Crossotarsus emorsus Beeson, 1937 Figures 5, 6

Crossotarsus emorsus Beeson, 1937: 87.

Material examined.

4 males, 1 female (JXAU) China: Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Jinghong City, Nabanhe River Watershed National Nature Reserve, Guomenshan, ca 1030 m, 22°14'46"N, 100°36'10"E, 27.I.2018, log dissection, host Dalbergia assamica, Shengchang Lai leg. ; 1 male, 1 female (RAB); 1 male (JXAU) China: Yunnan Province, Xishuangbanna Dai Autonomous Prefecture, Jinghong City, Damanmi Village, ca 580 m, 22°02'50"N, 100°48'27"E, 20.I.2018, log dissection, host Cassia siamea, Shengchang Lai leg.

Diagnosis.

C. emorsus is similar to C. terminatus but can be distinguished using the characters given in Table 3.

Distribution.

Myanmar, Thailand, Laos (Beaver and Liu 2013; Beaver 2016). New to China (Yunnan).

Host.

The species is recorded from trees in the families Lecythidaceae, Fabaceae, Sterculiaceae and Verbenaceae (Beeson 1937), and is presumably polyphagous (Beaver 2016). Host plants recorded here are: Senna siamea (Lam.) H.S.Irwin & Barneby and Dalbergia assamica Benth. ( Fabaceae).

Molecular data.

The phylogenetic tree for analyzing the evolutionary relationships of 13 taxa including the ingroups ( Crossotarsus species) and the outgroups ( P. contaminatus) was constructed based on four genes (Fig. 7). The BI tree shows the new species ( C. beaveri) and the new combination ( C. brevis) forming a clade, with high node support. These group with Schedl’s (1972a) ' Crossotarsi coleoptrati ' ( C. fractus Sampson, 1912, C. squamulatus and C. terminatus) and cluster with all remaining Crossotarsus species. It confirms that the taxonomic changes and the relationship of C. brevis and C. brevis are correct. It also indicates that C. emorsus, C. fractus, C. squamulatus and C. terminatus should be considered distinct species (as by Beaver and Liu 2013), and not considered synonyms or subspecies (Schedl 1972a).