Trisaria, new genus

Type species: Trisaria rex Shear, new species

Etymology: The name of the genus is an anagram (rearrangement of letters) of Striaria . It should be treated as feminine in gender.

Diagnosis: Unlike species of Striaria and Amplaria, Trisaria males lack labral spines, are usually smaller (5–10 mm long), and have a crest 4 (c4) on each side of the rings posterior to the fifth suppressed, much lower than c3 and c5. The gonopods are strikingly different and significantly more complex than those of striariines, and the ninth legpair is strongly reduced, with sternum, coxae and telopodites fused or partially so. For additional characters separating the two subfamilies, see the table above.

Description: Small striariid millipedes (Fig. 42; 5–10 mm long). Cranium of head with pebbled sculpture, males lack spine distolaterally on labrum. Two to four poorly formed, pigmented ocelli on each side of head. Mandibular stipes of males with low posteriorventral carina, with small carinal teeth. Collum (Figs. 36, 37) apparently with 10 crests, sixth crest on each side forming expanded lateral margin of collum. Male collum (Fig. 37) longer, broader than in female (Fig. 36). Second ring with broadly expanded pleural margin (Fig. 36). Succeeding rings with 10 crests, but posterior to ring 5, fourth crest on each side progressively suppressed, difficult to see on posteriormost rings—metazonites with strong ventrolateral marginal carinae possibly homologous to sixth crests (Figs. 42, 43). Telson (Fig. 16) lacking crests, short, lobes distinctly divided. Spinnerets short, projecting anteriorly. Paraparocts (para, Fig. 16) densely granular, with 5 smooth depressions and three marginal setae. Male first legpair (l1, Fig. 38, Fig. 17) robust, larger than second legpair, with 6 podomeres (trochanter lacking), prefemora and femora ventrally with very long, straight, blunt-tipped, 4-angled setae (Figs. 17); postfemora, tibiae and tarsi with shorter somewhat flattened setae. Male second legpair coxae with vas deferens opening on the anterior side at the base of a short, heavily setose process (Figs. 38, 39). Male legpair 3 (Fig. 18) with long, blunt extensions from the coxae, sides parallel, distally acuminate and smooth (Figs. 19, 40), without setae on posterior surface, anterior surface with scattered short appressed setae, prefemora with ectal lobes that interlock with pleural margin of corresponding pleurotergite. Male legpairs 4–7 distinctively enlarged and with flattened podomeres; lobes of coxae of legpair 7 covering bases of gonopods (cx7, Fig. 41). All legs with coarsely pebbled surface, scattered, flattened, scale-like setae (Fig. 25), becoming more like ordinary setae distally on tarsi.

Gonopods complex. Sternum (st, Fig. 32) broad, well-defined, not fused to coxae. Coxae small, displaced laterally, with or without setae (cx, Fig. 33). Two angiocoxite processes present, positioned mesally and laterally, each with anterior (mac1, lac 1, Fig. 21) and posterior branches (mac2, lac2, lac3, Fig. 21). Median (lac2) branch of lateral angiocoxite with many thin cuticular processes arranged in comb-like array. Colpocoxite (cc, Fig. 27) variable in form, poorly sclerotized, with scaly cuticle. Legpair 9 (Figs. 23, 28) substantially reduced, sternum, coxae and telopodite articles fused, telopodite interlocking with process from ring 7 (Figs. 20, 41) Legpair 10 (Fig. 24) with coxal glands, coxae conspicuously enlarged.

Vulvae as in Fig. 47.

Distribution: See Map 1. Known at this time only from western Washington.

Notes: The gonopods are described in more detail in the species accounts below.