Genus Paragryllodes Karny, 1909

(Figs 15–17)

Type species. Paragryllodes borgerti Karny, 1909

Other species included. See Kaltenbach (1982) and Desutter-Grandcolas (1999) for argumented species lists. Two additional species close to Paragryllodes amani Desutter-Grandcolas, n. sp. are available from Tanzania, Kimbosa and Tanghanika (MNHN, ZMC).

Distribution. Tropical Western and Eastern Africa, Madagascar, the Andaman island, Western India, Sri Lanka.

Diagnosis. See Kaltenbach (1982) and Desutter-Grandcolas (1999). This genus is characterized by its general shape and short FWs, dark uniform coloration and more or less shiny apparence (Fig. 15). TIII with four outer and three / four inner subapical spurs. Male. FWs thick and more or less corneous; stridulatory apparatus complete, but hardly visible and partly reticulate (Fig. 16 B). Male genitalia characterized by very long, thin and acute ectophallic dorsal valves, extending well beyond the phallic complexe; pseudepiphallus often densely covered with strong setae (Fig. 17). Female. FWs present, short, partly overlapping or not; venation weak, with longitudinal parallel veins (Fig. 15 F). Subgenital plate transverse, the apex bisinuate. Ovipositor with smooth apex. Female genitalia: copulatory papilla triangular, and more or less elongate (Fig. 16 O–N).

Phylogenetic relationships. In Chintauan-Marquier et al. 's (2013, 2015) molecular phylogeny, Paragryllodes is the sister group of a wide clade including present-day Paragryllinae, ( Caltathra Otte, 1987 — Protathra Desutter- Grandcolas, 1997), Brevizacla Gorochov, 2003 and Mikluchomaklaia Gorochov, 1986; the whole clade is the sister group of the neotropical Aclodae group ( Aclodes Hebard, 1928, Paraclodes Desutter-Grandcolas, 1992 and Uvaroviella Chopard, 1923, see Desutter-Grandcolas 1992). While this topology has to be confirmed with additional data, the position of Paragryllodes within this clade seems well-supported.