Affinities of Denticulobasis and Tuberculobasis

According to O'Grady & May (2003), the taxonomic characters commonly used for subfamily identification in Coenagrionidae are continuously distributed and subfamilies should not be recognized within Coenagrionidae until well-supported subdivisions are demonstrated. In view of this, I refrain from ascribing Denticulobasis and Tuberculobasis to any of the extant subfamilies. Denticulobasis and Tuberculobasis are close to Leptobasis and key out to this genus in Von Ellenrieder & Garrison (2008). Both genera share with Metaleptobasis the presence of mesepisternal processes that occur also in some species of Argia (Garrison, 1994) . Table 2 shows the characters that separate these four genera.