Tupala occulta sp. nov.

(Figs 1–50, 55)

Etymology. Specific epithet is derived from Latin ‘ occultus ’, meaning figuratively secret and refers to unclear provenance of the taxon (see Discussion).

Diagnosis. Face with lateral portions of frons yellowish; clypeus black. Tegmen with cell C1 shorter than cell C3; cell C5 more than twice as long as cell C3.

Description. General coloration dark brown. Lateral portion of frons yellowish, median streak brown; clypeus black. Lateral portion of head brownish, anterior portion of gena yellowish anteriorly; lora dark brown. Posterior margin of scutellum yellowish. Wings transparent, with brownish veins, indistinct lighter spots in median portion. Legs dark brown; metatibia brownish at base, yellowish toward apex. Ventral side of abdomen brown, dorsal side with pattern of spots and streaks. Anal tube brown, with lighter median band and yellowish apical margin. Genital styles brownish.

Total length 0.74–0.86 mm.

Head. Vertex: proportion A/B = 0.37– 0.47, Frons: proportion C/E = 0.15–0.16, proportion D/E = 0.38–0.40. Thorax. Pronotum: proportion F/B = 0.30–0.33; Mesonotum: proportion G/F = 2.82–2.85, proportion G/B+F = 0.55–0.70, proportion G/H = 0.68–0.70. Tegmina: proportion I/J = 2.95–3.11.

Type locality and occurrence. Madagascar, Antsiranana Province, Sava Region, Maroantsetra (Fig. 55). Type material. Holotype male and 2 male paratypes, all labeled: [Coll. R.I.Sc.N.B. / Madagascar / Maroantsetra II-1919 / Ex. Coll. Le Moult]. Specimens deposited in Royal Belgian Institute of Natural Sciences, Brussels, Belgium.