Incisus Grishin, new genus

http://zoobank.org/ 2ED74AC8-4CCF-4796-952B-0292620F0855

Type species. Antigonus incisus Mabille, 1878 .

Definition. A genus without close relatives. Formerly placed in Staphylus Godman and Salvin, [1896], incisus is not monophyletic with the Staphylus type species Helias ascalaphus Staudinger, 1876, nor with Scantilla opites Godman and Salvin, [1896] (a junior subjective synonym of Tagiades vincula Plötz, 1886) the type species of Scantilla Godman and Salvin, [1896], a genus-group name treated as a subjective synonym of Staphylus (Fig. 4). Keys to E.32.35b in Evans (1953). Readily distinguished from its relatives by the wing shape: forewing inner margin concave, hindwing outer margin excavate; forewing with hyaline apical spots; uncus long and narrow, pointed at the tip, without side processes, no defined gnathos, valva less than twice longer than wide, penis shorter that in Staphylus, with a spined cornutus. In DNA, a combination of the following base pairs is diagnostic: aly23605.8.3:C247 T, aly5294.23.1:A538C, aly9588.6.1: T843 C, aly 1370.7.2:A1760G, aly345.13.8:A62C .

Etymology. The name, a masculine noun in the nominative singular, echoes the type species name.

Species included. The type species, Staphylus fasciatus Hayward, 1933 and Pholisora (?) angulata Bell, 1937.

Parent taxon. Tribe Carcharodini Verity, 1940 .

Comments. Steinhauser doubted the placement of some of these species in Staphylus on the basis of morphological analysis (Steinhauser 1989), and he was correct.