Harpalus (Pseudoophonus) calceatus (Duftschmid, 1812)

(Figs 10, 86–87)

Carabus calceatus Duftschmid, 1812: 81 . Type locality: Linz, Austria.

Anisodactylus nonsignatus Krynicki, 1832: 72 . Type locality: "Polt.[ava] et Eka.[terinoslav] (Slavenoserbskiy uezd Sukhodol", Ukraine.

Harpalus calcitrapus Motschulsky, 1844: 216 . Type locality: "mont d'Oulou-Tau, Kirguises; Nertchinsk", Russia.

Anisodactylus propinquus Ballion, 1870: 328, syn. n. Type locality: "Bei Tschemkent", Kazakhstan.

Ophonus itoshimanus Habu, 1954: 281 . Type locality: Raizan-mura, Itoshima-gun, Fukuoka Prefecture, Kyushu, Japan.

Type material. Lectotype (present designation) of Anisodactylus nonsignatus: ♀, labeled with a small, lozengeshaped piece of white paper, " nonsignatus Kr. ", and "Museum of Nature, Kharkov National University" (MNKH).

Syntype of H. calcitrapus: 1 specimen (without head and pronotum), " Platus calcitratus Mots., Sib. or." (ZMM).

Additional material examined. More than 3400 specimens (more than 2500 from China), including the following specimens from Nei Mongol, Beijing and Guangxi. China. NEI MONGOL: 2 ♂, 1 ♀, E Bayan Hot, 38.835°N 105.768°E, 1750–1800 m, 3–6.VIII.2011, T. Zhura leg. (cIB&IK). BEIJING: 1 ♀, "Pekin", 22–29.VII.1907, Vasil'ev leg. (ZIN). GUANGXI: 1 ♂, Daqingshan Mountains, 7.V.1983, Liao Subai leg. (IOZ).

Diagnosis. The species is easily recognizable by having spines on ventral side of tarsomere 5 (Fig. 10). Internal sac of aedeagus (Figs 86–87) is armed with medial spiny patch; occasionally an additional small spiny patch is present in the apical portion of the median lobe. Body length 12.0– 14.2 mm.

Distribution. China: the northern, eastern and south-eastern parts. The species was recorded from Xinjiang, Liaoning, Hebei, Shanxi, Shaanxi, Sichuan, and Yunnan (Kataev et al. 2003); we examined additionally specimens from Nei Mongol, Beijing and Guangxi. According to Hua (2002), this species occurs also in Heilongjiang, Jilin, Henan, Gansu, Anhui, Jiangxi and Fujian. Harpalus calceatus is recorded here from Guangxi for the first time.

General distribution includes the Palaearctic from Portugal to Japan and North Korea (Kataev et al. 2003).

Remarks. Anisodactylus propinquus was described from Chemkent environments and formerly considered a synonym of A. nemorivagus Duftschmid though the type specimens have apparently never been reexamined (Noonan 1996). Ilya I. Kabak (pers. com.) studied the syntypes of this taxon in the former collection of E. Ballion at ZMO (1 ♂ and 1 ♀, each with pinned label "Tschemkent" and with common bottom label " Anisodactylus propinquus Ball. ") and found that they belong to Harpalus calceatus . Based on these data, we treat A. propinquus as a synonym of the latter species.