PODOSCIRTINI SAUSSURE, 1878

Podoscirtites Saussure, 1878: 693, Saussure, 1893: 257.

Podoscirtinae Desutter, 1987: 234 .

Podoscirtini Chopard, 1968: 365, Otte & Alexander, 1983: 310, Otte, 1994: 72, Otte, 1994: 86, Rentz, 1996: 138, Gorochov, 2004: 187–215, Gorochov, 2010: 206, Rentz & Su, 2019: 156.

Type genus: Podoscirtus Serville, 1838 .

Distribution: Afrotropics, Australasia, Indo-Malaya, Oceania and Palaearctic.

Diagnosis: Small to large size crickets. Male: FWs generally developed bearing stridulatory apparatus, sometimes without hv or mirror (or both). Male genitalia: pseudepiphallic sclerite generally capsular (Fig. 10D; Supporting Information, Fig. S7F); apex of LLophi frequently curved upwards (Supporting Information, Fig. S7F). Female: apex of dv of ovipositor pointed, its surface smooth, presence of bristles in apex inner margin (Fig. 9C); apex of vv generally rounded.

Included genera: Abaxitrella Gorochov, 2002, Adenopterus Chopard, 1951, Anaudus Saussure, 1874, Anisotrypus Saussure, 1878, Archenopterus Otte, 1987, Atrella Gorochov, 2003, Calscirtus Otte, 1987, Furcitrella Gorochov, 2002, Fryerius Uvarov, 1940, Hemiphonoides Chopard, 1951, Hemitrella Gorochov, 2003, Heterecous Saussure, 1897, Homalotrypus Brancsik, 1895, Idiotrella Gorochov, 2002 (transferred from Aphonomorphini), Indotrella Gorochov, 2003, Insulascirtus Otte & Rentz, 1985, Matuanus Gorochov, 1986, Mnesibulus Stål, 1877, Noctitrella Gorochov, 1990, Ocellotrella Gorochov, 2021, Parametrypa Brunner von Wattenwyl, 1873 (transferred from Paroecanthini), Paranaudus Saussure, 1878, Phyllotrella Gorochov, 1988, Pixipterus Desutter-Grandcolas, 2016, Poliotrella Gorochov, 1988, Posus Bolívar, 1890, ProzƲenella Gorochov, 2002, Pseudomadasumma Shiraki, 1930, Rupilius Stål, 1876, Scepastus Gerstaecker, 1863, Trelleora Gorochov, 1988, Valiatrella Gorochov, 2005, Varitrella Gorochov, 2003, Xuanaeua He & Gorochov, 2015, ZƲenella Gorochov, 1988; genus group Hemiphonus Otte & Alexander, 1983: Hemiphonus Saussure, 1878, Riatina Otte & Alexander, 1983; genus group Madasumma Otte & Alexander, 1983: Madasumma Walker, 1869, Tamborina Otte & Alexander, 1983

Subtribe Podoscirtina Saussure, 1878: Allotrella Gorochov, 2006, Atruljalia Gorochov, 1988, Brevitrella Gorochov, 2004, Eupodoscirtus Gorochov, 2004, Kilimagryllus Sjöstedt, 1910, Malawitrella Gorochov, 2021, Malgasotrella Gorochov, 2004, NeozƲenella Gorochov, 2004, Ombrotrella Gorochov, 2006, Podoscirtus Serville, 1838, Spinotrella Gorochov, 2004, Stenotrella Gorochov, 2005, Ultratrella Gorochov, 2004, ZƲenellomorpha Gorochov, 2004 .

Remarks: The genus Parametrypa Brunner von Wattenwyl, 1873 was recently transferred to the Neotropical tribe Paroecanthini (Gorochov,2021) . However, this genus is from the southern African continent, and there are no males described for any species. Male characters are crucial for the classification of Oecanthidae . The poor knowledge of this genus and its discrepant distribution could support the reallocation of Parametrypa to Podoscirtini again. There is a similar situation for Idiotrella Gorochov, 2002 . According to Cigliano et al. (2022), this genus belongs to Aphonomorphini, a Neotropical tribe; however, the distribution of Idiotrella is through southern China and Indonesia. Moreover, the male genital structures and a developed stridulatory apparatus match with Podoscirtini, not Aphonomorphini, whose forewings have only longitudinal veins (see Aphonomorphini diagnosis below).