The longicollis group
The longicollis group is characterized by the following combination of characters: the pronotal puncturation uniform on the whole surface, the pronotal pubescence at midline directed entirely anteriorly or posteriorly in apical half and anteriorly in basal half, the pronotal pubescence at posterolateral parts directed laterally, the pronotal pubescence pruinose in appearance, the pronotal basal impression relatively large and deep, well visible, and the posterior margin of tergite VIII without row of apical spines.