Speonemadus vandalitiae (Heyden, 1870)

Figs 10, 17, 37

Catops vandalitiae Heyden, 1870: 97 .

Ptomaphagus vandalitiae (Heyden); Heyden 1883: 368.

Anemadus vandalitiae (Heyden): Uhagón 1890: 32.

Hormosacus vandalitiae (Heyden); Jeannel 1936: 214.

Hormosacus vandalitiae pyrenaeus Jeannel (1936: 214); Blas 1984: 252 (synonymy)

Hormosacus vandalitiae pyrenaeus Jeannel;

Speonemadus vandalitiae (Heyden); Giachino & Vailati 1993: 183.

Type locality. "Sierra della Nieve prope Ronda Andalusiae" (Heyden 1870), Spain.

Lectotype (Fig. 37): ♂, "Ronda, Heyden" [hw L. von Heyden], " Syntypus " [red, printed], microscopic slide with aedeagus and "1563" [hw], " Coll. L. v. Heyden, DEI Eberswalde" [printed], " Lectotypus ♂, Catops Vandalitiae Heyden, P. M. Giachino & D. Vailati des. 1992" [red, printed] (SDEI).

Diagnosis. Apex of median lobe acuminated (Fig. 10); parameres very long, forming a marked angle, with apex round and expansion on inner side (Fig. 17) (see other characters under S. gracilis, see above). Molecular data place S. vandalitiae as sister species of S. pulchellus; although both species belong to the vandalitiae -group sensu Giachino & Vailati (1993), their aedeagi are considerably different (compare Figs 10 and 11), with much shorter and robust parameres relative to the length of the median lobe in S. pulchellus .

Distribution (Fig. 39; Appendix). Iberian Peninsula, extending to the French Pyrenees (Jeannel 1936; Giachino & Vailati 1993; Perreau 2000, 2004; Salgado et al. 2008; Agirre & Blas 2009; Fresneda & Salgado 2017). In Spain there are records from Albacete, Ciudad Real, Granada, Huesca and Madrid (Fresneda et al. 2007), Málaga (Heyden 1870; Fuente 1925; Giachino & Vailati 1993), Murcia (Fresneda et al. 2007), Tarragona and Toledo; and in France from Basses-Pyrénées (Jeannel 1936). However, some of these records may refer to S. gracilis, which has been considered to be a synonym of S. vandalitiae since Reitter (1885) (see section 3.3. above).