Similiphora intermedia (C.B. Adams, 1850)

Fig. 94

Cerithium intermedium C.B. Adams, 1850: 119 .

Triforis intermedius – Dautzenberg 1900: 195.

Triforis intermedia – Dall & Simpson 1901: 423.

Triphora pulchella auct. non C.B. Adams, 1850 – Lange-de-Morretes 1949: 80. — Parker & Curray 1956: 2434. — Porta & Porta 1960: 42. — Houbrick 1968: 14. — Rios 1970: 46; 1975: 51, pl. 13 fig. 190; 1985: 161, pl. 53 fig. 764; 1994: 95, pl. 31 fig. 377; 2009: 173. — Abbott 1974: 112, fig. 1135. — Vokes & Vokes 1983: 18, pl. 27 fig. 17. — Robinson & Montoya 1987: 384. — Absalão 1989: 3. — Absalão et al. 2006: 238. — Santos et al. 2007: 226. — Agudo-Padrón 2015: 65.

Cerithium intermedium – Clench & Turner 1950: 293, pl. 38 fig. 9.

Triphora intermedia – Parker & Curray 1956: 2434. — Warmke & Abbott 1962: 77. — Cooley 1978: 60. — Robinson & Montoya 1987: 384. — De Jong & Coomans 1988: 50. — Díaz 1994: 33. — Hess & Abbott 1994: 147.

Triphora sp. “C” – Rice & Kornicker 1962: 120, pl. 2 fig. 16.

Triphora (Cosmotriphora) pulchella – Odé 1989: 111, non C.B. Adams, 1850.

Similiphora intermedia – Rolán & Fernández-Garcés 1995: 12, figs 17–19; 2007: 24, pl. 5 figs 6–12; 2015: 54, pl. 4 figs m–n, not fig. k. — Redfern 2001: 68, pl. 33 fig. 286; 2013: 130, fig. 372. — Espinosa & Ortea 2001: 20. — Sevilla et al. 2003: 342. — Espinosa et al. 2007: 74, fig. 55; 2012: 257. — Lee 2009: 91. — Garcia & Lee 2011. — Zhang 2011: 101, fig. 303. — Lamy & Pointier 2018: 289, pl. 92 fig. 8. — Fernandes & Pimenta 2019a: 39, figs 2j, 25–27.

Material examined

BRAZIL – North Brazil • 2 specs; REVIZEE-Norte III st. 176; UFMA * • 1 spec.; REVIZEE-Norte III st. 211A; UFMA *. – Pará • 1 spec.; 00º10′ S, 46º00′ W; depth 35 m; Jan. 1981; MORG 52607. – Rio Grande do Norte • 1 spec.; BPot 1-MR11; MNRJ 31460 * • 1 spec.; BPot 1-MR22; MNRJ 31461 * • 1 spec.; BPot 1-MR23; MNRJ 31462 * • 2 specs; BPot 1-MR25; MNRJ 31463 * • 2 specs; BPot 1-MR31; MNRJ 31464 * • 4 specs; BPot 1-MR32; MNRJ 31465 * • 1 spec.; BPot 1-MR34; MNRJ 31466 * • 9 specs; BPot 1-MR41; MNRJ 31467 * • 5 specs; BPot 1-MR42; MNRJ 31468 * • 3 specs; BPot 1-MR43; MNRJ 31469 * • 1 spec.; BPot 1-MR45; MNRJ 31470 * • 2 specs; BPot 2-MR22; MNRJ 31471 * • 1 spec.; BPot 2-MR23; MNRJ 31472 * • 4 specs; BPot 2-MR32; MNRJ 31473 * • 1 spec.; BPot 2-MR33; MNRJ 31479 * • 1 spec.; BPot 2-MR34; MNRJ 31474 * • 2 specs; BPot 2-MR41; MNRJ 31475 * • 3 specs; BPot 2-MR42; MNRJ 31476 * • 4 specs; BPot 2-MR 44; MNRJ 31477 * • 4 specs; BPot 2-MR45; MNRJ 31478 * • 1 spec.; Genipabu; 1966; Cabral-Nasser leg.; MORG 12867. – Alagoas • 1 spec.; Maceió, Ponta Verde; 1992; IBUFRJ 4351 • 2 specs; Maceió, Jaraguá; MNRJ 18612 * • 20 specs; Maceió, Jaraguá; MORG 33738 • 2 specs; Petro/MAR st. 21; UFS. – Sergipe • 1 spec.; Petro/UFS st. E5-A2; with soft part preservation; UFS • 1 spec.; Petro/UFS st. E6-A2; UFS . – Bahia • 3 specs; Salvador, Itapuã; Apr. 1982; G. Oliveira leg.; MORG 22195 • 1 spec.; Salvador, Itapuã; Feb. 1990; P.M.S. Costa leg.; MORG 27669 • 5 specs; Salvador, Itapuã; MORG 33731 • 3 specs; Salvador, Itapuã; MORG 52616 • 1 spec.; Salvador, Itapuã; MZSP 63878 • 1 spec.; Salvador, Itapuã; MZSP 133313 • 1 spec.; Salvador, Itapuã; MZSP 133318 • 1 spec.; 13º29′22″ S, 38º48′43″ W; Jun. 2007; MNRJ 32914 * • 1 spec.; Cairu, Morro de São Paulo, Praia de Garapuá; depth 5 m; 2011; P. Coelho-Filho leg.; with soft part preservation; MZSP 100746 • 1 spec.; off Boipeba; depth 35–55 m; Dec. 2002; M. Ximenez leg.; MORG 43762 • 7 specs; Ilhéus; MNRJ 32371 * • 3 specs; Ilhéus; MNRJ 32974 * • 1 spec.; off Canavieiras; Apr. 2011; MNRJ 30711 * • 1 spec.; off Canavieiras; Apr. 2011; MNRJ 30721 * • 1 spec.; off Canavieiras; Apr. 2011; MNRJ 30755 * • 3 specs; off Canavieiras; Apr. 2011; MNRJ 33121 * • 1 spec.; Abrolhos; Feb. 1978; MORG 52609 • 1 spec.; Abrolhos; Feb. 1978; MORG 20266 • 9 specs; Abrolhos; Jan. 1985; MORG 23827 • 1 spec.; Abrolhos; Jan. 1985; MORG 52606. – Espírito Santo • 9 specs; REVIZEE-Central C1-VV31; IBUFRJ 19483 • 1 spec.; REVIZEE-Central C1-VV33; MORG 39115 • 3 specs; REVIZEE-Central C1- C65; IBUFRJ 19460 • 1 spec.; REVIZEE-Central C1-C65; IBUFRJ 19498 • 1 spec.; REVIZEE-Central C1-C65; MORG 40895 • 7 specs; REVIZEE-Central C1-C65; MORG 52599 • 2 specs; REVIZEE-Central C1-C64; IBUFRJ 19494 • 1 spec.; REVIZEE-Central C1-VV38; IBUFRJ 19524 • 5 specs; REVIZEE-Central C1-VV38; IBUFRJ 19606 • 3 specs; 19º22′46″ S, 39º39′26″ W; depth 12 m; Oct. 2003; MNRJ 31166 * • 1 spec.; 19º24′23″ S, 39º34′42″ W; depth 24 m; Oct. 2003; MNRJ 31162 * • 6 specs; 19º24′52″ S, 39º22′00″ W; depth 42 m; Oct. 2003; MNRJ 30753 * • 4 specs; 19º25′ S, 39º22′ W; depth 55 m; Oct. 2003; MNRJ 30763 * • 3 specs; 19º25′05″ S, 39º15′54″ W; depth 65 m; Oct. 2003; MNRJ 30701 * • 6 specs; 19º25′08″ S, 39º15′59″ W; depth 65 m; Oct. 2003; MNRJ 30768 * • 3 specs; 19º25′34″ S, 39º22′16″ W; depth 42 m; Oct. 2003; MNRJ 30717 * • 4 specs; 19º25′34″ S, 39º22′16″ W; depth 42 m; Oct. 2003; MNRJ 31163 * • 9 specs; 19º25′37″ S, 39º22′22″ W; depth 43 m; Oct. 2003; MNRJ 30748 * • 4 specs; 19º25′37″ S, 39º22′22″ W; depth 43 m; Oct. 2003; MNRJ 30764 * • 1 spec.; 19º25′37″ S, 39º22′22″ W; depth 43 m; Oct. 2003; MNRJ 30891 * • 2 specs; 19º25′45″ S, 39º24′58″ W; depth 35 m; Oct. 2003; MNRJ 30745 * • 7 specs; 19º25′45″ S, 39º24′58″ W; depth 35 m; Oct. 2003; MNRJ 30759 * • 1 spec.; 19º25′45″ S, 39º24′58″ W; depth 35 m; Oct. 2003; MNRJ 32126 * • 10 specs; 19º25′54″ S, 39º24′58″ W; depth 50 m; Oct. 2003; MNRJ 30747 * • 1 spec.; 19º26′ S, 39º22′ W; depth 46–55 m; Oct. 2003; MNRJ 30705 * • 2 specs; 19º26′ S, 39º22′ W; depth 46–55 m; Oct. 2003; MNRJ 30724 * • 7 specs; 19º26′ S, 39º22′ W; depth 46–55 m; Oct. 2003; MNRJ 30930 * • 1 spec.; 19º26′ S, 39º22′ W; depth 46–55 m; Oct. 2003; MNRJ 32150 * • 2 specs; 19º26′ S, 39º22′ W; depth 46–55 m; Oct. 2003; MNRJ 32163 * • 3 specs; 19º26′ S, 39º22′ W;depth 46–55 m; Oct.2003 MNRJ 32165 * • 3 specs; 19º26′03″ S, 39º22′35″ W; depth 44 m; Oct. 2003; MNRJ 32431 * • 1 spec.; 19º26′03″ S, 39º22′35″ W; depth 44 m; Oct. 2003; MNRJ 32467 * • 4 specs; 19º26′19″ S, 39º15′17″ W; depth 53 m; Oct. 2003; MNRJ 19035 * • 1 spec.; 19º26′19″ S, 39º15′17″ W; depth 53 m; Oct. 2003; MNRJ 30712 * • 2 specs; 19º26′22″ S, 39º15′22″ W; 51 m; Oct. 2003; MNRJ 31167 * • 1 spec.; 19º26′36″ S, 39º22′02″ W; depth 50 m; Oct. 2003; MNRJ 30765 * • 4 specs; REVIZEE-Central C6-R2#1-1; IBUFRJ 16257 • 6 specs; REVIZEE-Central C1-VV24; IBUFRJ 19471 • 6 specs; REVIZEE-Central C1-VV24; MORG 40544 • 5 specs; 19º40′26″ S, 39º36′19″ W; depth 40 m; 2 Dec. 2011; OC. Ship Seward Johnson; MNRJ 34829 * • 1 spec.; 20º10′02″ S, 40º08′31″ W; depth 25 m; 2 Dec. 2011; OC. Ship Seward Johnson; MNRJ 34981 * • 12 specs; 20º14′ S, 40º12′ W; Jun. 2008; MNRJ 32647 * • 7 specs; 20º18′ S, 40º14′ W; Mar. 1993; V. Abud leg.; MORG 40025 • 4 specs; Guarapari; 1970; MORG 16296 • 2 specs; Guarapari; Jan. 2013; MNRJ 34027 * • 1 spec.; Guarapari, off Meaípe; depth 20–25 m; Sep. 1997; A. Bodart leg.; MZSP 64403 • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Sep. 2007; MNRJ 30836 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Sep. 2007; MNRJ 30897 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Sep. 2007; MNRJ 30907 * • 26 specs; Porto de Ubú; 20º47′ S, 40º32′ W, Nov. 2007; MNRJ 32410 * • 5 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31011 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31013 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31014 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31021 * • 3 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31022 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Oct. 2008; MNRJ 31026 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Nov. 2009; MNRJ 31006 * • 4 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Nov. 2009; MNRJ 31008 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31035 * • 3 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31037 * • 3 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31049 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31055 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31059 * • 4 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31066 * • 3 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31067 * • 3 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31072 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31074 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31084 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31090 * • 2 specs; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31095 * • 1 spec.; Porto de Ubú; 20º47′ S, 40º32′ W; Mar. 2010; MNRJ 31098 * • 5 specs; Piúma; 1993; IBUFRJ 5595 • 29 specs; Piúma; 1993; IBUFRJ 8624 • 1 spec.; Piúma; 1998; L.R.L. Simone leg.; MZSP 84798 . – Campos Basin (Espírito Santo/ Rio de Janeiro) • 2 specs; 21º09′ S, 40º31′ W; 27 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7557 • 4 specs; 20º47′ S, 40º26′ W; 29 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7560 • 9 specs; 20º53′ S, 40º12′ W; 26Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7561 • 3 specs; 20º42′ S, 40º06′ W; 27 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7565 • 5 specs; 20º38′ S, 40º08′ W; 26 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7566 • 1 spec.; 21º32′ S, 40º25′ W; 28 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7634 • 1 spec.; 21º40′ S, 40º02′ W; 28 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 7684 • 4 specs; REVIZEE-Central C1-VV17; IBUFRJ 9643 • 1 spec.; REVIZEE-Central C1-VV22; IBUFRJ 11368 • 1 spec.; REVIZEE-Central C2- 35R; IBUFRJ 12895 • 3], IBUFRJ 12896; 9], IBUFRJ 19489: REVIZEE-Central C1-VV16 • 3 specs; 21º15′ S, 40º20′ W; 28 Aug. 1979; OC. Ship Almirante Câmara leg.; IBUFRJ 19512 • 17 specs; REVIZEE-Central C1-D1-2; IBUFRJ 19528 • 4 specs; REVIZEE-Central C1-VV21; IBUFRJ 19541 • 3 specs; REVIZEE-Central C1-D3; IBUFRJ 19554 • 15 specs; REVIZEE-Central C1-D3; IBUFRJ 19588 • 1 spec.; REVIZEE-Central C1-VV22; IBUFRJ 19570 • 1 spec.; HAB 13-I4; MNRJ 17944 * • 1 spec.; HAB 13-H4; MNRJ 17948 * • 2 specs; HAB 11-G3; MNRJ 17949 * • 1 spec.; HAB 11-G4; MNRJ 17950 * • 4 specs; HAB 13-I2; MNRJ 17955 * • 15 specs; HAB 13-H2; MNRJ 17967 * • 2 specs; HAB 13-H3; MNRJ 17969 * • 1 spec.; HAB 11-D3; MNRJ 17972 * • 2 specs; HAB 11-C3; MNRJ 17974 * • 4 specs; HAB 11-B5; MNRJ 17976 * • 1 spec.; HAB 11-B4; MNRJ 17977 * • 9 specs; HAB 13-H1; MNRJ 17982 * • 2 specs; HAB 16-C4; MNRJ 18376 * • 6 specs; HAB 16-C3; MNRJ 18380 * • 1 spec.; HAB 16-G4; MNRJ 18396 * • 15 specs; HAB 17-I2; MNRJ 18411 * • 2 specs; HAB 16-H4; MNRJ 18413 * • 3 specs; HAB 16-H3; MNRJ 18416 * • 4 specs; HAB 16-G3; MNRJ 18418 * • 8 specs; HAB 16-H2; MNRJ 18433 * • 1 spec.; 22º42′ S, 40º40′ W; depth 5–10 m; 14 Apr. 2005; MNRJ 32405 * • 4 specs; 22º42′ S, 40º40′ W; depth 12 m; Oct. 2005; MNRJ 32618 * • 22 specs; 22º42′ S, 40º40′ W; depth 110 m; 2006; MNRJ 18751 * • 3 specs; 22º42′ S, 40º40′ W; depth 5–10 m; Mar. 2007; MNRJ 15402 * • 4 specs; 22º42′ S, 40º40′ W; depth 5–10 m; MNRJ 18953 * • 3 specs; 23º04′ S, 40º59′ W; 17 Dec. 2004; MNRJ 18969 * • 65 specs; 22º42′ S, 40º40′ W; 2007; MNRJ 29763 * • 2 specs; 23º04′ S, 40º59′ W; 1 Apr. 2005; MNRJ 31111 * • 1 spec.; 23º03′18″ S, 41º02′06″ W; Oct. 2008; MNRJ 32063 * • 2 specs; 23º15′00″ S, 41º06′00″ W; Oct. 2008; MNRJ 32064 * • 1 spec.; 23º04′ S, 40º59′ W; Jul. 2005; MNRJ 32353 * • 3 specs; REVIZEE-Central C1-VV22; MORG 40425 • 1 spec.; REVIZEE-Central C1-VV22; MORG 52260 • 1 spec.; REVIZEE-Central C1-VV21; MORG 52201 . – Almirante Saldanha Seamount • 3 specs; REVIZEE-Central C1-D1; MNRJ 33760. – Rio de Janeiro • 11 specs; 21º49′04″ S, 41º00′04″ W, São João da Barra, Porto do Açu; Jul. 2012; MZSP 111122 • 4 specs; 21º49′04″ S, 41º00′04″ W, São João da Barra, Porto do Açu; Jul. 2012; MZSP 111181 • 7 specs; Búzios, Praia da Ferradura; MNRJ 18602 * • 1 spec.; Cabo Frio; 17 Feb. 1970; J. Colella leg.; MZSP 57208 • 1 spec.; Arraial do Cabo; depth 25–30 m; Aug. 2003; P. Gonçalves leg.; MZSP 39606 • 1 spec.; Ilha do Calombo, Angra dos Reis; RAP-BIG st. 26; UERJ 5394 • 1 spec.; Ilha de Búzios, Angra dos Reis; RAP-BIG st. 23; UERJ 5452 • 1 spec.; Ilha de Búzios, Angra dos Reis; RAP-BIG st. 23; UERJ 7567 • 1 spec.; RAP-BIG st. 22; UERJ 7557 • 1 spec.; Ilha Grande, Enseada de Lopes Mendes; RAP-BIG st. 34; UERJ 5297 • 1 spec.; Ilha Grande, Enseada de Lopes Mendes; RAP-BIG st. 34; UERJ 7547. – São Paulo • 1 spec.; Ubatuba; MZSP 78290 • 1 spec.; São Sebastião, Praia do Guaecá; 23º49′14″ S, 45º27′45″ W; 9 Aug. 2011; MZSP 118845 • 1 spec.; PADCT st. 6571; MZSP 133490 • 2 specs; PADCT st. 6577; MZSP 133494 • 5 specs; REVIZEE-Sul st. 6657; MNRJ 60193 . – Santa Catarina • 2 specs; Bombinhas, Praia da Sepultura; 27º08′24″ S, 48º28′44″ W; depth 0–5 m; 17 Dec. 2019; M.R. Fernandes and A.D. Pimenta leg.; MNRJ 23366 .

COLOMBIA • 1 spec.; Cartagena, Isla Baru, Playa Blanca; 10º10′38″ N, 75º39′06″ W; 21 Mar. 2011; MNRJ 18629 *.

UNITED STATES OF AMERICA – North Carolina • 6 specs; 34º11′00″ N, 76º10′30″ W; depth 95 m; 19 Oct. 1885; R/V Albatross leg.; USNM 1438850 • 3 specs; 33º32′18″ N, 77º25′00″ W; depth 30 m; 21 Oct. 1981; with soft part preservation; USNM 850622 • 2 specs; 33º32′18″ N, 77º25′00″ W; depth 30 m; 21 Oct. 1981; with soft part preservation; USNM 850624 . – South Carolina • 1 spec.; 32º48′36″ N, 78º39′36″ W; depth 34 m; 6 May 1981; Oregon R/V leg.; with soft part preservation; USNM 850958 . – Georgia • 2 specs; 31º41′06″ N, 80º20′48″ W; depth 27 m; 14 May 1981; Bagby R/V leg.; with soft part preservation; USNM 850621. – Florida • 3 specs; off Boynton Beach; depth 27–55 m; Sep. 1967; BMSM 67476 • 1 spec.; Brower County, Deerfield Beach; depth 3–4 m; Jul. 1998; P. Souza leg.; MZSP 41482 .

Remarks

The names Cerithium pulchellum C.B. Adams, 1850 or Triphora pulchella were used by many authors (e.g., Abbott 1974; Rios 2009). It is a junior homonym of Cerithium pulchellum Sowerby, 1832, a senior synonym of the cerithiopsid Horologica pupa (Dall & Simpson, 1901) (De Jong & Coomans 1988; Rolán & Espinosa 1992) and its type material is lost (Clench & Turner 1950). Thus, C. pulchellum C.B. Adams, 1850 is not a synonym of S. intermedia, as suggested by Rolán & Fernández-Garcés (2007, 2008), but T. pulchella auct. non C.B. Adams, 1850 is indeed a synonym.

Another problematic name is Triphoris pulchellus A. Adams, 1854, which Albano et al. (2019: 171) wrongly indicated was regarded as a synonym of Similiphora intermedia by Rolán & Fernández-Garcés (2008); actually, Rolán & Fernández-Garcés (2008) referred to the species of C.B. Adams (1850), not A. Adams (1854). Because the single type of Triphoris pulchellus A. Adams, 1854 has a broken apex (without protoconch, preventing a precise identification) and the type locality is unknown, it must be regarded as a nomen dubium (but not regarded as homonym of the cerithiopsid Cerithium pulchellum C.B. Adams, 1850). If this is indeed a Caribbean shell, the most similar species is Triphora atlantica Smith, 1890, although apparently without the spiral microsculpture observed in T. pulchellus A. Adams, 1854 (Albano et al. 2019: fig. 7h).