Pusillina cf. radiata (Philippi, 1836)

(Fig. 21a–c)

Material examined. (23 empty shells and 2 with soft parts, in 8 samples): SPAIN • 1 sPc (juv); 43° 35.451′N, 008° 34.432′W to 43° 34.810′N, 008° 35.407′W; 153– 151 m; 08–15 SeP. 2002; DIVA-Artabria I EBS-150 • 2 sh; 43° 48.421′N, 008° 51.453′W to 43° 49.160′N, 008° 51.091′W; 599–607 m; 10–20 SeP. 2003; DIVA-Artabria I DRN-600 • 1 sPc (juv); 43° 34.127′N, 008° 36.562′W to 43° 34.820′N, 008° 35.585′W; 152– 149 m; 10–20 SeP. 2003; DIVA-Artabria I EBS-150 • 6 sh; 43° 38.812′N, 009° 07.949′W to 43° 39.841′N, 009° 07.405′W; 999–1001 m; 17–28 Sep. 2004; VERTIDOS GA-DRN-1000 • 6 frg; 43° 38.105′N, 009° 02.006′W to 43° 37.738′N, 009° 00.905′W; 836– 826 m; 17–28 SeP. 2004; VERTIDOS GA-AT-800 • 2 sh; Polygon delimited by Points 44° 10.000′ N, 009° 00.000′ W /44° 10.000′ N, 008° 35.000′ W/44° 00.000′ N; 009° 00.000′ W/44° 07.000′ N; 008° 35.000′ W; 480–600 m; SARRIDAL (2006–2007) SARRI-2 • 6 sh; 44° 11.652′N, 008° 58.152′W to 44° 11.539′N, 008° 57.574′W; 908–1106 m; 15–24 Jul. 2008; A SELVA DRN-7 • 1 sh; 44° 08.461′N, 009° 04.634′W to 44° 08.607′N, 009° 04.413′W; 917– 896 m; 15–24 Jul. 2008; A SELVA DRN-7-1 .

Remarks. It is a Mediterranean species with isolated records in the Atlantic coasts (Verduin 1976) usually in seagrass meadows (Gofas & Oliver 2011). Verduin (1976) pointed out the extreme variability of this species (with some local forms), whose precise delimitation is not clear. A thorough review of most species of genus Pusillina, including genetic analysis, is necessary to elucidate their taxonomic status.

The presence of some juvenile shells in our samples, attributed with doubts to P. radiata (Figs. 21a–b) is considered accidental; probably washed away by currents. Only two shells with soft parts were found in samples obtained at 150 m. The protoconch is shown in Figure 21c.