27. "Gulella" (Aenigmigulella) aenigmatica (E. A. Smith, 1890) Figs 57-59

Ennea aenigmatica Smith 1890: 164; pl. 6, fig. 11

Notes.

Pemba material matches Usambara material (NMW) of this Eastern Arc species very well. No species of Aenigmigulella has previously been reported from the coastal region (Verdcourt 2000, 2006) and it is unknown from Unguja. If native to Pemba it would thus suggest a special Eastern Arc affinity. However we note it was not found in forest reserves apart from a juvenile at Msitu Mkuu FR (Table 2). On Pemba, juvenile shells have complex dentition at a variety of stages. The living animal is cream-coloured. Recent systematic work (Rowson 2010) indicates "Gulella" aenigmatica does not belong in the genus Gulella L. Pfeiffer and a genus-level revision is in progress.