Key to the Colombian species of Triplocania

1. Males............................................................................................... 2

- Females............................................................................................ 35

2. Forewing M3 branched (Figs 305, 317, 323, 329, 335, 341, 347, 353 and 359)..................................... 3

- Forewing M3 simple (Figs 3, 9, 75, etc.)................................................................... 8

3. Hypandrium with only the central sclerite, this with short caudo-lateral lobular processes and one central narrow elongate process having two small apical lobes (Fig. 326); phallosome as in figure 328..................... T. lamensuraensis n. sp.

- Hypandrium with two to four sclerites (Figs 308, 338, 365, 365 and 367); central sclerite of hypandrium and phallosome variable (Figs 310, 371 and 340)............................................................................. 4

4. Hypandrium of three sclerites, central sclerite with two pairs of posterior, acuminate processes, and a posterior median, elongate, acuminate process (Figs 308 and 365)................................................................. 5

- Hypandrium of two or four sclerites, the anterior-central sclerite with a pair of long acuminate, horn-shaped processes, strongly curved distally (Figs 338, 366, and 367); posterior sclerite with an acuminate or spatulate process............... 6

5. Central sclerite of hypandrium with median, posterior, long acuminate process, almost as long as the lateral ones (Fig. 365); phallosome with phallobase almost straight proximally, posterior endophallic sclerite sinuous distally, with strongly curved tapered distal process (Fig. 370)............................................................... T. furcata New

- Central sclerite of hypandrium with median, posterior, stout process, longer than the lateral ones (Fig. 308); phallobase curved proximally toward the mesal line; posterior endophallic sclerites almost straight distally, with curved tapered apical process and two curved anteapical small teeth (Fig. 310).............................................. T. furcatoides n. sp.

6. Hypandrium of two sclerites, acuminate processes of anterior sclerite each with a basal tooth; posterior sclerite with spatulate posterior process, dilated distally (Fig. 338); phallosome with side struts independent, each widened basally (Fig. 340)........................................................................................ T. leguizamoensis n. sp.

- Hypandrium of four sclerites, spines of anterior sclerite without basal tooth (Fig. 366 and 367); posterior sclerite with a short, stout tapered posterior process........................................................................... 7

7 Posterior sclerite of hypandrium with posterior process stout, about as long as the antero-posterior length of the anterior sclerite; sickle-shaped lateral processes of the anterior sclerite long, extending beyond the side sclerites (Fig. 366)............................................................................................. T. lamasi Silva Neto et al.

- Posterior sclerite of hypandrium slender, with posterior process slender, much longer than the antero-posterior length of the anterior sclerite (Fig. 367); sickle-shaped lateral processes of the anterior sclerite short, not extending beyond the side sclerites............................................................................ T. lamasoides Silva Neto et al.

8. Central sclerite of hypandrium asymmetric, with apex directed dorsally (Figs 79, 138, 180, 222); forewings with a brown marginal band, from R4+5 to end of CuP; anterior endophallic sclerite of three arms or processes, one central................ 9

- Central sclerite of hypandrium symmetric (Figs 7, 13, 18), forewings hyaline or with brown marginal band; anterior endophallic sclerites paired, usually with two central processes........................................................ 12

9. Phallosome with antero-mesal endophallic sclerites projected caudally to about the level of the apex of the external parameres (Figs 80 and 139); pair of posterior endophallic sclerites with arms not distally overlapping.......................... 10

- Phallosome with short antero-mesal endophallic sclerites, not projected caudally to the level of the apex of the external parameres (Figs 181 and 223); pair of posterior endophallic sclerites asymmetrical, each arm anteriorly curved and distally overlapping......................................................................................... 11

10. Central sclerite of hypandrium with a stout, median posterior projection, arrow-shaped distally, slanted to the left (Fig. 1 38). Endophallic sclerites slender, mesal sclerite smooth, narrowing distally, a round asymmetric sclerite anteriorly on the left, bearing a row of teeth along anterior border (Fig. 139)......................................... T. felidiaensis n. sp.

- Central sclerite of hypandrium with an acute, median posterior projection, slanted to the right (Fig. 79). Anterior pair of endophallic sclerites distally narrow, acuminate, with arms long and slender; mesal sclerite dilated proximally, narrowing distally, acuminate, bearing a row of short straight spines along side, bending to the left (Fig. 80)............. T. calima n. sp.

11. Posterior pair of endophallic sclerites curved and acuminate distally; mesal sclerite with median projection stout, distally forked, with both arms acuminate, one straight and the other curved, bearing an anteapical denticle on the inner border (Fig. 223)............................................................................. T. mariacarmenae n. sp.

Posterior pair of endophallic sclerites with right one distally curved, acuminate; left one distally curved, apically dilated, with a row of teeth on outer border; mesal pair with arms slender, and a stout median projection bearing distally a row of teeth (Fig. 181)............................................................................... T. humboldtiana n. sp.

12. Central sclerite of hypandrium with a deep, U-shaped posterior concavity; postero-lateral processes elongate and tapered, widely separated; posterior endophallic sclerite tapered and curved outwards, mesal sclerite transverse (Figs 18 and 119).. 13

- Central sclerite of hypandrium variable (Figs 7, 43, 48, 61, 91, 96, 114, 150, 168, 192, 210, 234, 251, 257, 291, and 368), if Ushaped posterior concavity, then lateral processes not elongated and tapered as the previous one; endophallic sclerites variable................................................................................................... 14

13. Central sclerite of hypandrium with short acuminate teeth proximally on the inner side (Fig. 18); posterior endophallic sclerite strongly enlarged basally and with short process outwards; mesal sclerite transverse (Fig. 20)............ T. andaqui n. sp.

- Central sclerite of hypandrium with two teeth or mesal processes (Fig. 119); posterior endophallic sclerite enlarged basally but not as the previous one (Fig. 121)............................................................. T. dimitrii n. sp.

14. Vein A2 not reaching the wing margin.................................................................... 15

- Vein A2 reaching the wing margin....................................................................... 30

15. Central sclerite of hypandrium with caudo-lateral and central projections (Figs 7, 96, 150, 257 and 291)................ 16

- Central sclerite of hypandrium with caudo-lateral projections only, widely separated or close and centered, or reduced to a single mesal lobe (Figs 43, 48, 61, 91, 114, 168, 192, 210, 234, 251 and 368)........................................ 20

16. Central sclerite of hypandrium on inner margin of lateral processes and between these and median lobes with short sclerotized teeth; median lobes wide, occupying more than 1/3 of the posterior margin, distally rounded and directed laterally (Fig. 257); phallosome with mesal endophallic sclerite stout, X-shaped, anterior sclerites with inner arms straight, slender, acuminate, and outer arms stout, bow shaped (Fig. 258).................................................... T. pericosensis n. sp.

- Central sclerite of hypandrium without teeth as described above, median lobes not as wide; phallosome with mesal sclerite Wshaped (Fig. 96) or as two separate sclerites (Fig. 8)......................................................... 17

17. External parameres distally rounded, directed posteriorly; mesal sclerites laminar, proximally wide, narrowing distally, with apical teeth; anterior sclerites with inner arms globose, outer arms curved, distally blunt (Figs. 8)... T. amacayacuensis n. sp.

- External parameres not as above; mesal sclerite W-shaped (Fig. 96)............................................. 18

18. Central sclerite of hypandrium with well developed postero-lateral processes (Figs 150, 291)........................ 19

- Central sclerite of hypandrium with postero-lateral processes small or not evident (Fig. 96); endophallic sclerites as in figure 98..................................................................................... T. cantatis n. sp.

19. Central sclerite of hypandrium with inwards curved postero-lateral processes, with stout, deeply obtusely concave median Ushaped process (Fig. 291); anterior endophallic sclerites with inner arms stout, straight, outer arms pedunculate, distally dilated, papillose (Fig. 292)................................................................ T. yanacona n. sp.

- Central sclerite of hypandrium with outwardly directed postero-lateral processes, with stout, deeply obtusely concave median process, with setose lobes (Fig. 150); phallosome without anterior endophallic sclerite projected laterally and enlarged distal area (Fig.151)..................................................................... T. garciamarquezi n. sp.

20. Forewings with broad marginal band from R2+3 to Cu1a (Fig. 206). Postero-lateral processes of central sclerite of hypandrium distally bulged (Fig. 210); anterior endophallic sclerite with arms L-shaped, the longer stem projected posteriorly, ending in a short process and bearing a large subapical tooth on inner border; mesal pair with outer arm curved, distally acuminate, outer border with a row of small pointed processes; inner arm straight, distally acuminate; posterior pair with arms curved, elongate, distally acuminate (Fig. 211)............................................................... T. lithophila n. sp.

- Forewings without marginal pigmented band, or, if present, poorly defined (Fig. 39); postero-lateral processes of central sclerite of hypandrium variable (Figs 43, 48, 61, 91, 114, 168, 192, 234, 251 and 368); mesal endophallic sclerite, H, W, X, or Yshaped............................................................................................. 21

21. Central sclerite of hypandrium elongate, with outer subparallel borders that continue in two processes or elongate lateral lobes widely separated or relatively close (Figs 61 and 168)........................................................ 22

- Central sclerite of hypandrium short and narrow or wider than long, with rounded or irregular outer borders (Figs 43, 48, 91, 114, 192, 234, 251 and 368)............................................................................ 23

22. Central sclerite of hypandrium with sides almost parallel, lateral processes widely separated, rounded, each bearing distally two lateral acuminate processes, and a small acuminate process on inner border (Fig. 61); endophallic sclerites as in figure 62...................................................................................... T. bicornuta n. sp.

- Central sclerite of hypandrium with sides convergent, lateral processes blunt ended; endophallic sclerites as in figure 169............................................................................................ T. huitota n. sp.

23. Hypandrium of one sclerite, wider than long, with a slender, sclerotized process in the middle (Fig. 48); anterior endophallic sclerite transverse, sausage-shaped, with a row of short denticles along posterior border; mesal sclerite broadly H-shaped, with anterior arms stout; posterior pair with each arm of two pieces, the anterior one wide, anteriorly acuminate, the posterior one elongate, curved, distally wider, with denticles along posterior border (Fig. 50)............................ T. awa n. sp.

- Hypandrium of one or three sclerites; central sclerite with double processes, central or lateral (Figs 43, 91, 114, 192, 234, 251 and 368); endophallic sclerites not as above................................................................ 24

24. Central sclerite of hypandrium with central short or elongate lobular projections (Figs 43, 114, 192 and 368)............ 25

- Central sclerite of hypandrium with only lobed lateral projections (Figs 91, 234 and 251)............................ 28

25. Central sclerite of hypandrium with long posterior process in the middle, deeply obtusely concave distally (Fig. 43); anterior endophallic sclerite small, transverse, close together in the middle, mesal pair with each arm T-shaped, with a median stout process flanked by small slender processes formed by two lateral sclerites; posterior pair proximally slender, acuminate, wider in the middle, distally curved inward, with the border denticulate (Fig. 44)........................... T. asisensis n. sp.

- Central sclerite of hypandrium with short lobular processes, originated in a mesal lobe wider than long (Figs 114, 192 and 368); endophallic sclerites not as above (Figs 115, 193 and 371)................................................ 26

26. Central sclerite of hypandrium with two small median processes, almost closing a round concavity, sclerotized along its border (Fig. 114); anterior endophallic sclerites M-shaped, stout; mesal sclerite broadly X-shaped, underlaid by elliptic round areas; posterior pair slender, curved outwards, bearing denticles on outer border (Fig. 115).................. T. chocoensis n. sp.

- Central sclerite of hypandrium uniformly sclerotized, wider than long, with short lobular projections, originating in a mesal lobe wider than long (Figs 192 and 368)................................................................... 27

27. Central sclerite of hypandrium with a median, bilobed posterior process, postero-lateral corners not projected (Fig. 192); posterior endophallic sclerites of two symmetrical T-shaped arms (Fig. 193).............................. T. kichwa n. sp.

- Central sclerite of hypandrium with two small, median posterior processes, postero-lateral corners projected (Fig. 368); posterior endophallic sclerites triangular (Fig. 371)........................................... T. erwini Silva-Neto et al.

28. Central sclerite of hypandrium U-shaped, stout (Fig. 251); mesal endophallic sclerite wide, H-shaped (Fig. 252)................................................................................................... T. panchei n. sp.

- Central sclerite of hypandrium not as above (Figs 91 and 234); mesal endophallic sclerite wide, H or sea star-shaped (Figs 92 and 235)............................................................................................ 29

29. Central sclerite of hypandrium with two elongate postero-lateral lobes, slightly curved inwards, blunt ended (Fig. 234); mesal endophallic sclerite broadly H-shaped, posterior arms distally denticulate; anterior pair robust, U-shaped (Fig. 235).............................................................................................. T. mocoaensis n. sp.

- Central sclerite of hypandrium with two stout, distally truncate postero-lateral lobes (Fig. 91); mesal endophallic sclerite starshaped (Fig. 92)......................................................................... T. camentsa n. sp.

30. Central sclerite of hypandrium with central and lateral processes (Figs 73 and 269)................................ 31

- Central sclerite of hypandrium with lateral or central processes (Figs 13, 31, 132 and 263)........................... 32

31. Central sclerite of hypandrium with a median, distally bilobed posterior projection, and a broad process on each postero lateral corner, each bearing a rounded lateral projection, with a row of short, sclerotized teeth along the posterior border (Fig. 269); mesal pair of endophallic sclerites with central, fused area arrow shaped, posteriorly bifid, anteriorly with two arms, each one straight in the middle, then bow-shaped, with outer border irregular distally bearing denticles along both sides, posterior endophallic sclerites transverse, slender, with inner ends pointed and outer ends bifid, acuminate (Fig. 270)....................................................................................................... T. rugosa n. sp.

- Central sclerite of hypandrium anteriorly straight, with postero-lateral corners sclerotized, projected posteriorly; posterior border with two short projections in the middle, apically blunt, directed outwards, the space between the middle and the lateral projections denticulate, strongly sclerotized (Fig. 73); anterior pair of endophallic sclerites broadly M-shaped, mesal pair Xshaped, with posterior arms curved, acuminate; posterior pair close to the inner border of the external parameres, each half elongate, with a row of four teeth along the inner border (Fig. 74)................................... T. bubuae n. sp.

32. Central sclerite of hypandrium with latero-caudal processes separated by a deep or shallow concavity (Figs 13 and 132); posterior endophallic sclerites H-shaped (Figs 14 and 133)........................................................ 33

- Central sclerite of hypandrium with median processes, small or well developed (Figs 31, 263); mesal endophallic sclerites Y or H-shaped (Figs 32 and 264)............................................................................ 34

33. Central sclerite of hypandrium with two short, stout posterior projections, with a deep concavity between them (Fig. 132); two pairs of endophallic sclerites, anterior pair with inner arms slender, acuminate; outer arms club-shaped; mesal pair X-shaped, with posterior arms distally rounded, with an acuminate lateral projection (Fig. 133).................... T. embera n. sp.

- Central sclerite of hypandrium with two short, blunt ended posterior projections, with a narrow concavity between them (Fig. 13); three pairs of endophallic sclerites, mesal pair stout, H-shaped, anterior pair with inner arms straight, slender, acuminate, outer arms curved, stout (Fig. 14)....................................................... T. anchicayaensis n. sp.

34. Forewing M1 with Y-shaped spur vein (Fig. 27); central sclerite of hypandrium oval, with a small, concave posterior projection in the middle (Fig. 31); anterior pair of endophallic sclerites with fan-shaped processes, postero-mesal pair dilated posteriorly, fork-shaped (Fig. 32)...................................................................... T. arhuaca n. sp.

- Forewing M1 without spur vein (Fig. 259); central sclerite of hypandrium convex anteriorly, bilobed posteriorly, with apex widely rounded and closely spaced (Fig. 263); mesal endophallic sclerite transverse, H-shaped, with two long anterior arms, posterior arms straight, acuminate, and close together (Fig. 264)................................. T. robustoides n. sp.

35. Forewing M3 simple.................................................................................. 36

- Forewing M3 branched, bi- or trifurcated (Figs 305, 311, 317, 323)............................................. 60

36. Forewings with dark brown pigmentation proximally, sometimes with a wide, marginal pigmented band (Figs 194 and 200); A2 reaching the wing margin; IX sternum convex, emarginate, with clearly defined antero-lateral lobe (Figs 198 and 205)...................................................................................................... 37

- Forewings hyaline, or slightly pigmented proximally, sometimes with wide marginal pigmented band; A2 reaching or not the wing margin; IX sternum variable........................................................................ 38

37. Forewings without marginal pigmented band, deeply pigmented proximally (Fig. 200); labral sclerites evident (Fig. 202); ninth sternum concave anteriorly in the middle (Fig. 205).......................................... T. lapayaensis n. sp.

- Forewing with broad marginal pigmented band from R4+5 to areola postica, with large dark brown proximal area, pterostigma with proximal and distal brown bands; A2 joining A1 (Fig. 194); labral sclerites not evident (Fig. 196); IX sternum trapeziform, peripherally pigmented (Fig. 198)....................................................... T. korebaju n. sp.

38. Forewings with well defined pigmented marginal band, at least between R4+5 and Cu1b (Figs 158, 224, and 281)........ 3 9

- Forewings hyaline or with pigmented marginal band not well defined, if it continues, then with hyaline or clear spaces near margin of the respective cells (Figs 21, 33,122, 182, 242, and 277).............................................. 44

39. Forewings with pigmented marginal band without setae; areola postica with ratio length /width greater than 1.5, with strong concavity on the part distal to Cu1a (Fig. 151); IX sternum with two well defined areas, the anterior widened, with lateral lobes and leading edge convex, emarginate (Fig. 162)............................................................. 40

- Forewings with pigmented marginal band with setae; areola postica with ratio length/width smaller than 1.5; IX sternum variable............................................................................................... 42

40. Forewing with A2 reaching the wing margin (Figs 224 and 281); IX sternum with anterior area distinctly delimited (Figs 228 and 286), with oval laminar structure in central area........................................................ 41

- Forewing with A2 not reaching the wing margin (Fig. 158); IX sternum slender posteriorly, with anterior, transverse, pigmented narrow band, without central laminar structure (Fig. 162)..................................... T. guane n. sp.

41. Ninth sternum with three well delimited areas, a pigmented transverse anterior one, a mesal area projected posteriorly in the middle, with one transverse sickle-shaped pigmented area on each side of the longitudinal midline, and one mostly hyaline posterior area (Fig. 286)................................................................. T. sevillaensis n. sp.

- IX sternum broader posteriorly, with the anterior, transverse pigmented band wider (Fig. 228), posterior area distinctly wide, trapezoida.............................................................................. T. matildae n. sp.

42. Forewings with transverse pale brown band from rs-m to apex of cell cup; veins M1 to Cu1b ending in a hyaline fenestra, with dark sides (Fig. 212); IX sternum with a well-defined, anterior transverse area and two posterior lobes, anterior margin distinctly emarginated (Fig. 217).............................................................. T. lithophila n. sp.

- Forewings without transverse pale brown band from rs-m to apex of cell cup, longitudinal veins with brown areolae at setal insertions (Figs 81 and 140); IX sternum variable (Figs 86 and 145)............................................. 43

43. IX sternum with three well defined areas, the anterior one oval; mesal area with an anterior V-shaped concavity, and a posterior, trapeziform pigmented process; posterior area membranous, unpigmented (Fig. 86)................. T. calima n. sp.

- IX sternum trapeziform, with a distinct, pigmented transverse band, projected posteriorly in the middle (Fig. 145)............................................................................................... T. felidiaensis n. sp.

44. Forewings with vein A2 not reaching the wing margin....................................................... 45

- Forewings with vein A2 reaching the wing margin.......................................................... 57

45. Forewings hyaline, membrane with spots at the ends of the veins, with or without short spur veins on margin, on cells m2 and m3 (Figs 21 and 122); IX sternum with widened anterior area, rounded edges and front side emarginated (Figs 26 and 127)..................................................................................................... 46

- Forewings hyaline or with pigmented areas, without spur veins on margin (Figs 33, 182, 242, and 277); IX sternum with anterior area variable..................................................................................... 47

46. Forewings with short spur veins on margin, on cells m2 and m3 (Fig. 21); IX sternum without sclerotized or pigmented area in anterior area (Fig. 26)..................................................................... T. andaqui n. sp.

- Forewings without spur veins as above (Fig. 122); IX sternum with two sclerotized areas in the posterior margin of the anterior area (Fig. 127)............................................................................ T. dimitrii n. sp.

47. Forewings with a transverse dark or pale brown band from rs-m to apex of cell cup (Figs 63 and 370); distal part of Cu1a with or without distinct emargination......................................................................... 48

- Forewings without a transverse dark brown band between rs-m to apex of the cell cup (Figs 182, 277); distal part of areola postica almost straight, sometimes emarginated................................................................ 49 48. IX sternum broadly pentagonal (Fig. 68); Cula concave in the middle (Fig. 63)....................... T. bicornuta n. sp.

- IX sternum oval (Fig. 373); Cu1a almost straight (Fig. 370).............................. T. colombiana García Aldrete

49. IX sternum trapeziform, with convex anterior margin (Figs 186 and 280)........................................ 50

- IX sternum cup-shaped, with anterior margin convex or straight (Figs 55, 103, 109, 157, 175, 298 and 303); if trapeziform, then anterior margin fully emarginate..................................................................... 51

50. IX sternum broadly trapeziform, convex anteriorly; posteriorly with a triangular, sclerotized area on each side of longitudinal midline (Fig. 280); forewings hyaline, with a submarginal pale brown band from R2+3 to M3; Cu1a straight (Fig. 277)............................................................................................. T. sarmaca n. sp.

- IX sternum trapeziform, with pigmented band proximally in the middle (Fig. 186); forewings hyaline, with brown pigmented areas as in figure 182......................................................................... T. inga n. sp.

51. IX sternum quadrangular, with oval anterior area and cup-shaped posterior area (Fig. 103); Cu1a concave in the middle (Fig. 99)..................................................................................... T. cantatis n. sp.

- IX sternum subquadrangular, without oval anterior area (Figs 55, 109, 157, 298, and 303); Cu1a almost straight or gently concave (Figs 45, 152, 164, 293, and 299).................................................................... 52

52. IX sternum quadrangular: anterior area projected posteriorly in the middle (Fig. 298); subgenital plate with well defined pigmented area (Fig. 297)................................................................... T. yanacona n. sp.

- IX sternum not as above; subgenital plate with pigmented area variable.......................................... 53

53. IX sternum cup-shaped, pigmented, anterior margin slightly convex, almost straight, with ventro-anterior V-shaped pigmented area (Fig. 55), posteriorly rounded, with two rounded processes........................................ T. awa n. sp.

- IX sternum cup or trapeziform-shaped, without sclerotized areas as above........................................ 54

54. IX sternum cup-shaped, anteriorly wide; anterior and posterior margins almost straight, with two sclerotized large sublateral areas distally (Fig. 157).............................................................. T. garciamarquezi n. sp.

- IX sternum cup-shaped or trapeziform, with anterior margin distinctly emarginate, or without sclerotized areas as above... 55

55. IX sternum trapeziform, wider anteriorly, slightly concave in the middle, narrower posteriorly, deeply concave in the middle (Fig. 109); a mesal, transverse, bow-shaped pigmented band, and one small pigmented area posteriorly, on each side of the longitudinal midline........................................................................... T. caribe n. sp.

- IX sternum trapeziform, anterior margin clearly emarginated (Figs 175 and 303)................................... 56

56. IX sternum cup-shaped, concave anteriorly, projected posteriorly, with membranous areas on sides (Fig. 175)..................................................................................................... T. huitota n. sp.

- IX sternum trapeziform, anterior margin widely emarginated; posteriorly without lateral concavity or oval areas as in T. huitota (Fig. 303)........................................................................ T. erwini Silva-Neto et al.

57. Forewing M1 with a Y-shaped spur vein (Fig. 33); IX sternum broadly trapeziform, with a transverse, pigmented band anteriorly, and an irregular pigmented area on each postero-lateral corner (Fig. 38)......................... T. arhuaca n. sp.

- Forewing M1 without spur vein (Figs 236, 242 and 271); IX sternum variable (Figs 241, 246, and 276)................ 58

58. Forewings with slender, pale brown marginal band; areola postica slanted posteriorly (Fig. 242); IX sternum wider anteriorly, concave on the sides, and anteriorly and posteriorly; a transverse pigmented band along anterior border (Fig. 246)................................................................................................. T. motilona n. sp.

- Forewings not as above, areola postica not slanted (Figs. 236 and 271); IX sternum variable (Figs 241 and 276).......... 59

59. IX sternum long, trapeziform, wider anteriorly, slightly concave in the middle (Fig. 276)................. T. rugosa n. sp.

- IX sternum cup-shaped, anterior margin slightly emarginated (Fig. 241)........................... T. mocoaensis n. sp.

60. Forewings hyaline, without dark marginal band (Figs 335, 353 and 359); IX sternum of two large areas, anterior one with lateral margins oval, anterior margin emarginate centrally, with mesal, narrow and elongate concavity projected mesally; epiproct with oval bulge in mesal area, sometimes not evident in dorsal view (Figs 316, 358 and 364)......................... 61

-..... Forewings with pale brown area proximally, with dark marginal band from R2+3 to Cu1a (Figs 317, 329, 341 and 347); pterostigma brown, with cross pale brown spot from rs-m to anal margin; IX sternum not as above; epiproct without oval bulge in mesal area (Figs 321, 333, 344 and 350)................................................................ 64

61. IX sternum posteriorly without longitudinal grooves or clearly delimited by lateral sclerotized area, posterior border emarginate (Figs 364 and 373)................................................................................. 62

- IX sternum posteriorly with longitudinal grooves, posterior border rounded, not emarginate (Figs 316 and 358).......... 63

62. IX sternum with posterior area pigmented, thicker than the anterior one, with a strongly sclerotized latero-posterior band, and a small, strongly pigmented oval area mesally on each side (Fig. 373)..................... T. lamasoides Silva-Neto et al.

- IX sternum with posterior area not pigmented, without a strongly sclerotized latero-posterior band, slightly pigmented or without pigmented oval area mesally on each side (Fig. 364)................................... T. lamasi Silva-Neto et al.

63. IX sternum posteriorly with slightly developed longitudinal grooves mesally (Fig. 358), V2 +3 proximally wide, V3 with widely rounded apex........................................................................ T. furcata New

- IX sternum posteriorly with many longitudinal grooves, elongated mesally, more than in T. furcata (Fig. 316); v2 +3 proximally not as wide as above, v2 with apex less rounded.............................................. T. furcatoides n. sp.

64. A2 not reaching the wing margin, IX sternum with elongate lateral and sclerotized brown area, lateral borders well sclerotized (Figs 322 and 334).................................................................................... 65

- A2 reaching the wing margin; IX sternum without sclerotized lateral area, lateral borders well sclerotized, irregularly oval or cup-shaped (Figs 346 and 352).......................................................................... 66

65. IX sternum with semi-oval anterior area, delimited by latero-posterior brown slender bands; posteriorly emarginate (Fig. 322); fore- coxa pale brown, with brown ochre spot, middle and posterior coxae cream..................... T. huilaensis n. sp.

- IX sternum with trapeziform; anterior area delimited by latero-posterior brown bands; anterior border emarginate (Fig. 334); fore- and mid coxae dark brown, hind- coxae cream....................................... T. lamensuraensis n. sp.

66. IX sternum shield-shaped, wider anteriorly, with lateral borders limited by a sclerotized band, anterior border gently emarginated; latero-posteriorly with two oval, projected membranous areas (Fig. 346)................ T. otunquimbayaensis n. sp.

- IX sternum broadly oval, lateral borders limited by a sclerotized band, anterior border projected in the middle, posterior border with corners protruding (Fig. 352)............................................................ T. sarriae n. sp.