The Eulichas fasciolata species complex

The complex contains four species from the Greater Sunda Island of Borneo. They differ from all other Eulichas species in the relatively short parallel sided parameres with a characteristic twisted apex. The subapical parameral hook is well developed, the subbasal hook is reduced. The median lobe is narrowly lanceolate, always longer than the parameres. The phallobase is about the same length as parameres and median lobe combined.

Eulichas fasciolata (Fairmaire, 1898)

(Figs. 21, 41, 61, 81)

Lichas fasciolata Fairmaire, 1898: 388 (original description, Kina-Balu).

Eulichas fasciolata (Fairmaire, 1898): Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue).

Type locality. “Kina-Balu” [Sabah state, Malaysia].

Type material. Not found in MNHN.

Additional material studied. 121 specimens — MALAYSIA: SABAH: 1Ƥ (SMTD); 2ƤƤ, coll. C. Felsche (SMTD); 2633 13ƤƤ, Kinabalu (DEIC, HNHM, MNHN, NHMB, SMTD); 633 8ƤƤ, Mount Kinabalu, v.–viii.1903, J. Waterstradt (MNHN, RMNH); 233, Kinabalu National Park, Poring, 500 m, 29.xi.–2.xii.1996, W. Schawaller leg. (SMNS); 433 2ƤƤ, Crocker Range [Mts.], iv.1990 (MCSN); 433, Crocker Range Mts., 50 km E Kota Kinabalu, Gunung Emas [Mt.], 16.–27.iv.1993, I. Jeniš leg. (IJCN, NHMW, NMPC); 13, Keningau, 800 m, iv.1987 (NMPC); 633 1Ƥ, Crocker Range, Keningau, v.1993 (NMPC); 13, 8, 5 miles NW of Keningau, Kimanis road, at light, 11.v.1982, D. Burckhardt leg. (MNHG); 333, Keningau – Kimanis, km 25, iv.1994 (NHMW); 13, NW of Keningau, C.R.Park, Kimanis road, 10–15 km oberh., Park HQ, 05°26’N 116°05’E, 1100–1300 m, 2.–4.ix.1998, D. Bratsch & C. Häuser leg. (SMNS); 433 3ƤƤ, Crocker Range, 16 miles NW of Keningau, 8.ix.1982, S. Nagai leg. (EUMC); 13, same label data, but 12.ii.1982 (EUMC); 2 33, same label data, but 1400 m, 2.–26.iv. 1984 (EUMC); 1Ƥ, same label data, but 4.iv.1984 (EUMC); 13, same label data, but 9.iv.1984 (EUMC); 13, same label data, but 20.iv.1984 (EUMC); 13, Bunsit, 13.iii.1983, S. Nagai leg. (EUMC); 233, 105 km S Beaufort, Long Pasia Area, airstrip Long Pasia, 04°24’N, 115°43’E, 1000 m, at light, Recultivated area, 16.iv.1987, J. van Tol & J. Huisman leg. (RMNH); 333, 70 km W of Lahad Datu, Danum valley, 150 m, at light, 13.xii.1989, M.J. & J.P. Duffels leg. (ZMAN); 13, Batu Punggul [ca. 04°38’N 116°35’E], Banjaran Maitland, 25.–27.v.1995, I. Jeniš leg. (MNHG); 13, 50 km S Tomani, 4.v.1999, M. Snížek leg. (VKCZ); 1Ƥ, Trus Madi, 2200 m, 22.iv.1990, Martini leg. (NMPC); 233 1Ƥ, Mount Trus Madi, 1100 m, iv.2000, K. & B. Martini leg. (VKCZ); 633, Mt. Trus Madi, v.2004, S. Chew leg. (NMPC); 13, same data, but 3.iii.2005 (BMNH); 233, Mamut, 17.v.1979, N. Nishikawa leg. (EUMC); 13, Sepilok, Sandakan, 6.ix.1982, S. Nagai leg. (EUMC); 13, 80 km E of Ranau, Telupid, 800 m, 20.iv.1979, S. Nagai leg. (EUMC); 13, Tinamantawaran, near Ranau, 16.iv.1983, S. Nagai leg. (EUMC). SARAWAK: 13, Gunong Mulu National Park, 150 m, at light, ii.1978, J.D. Holloway et al. (BMNH); 2ƤƤ, same label data, but 4th Division, 150–200 m, v.–viii.1978, P.M. Hammond & J.E. Marshall leg. (BMNH).

Description. Habitus elongate, fusiform. Body colouring redish-brown to brown. Pale part of setation consists of recumbent brownish-yellow setae covering uniformly head, pronotum and ventral part, and forming rather indistinct ocellations on elytra (Fig. 21).

Measurements. Males: 23–32 mm; females: 30–35 mm.

Head punctation consists of sparse moderately large setigerous punctures. Antenna long, slender, last antennomere narrowly oval, 3.19–3.58 times as long as wide (Fig. 41), its ventral side smooth.

Pronotum trapezoidal, ca. 1.90–1.91 times as wide as long. Sides almost straight, in some specimens with indistinct sinuation behind the middle, hind angles prominent (Fig. 61). The disc convex with two lateral and one basal shallow depression. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally.

Elytra with numerous longitudinal rows of moderately large punctures, and fine interstitial setigerous punctures.

Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally regularly rounded to apex.

Male. Aedeagus with phallobase shorter than parameres. Parameres parallel sided, with twisted emarginated apex; their distal part rather short. Median lobe lanceolate, with thin apical part; exceeding parameres (Fig. 81).

Female. Similar to male in habitus, but larger. Antenna shorter, and more slender.

Remarks. Although the type material of the species was not found/recognised in the MNHN, the large number of additional specimens from Kinabalu deposited in various collections has allowed confident identification and redescription of the species.

Collection circumstances. Collected at light in the primary forest.

Distribution. Known from the Malay part of the Greater Sunda Island of Borneo – Sabah and Sarawak.