Pulvinaria psidii Maskell, 1893

(Figs 43, 44)

Pulvinaria psidii Maskell 1893: 223 .

Diagnosis. Dorsal derm without polygonal reticulations; tubular ducts absent; dorsal tubercles present (Fig. 44D). Marginal setae short, mostly with fimbriate apices (Fig. 44E). Stigmatic clefts distinct, each containing 3 stigmatic spines (Figs 43C, 44C). Venter with multilocular disc-pores usually each with 10 loculi, mainly present around vulvar area, a few pores also present laterad of each meta-, meso- and procoxa (Fig. 44H); tubular ducts of 2 types: type I each with a broad inner ductule, mainly present on medial area of head, thorax and abdomen; and type II each with a longer and thinner inner ductule, mainly present in submarginal areas (Figs 43D, 44I); antenna 8 segmented (Fig. 44N) (partially adopted from Williams & Watson, 1990).

Material examined. 29 ♀♀, LAOS, Nan Dist., Sayaboury Prov., 30.x.2014, coll. J.Y. Choi, on Psidium guajava L. ( Myrtaceae); 2 ♀♀, Paksong Dist., Champasak Prov., 16.vii.2016, coll. P.P. Soysouvan, on Coffea sp. ( Rubiaceae); 2 ♀♀, Beng Dist., Oudomxay Prov., 24.viii.2016, on Nephelium hypoleucum Kurz (Sapindaceae) (same collector); 2 ♀♀, on Morinda citrifolia L. ( Rubiaceae) (same locality, date and collector); 2 ♀♀, Pakgnum Dist., Vientiane Capital, 16.vii.2016, on Psidium guajava L. ( Myrtaceae) (same collector).

Hosts. Polyphagous. According to García Morales et al. (2016), P. psidii has been recorded from plants belonging to 141 genera in 67 families.

Distribution. All zoogeographical regions; Oriental Region (India, Indonesia, Malaysia, Philippines, Singapore, Taiwan and Thailand) (García Morales et al. 2016); Laos (new country record).

Economic importance. In Florida, Pulvinaria psidii is an economic pest of ornamentals, especially Ficus spp. (Hamon & Williams 1984); in India it attacks red ginger ( Alpinia purpurata) (Mani et al. 2009). In addition, it was considered to be an important pest of coffee in the tropical South Pacific Region (Williams & Watson 1990), and of citrus and guava (Psidium guajava) in Bangladesh (Bhuiya 1998).

Remarks. Pulvinaria psidii is similar to P. acericola (Walsh & Riley), but is easily distinguished by having strongly expanded and fimbriate marginal setae, and sharply spinose dorsal setae; P. acericola also has fimbriate marginal setae but these are never strongly expanded, and it has bluntly spinose dorsal setae (Hamon & Williams 1984).