Doryscus indochinensis sp. nov. (Figs 4 A, 5)

Types (n= 245). Holotype ♂ (NHMB). THAILAND. Chiang Mai Prov.: Doi Chiang Dao, 1200± 50 m, 19°24’45”N 98°51’30”E, 9–13.V.2009, leg. L. Dembický . Paratypes: 24♂♂, 47♀♀ (NHMB), same data as holotype; 4♂♂, 10♀♀ (NHMB), same localty, 10–16.V.1991, leg. V. Kubáň; 4♂♂, 7♀♀ (NHMB), same locality, 17–24.V.1991, leg. V. Kubáň; 1♀ (SEHU), Doi Inthanon, 5.V.1984, leg. H. Akiyama ; 1♂ (SEHU), Doi Inthanon, Mae-Pan, 5.V.1984, leg. M. Takakuwa ; 1♀ (NHMB), Lansang Nat. Park, 16°48’N 98°57’E, 500 m, 18–24.IV.1991, leg. D. Král; 1♂ (NHMB), Mae Hong son, 28.IV.1991, leg. S. Bily; 1♂, 4♀♀ (NHMB), Mae Hong Son, Ban Huai Po, 1600 m, 9–16.V.1991, leg. J. Horák ; 1♀ (NHMB), same locality, 30.IV.–14. V.19 91, leg. J. Parkač; 1♂ (NHMB), same locality, 1–15.V.1991, leg. S. Bily; 1♂ (UCB), same locality, 1–5.V.1992, leg. S. Jan; 9♂♂, 10♀♀ (NHMB), same locality, 8–17.V.1992, leg. S. Bily; 1♀ (SEHU), Phuping Place, 29.V.1983, leg. Y. Komiya ; 1♂, 1♀ (SEHU), Whuay Kaeo Falls, 1.VI.1983, leg. K. Ikeda ; 2♀♀ (SEHU), same locality, 1.VI.1983, leg. Y. Komiya; 1♀ (SEHU), same locality, 2.VI.1983, leg. H. Akiyama; Kanchanaburi Prov.: 2♀♀ (SEHU), Ban Non Bang, 12.V.1985, leg. Y. Komiya ; 1♂, Daw Dung Cava, 13. V.1 985, leg. Y. Komiya; Nan Prov.: 3♂♂, 2♀♀ (BMNH), Doi Phuka N. P., V.2000, leg. local collector; 2♂♂, 2♀♀ (SEHU), Wiang Sa, 15.V.1993, leg. S. Ohmomo ; Phrae Prov.: 1♀, Wieng Ko Sai N. P., 18.V.1985, leg. Y. Komiya; 3♂♂, 1♀ (NHMB), Soppong-Pai, 1800 m, 1–6.V.1991, leg. Pacholátko ; 5♂♂, 6♀♀ (NHMB), Umphang riv., 1000 m, 16°07’N 99°00’E, 28.IV.–6.V.1991, V. Kubáň ; CHINA. Yunnan: 1♂ (NHMB), Jizushan, 25°58’N 100°21’E, 2500–2700 m, 6–10.VII.1994, leg. V. Kubáň ; 1♂ (NHMB), Weibaoshan E slope, 25°16’N 100°27’E, 2500– 1800 m, 1.VII.1992, leg. D. Král; 21♂♂, 22♀♀ (NHMB), Yipinglang, 25°03’N 101°55’E, 2000 m, 8–10.VI.1993, leg. V. Kubáň ; 6♂♂, 4♀♀ (NHMB), same but with “Bolm”; 1♂, 9♀♀ (NHMB), same locality, 17–20.VI.1994, leg. V. Kubáň; INDIA. 1♂ (NHMB), Karnataka, W. Ghats, 20 km W Talguppa, Jog Falls, 14°14’N 74°44’E, 500± 200 m, 22–28.V.2002, leg. P. Pacholáko ; LAOS. Bolikhamxay Prov.: 1♀ (RBCN), Ban Nok env., 220 m, Route no 8, 18°08’7 N 104°28’1 E, 9–14.V.1998, leg. E. Jendek & O. Sausa ; Houa Phan Prov.: 1♂ (NHMB), Ban Saluei + Phou Pane Mt., 20°12–13.5’N 103°59.5’– 104°01’E, 1340–1870 m, 15.IV.–15.V.2008, leg. Lao collectors ; 1♂, 2♀♀ (NHMB), same locality,

9–17.VI.2009, leg. M. Geiser; 1♂ (EUMJ), Ban Saluei, Xam Neua, 27.IV.–22.V.2007, leg. J. Yamasako ; Xieng Khouang Prov.: 1♀ (EUMJ), Nong Het, 16–18.VI.2005, leg. J. Yamasako ; 1♂ (NHMB), 30 km NE Phonsavan: Ban Na Lam + Phou Sane Mt., 19°37–8’N 103°20–1’E, 1300–1700 m, 10–30.V.2009, leg. M. Geiser ; VIETNAM. 3♀♀ (NHMB), Buonloi, 1982, leg. L. Medvedev; 1♂ (NME), 100km SSW Hanoi, 40km SW Than Hoa, Ben En N. P., 50 m, 5–8.VII.1997, leg. A. Napolov ; 2♂♂ (NME), same but with “ 23–28.VII.1997 ”; 1♂ (NME), same but with “ 5.VII.1990 ”; Ninh Binh Prov.: 1♂, 1♀ (NME), 90km SW Hanoi, Cucu Phuoung N. P., primat rescue centre, 25.IV.2012, 20°14’24”N 105°42’53”E, leg. A. Weigel .

Description. Length 5.0– 6.5 mm, width 2.1–2.6 mm. General color yellowish brown (Fig. 4 A). First antennomere not modified in males (Fig. 5 A), length ratios of antennomeres I–XI 1.0: 0.4: 0.5: 0.8: 0.8: 0.8: 0.8: 0.8: 0.7: 0.7: 0.8, length to width ratios of antennomeres I–XI 3.7: 1.8: 2.7: 3.9: 3.9: 3.9: 4.0: 3.9: 3.8: 3.5: 4.1; first antennomere in female more slender (Fig. 5 B), length ratios of antennomeres I–XI 1.0: 0.4: 0.6: 0.8: 0.7: 0.7: 0.7: 0.7: 0.7: 0.7: 0.9, length to width ratios of antennomeres I–XI 4.0: 1.9: 2.9: 3.9: 3.8: 3.6: 3.8: 4.1: 3.8: 3.6: 4.7. Elytra 1.8 x longer than wide; surface with erect, yellowish brown, dense, elongate setae; punctures coarse. Apical margin of last abdominal ventrite with two well developed incisions in male (Fig. 5 G); weakly and medially depressed in females (Fig. 5 H). Penis (Figs 5 C–5D) slender, gradually narrowed towards apex; apex of dorsal surface with deep medial notch; apex of ventral surface widely rounded; almost straight in lateral view, apex with extremely narrow notch between dorsal and ventral surfaces; ventral surface smooth. Internal sac without dense setae; median endophallic spiculae elongate, about 0.7 x as long as penis, inner margin straight, covered with minute spines, basally conjoined; one short endophallic spiculum between median endophallic spiculae, basally covered with setae, apically tapering, ventrally curved in lateral view; two pairs of dorsal endophallic spiculae elongate, basally conjoined, apex rounded, one pair a slightly shorter than the other; 11 pair of lateral endophallic spiculae that are horn-like and small, located from near base to on-half length; one ringlike spiculum anteriorly connected with first horn-like lateral endophallic spiculae, anteriorly open. Gonocoxae (Fig. 5 E) elongate, 8.8 x longer than wide, sides narrowed in apical one-third; apices narrowly rounded; apex with nine setae; slender from midpoint to base but abruptly widened at base. Ventrite VIII (Fig. 5 F) well sclerotized; spiculum long; apex square, apical margin truncate, with several short setae at sides; disc with several elongate setae at sides. Receptacle of spermatheca (Fig. 5 I) wide, narrowly separated from pump; pump strongly curved and apically tapering; proximal spermathecal duct short, connected with receptacle.

Color variability. Some specimens have darkened tibiae and tarsi.

Differential diagnosis. See diagnosis of Doryscus geiseri sp. nov.

Etymology. The species name refers to the distribution of this new species.

Distribution. China (Yunnan), India, Laos, Thailand, Vietnam.