nov.