Updated key couplets to Ochodaeus spp. in Madagascar

The following couplet changes to the species key in Paulsen (2019) allow for determination of all Madagascan species having a granulate pronotum, including the three newly described species.

2(1). Elytral intervals with setose tubercles of 2 sizes, with setae of larger tubercles flattened, scale-like........................................................ O. pygmaeus Paulian — Elytral intervals with tubercles either absent or of uniform size and without flattened setae. 3

3(2). Clypeus with median tubercle or small horn that is distinct from anterior margin (Fig. 4–6)........................................................................... 4 — Clypeus lacking free tubercle or horn, anterior margin usually thickened medially (Fig. 7–8). 6

4(3). Mentum with a knobby protuberance on each side (Fig. 1d).... O. meridialis Paulsen, n. sp. — Mentum lacking protuberances laterally............................................ 5

5(4). Clypeus (Fig. 5) with median tubercle anterior to frontoclypeal suture; anterior margin of clypeus indistinct and eroded medially........................ O. umbonulus Paulsen, n. sp.

— Clypeus (Fig. 6) with small horn located on frontoclypeal suture; anterior margin of clypeus distinct and entire throughout.......................... O. iniquipes Paulsen, n. sp.

6(3). Median thickening of clypeal bead variable, lateral thickening always evident (Fig. 7); body length usually> 9.0 mm........................................ O. isoanalensis Paulian

— Median thickening of clypeal bead simply convex, bead slender laterally (Fig. 8); body length usually 4.5–8.0 mm............................................ O. miliaris Klug