Alvania testae (Aradas & Maggiore, 1844)

(Figs. 5 e–g)

Material examined. (nearly 2000 shells, most of them with soft parts, in 33 samples): SPAIN • 1 s; 43° 40.165′N, 008° 43.697′W to 43° 40.513′N, 008° 43.159′W; 204–209 m; 08–15 SeP. 2002; DIVA-Artabria I DRN-200 • >80 sPc; 43° 35.451′N, 008° 34.432′W to 43° 34.810′N, 008° 35.407′W; 153– 151 m; 08–15 SeP. 2002; DIVA-Artabria I EBS-150 • 40 spc; 43° 40.192′N, 008° 43.760′W to 43° 40.943′N, 008° 42.366′W; 207–212 m; 08–15 SeP. 2002; DIVA-Artabria I EBS-200 • 10 sPc; 43° 41.689′N, 008° 45.195′W to 43° 42.556′N, 008° 44.226′W; 298–303 m; 08–15 SeP. 2002; DIVA-Artabria I EBS-300 • 1 spc (juv); 43° 53.575′N, 008° 56.868′W to 43° 54.015′N, 008° 56.959′W; 965–974 m; 10–20 SeP. 2003; DIVA-Artabria I DRN-1000 •> 80 sPc; 43° 34.116′N, 008° 36.535′W to 43° 34.689′N, 008° 35.589′W; 149– 148 m; 10–20 Sep. 2003; DIVA-Artabria I AT-150 •> 200 spc; 43° 34.127′N, 008° 36.562′W to 43° 34.820′N, 008° 35.585′W; 152– 149 m; 10–20 Sep. 2003; DIVA-Artabria I EBS-150 •> 50 spc; 43° 40.250′N, 008° 43.755′W to 43° 40.760′N, 008° 42.120′W; 207– 197 m; 10–20 Sep. 2003; DIVA-Artabria I EBS-200 •> 300 spc; 43° 41.590′N, 008° 45.328′W to 43° 42.396′N, 008° 44.286′W; 301– 301 m; 10–20 SeP. 2003; DIVA-Artabria I EBS-300 • 1 sh; 43° 42.348′N, 008° 45.889′W to 43° 43.269′N, 008° 45.289′W; 344–354 m; 10–20 SeP. 2003; DIVA-Artabria I EBS-350 • 30 spc + 200 sh; 43° 29.412′N, 008° 28.362′W to 43° 29.779′N, 008° 28.240′W; 110–111 m; 17–28 SeP. 2004; VERTIDOS GA-AT-110 • 50 sPc; 43° 32.026′N, 008° 37.520′W to 43° 32.806′N, 008° 35.993′W; 150–151 m; 17–28 SeP. 2004; VERTIDOS GA-AT-150 • 1 spc (juv); 43° 38.105′N, 009° 02.006′W to 43° 37.738′N, 009° 00.905′W; 836– 826 m; 17–28 SeP. 2004; VERTIDOS GA-AT-800 • 30 sPc; 42° 29.455′N, 009° 17.472′W to 42° 31.292′N, 009° 19.660′W; 149– 147 m; 17–28 SeP. 2004; VERTIDOS AG-AT-150 •>40 sPc; 42° 47.353′N, 009° 25.161′W to 42° 44.866′N, 009° 25.913′W; 147– 146 m; 17–28 SeP. 2004; VERTIDOS CA-AT-150 • 30 spc; 43° 31.970′N, 008° 37.670′W to 43° 32.697′N, 008° 3.171′W; 151– 151 m; 17–28 SeP. 2004; VERTIDOS GA-EBS-150 • 25 spc; 43° 36.222′N, 008° 52.845′W to 43° 36.797′N, 008° 52.112′W; 201– 201 m; 17–28 Sep. 2004; VERTIDOS GA-EBS-200 • >800 spc; 42° 30.391′N, 009° 19.517′W to 42° 29.428′N, 009° 17.924′W; 147–148 m; 17–28 Sep. 2004; VERTIDOS AG-EBS-150 •>50 spc; 42° 31.176′N, 009° 23.380′W to 42° 32.132′N, 009° 24.151′W; 242–248 m; 17–28 Sep. 2004; VERTIDOS AG-EBS-250 •>200 sPc; 42° 50.507′N, 009° 25.773′W to 42° 48.810′N, 009° 25.198′W; 151– 148 m; 17–28 SeP. 2004; VERTIDOS CA-EBS-150 • 30 sPc; 42° 15.780′N, 009° 10.500′W to 42° 17.100′N, 009° 10.630′W; 149–150 m; 17–28 SeP. 2004; VERTIDOS CH-EBS-150 • 8 sh; 43° 29.13′N, 008° 43.233′W to 43° 29.596′N, 008° 42.838′W; 149–151 m; 15–24 Jul. 2008; A SELVA DRN-22 • 14 spc; 43° 31.727′N, 008° 52.374′W to 43° 32.678′N, 008° 52.687′W; 173–179 m; 15–24 Jul. 2008; A SELVA DRN-23 • 12 sPc; 44° 06.496′N, 008° 23.522′W to 44° 07.16′N, 008° 23.201′W; 337– 324 m; 15–24 Jul. 2008; A SELVA AT-13 • 1 spc + 1 sh; 44° 08.516′N, 008° 40.788′W to 44° 09.688′N, 008° 39.721′W; 436– 428 m; 15–24 Jul. 2008; A SELVA EBS-11 • 14 sPc; 44° 05.687′N, 008° 24.558′W to 44° 07.253′N, 008° 24.302′W; 337– 330 m; 15–24 Jul. 2008; A SELVA EBS-13 • 15 sPc; 44° 00.806′N, 008° 33.098′W to 44° 01.967′N, 008° 32.57′W; 346– 344 m; 15–24 Jul. 2008;A SELVA EBS-14 • 17 sPc + 10 sh; 43° 58.84′N, 008° 15.625′W to 43° 59.621′N, 008° 14.285′W; 233–237 m; 15–24 Jul. 2008; A SELVA EBS-20 •> 50 sPc; 43° 31.687′N, 008° 52.188′W to 43° 33.149′N, 008° 53.142′W; 171–182 m; 15–24 Jul. 2008; A SELVA EBS-23 • 5 sPc; 43° 48.252′N, 008° 51.427′W to 43° 49.707′N, 008° 51.164′W; 575–584 m; 15–24 Jul. 2008; A SELVA EBS-30-1 • 4 sPc + 3 sh; 43° 33.20′N, 008° 58.78′W to 43° 34.57′N, 008° 56.90′W; 230–236 m; 15–30 SeP. 2008; DIVA-Artabria II EBS-01 • 1 sh; 43° 34.86′N, 009° 21.95′W to 43° 36.44′N, 009° 20.71′W; 1010–1112 m; 15–30 SeP. 2008; DIVA-Artabria II EBS-04 • 2 sPc; 43° 19.92′N, 009° 38.13′W to 43° 21.25′N, 009° 37.11′W; 1267–1286 m; 15–30 SeP. 2008; DIVA-Artabria II EBS-15 • 1 sh; 42° 31.58′N, 009° 40.14′W to 42° 33.87′N, 009° 38.75′W; 2030– 1886 m; 15–30 SeP. 2008; DIVA-Artabria II EBS-30 .

Remarks. Very common species that extends from Norway to off western Morocco and throughout the Mediterranean, in a wide bathymetric range 30–1000 m, occasionally deeper (Oliverio et al. 1992; Bouchet & Warén 1993; Warén 1996; Cachia et al. 1996; Giribet & Peñas 1997; Gofas & Oliver 2011; Negri & Corselli 2016; Bitlis & Öztürk 2017). The shell is characterized by the opisthocline orientation of the peristome (Fig. 5e). Protoconch sculptured by an irregular network (Figs. 5f–g).

It was the most abundant species in studied the samples along with Obtusella macilenta, predominantly between 100 and 400 m in muddy sand bottoms. Its putative planktrotrophic larval development (deduced from its protoconch) enables a large dispersal capacity.