taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
FE5D87EC26156D7DDE0EFCAEFBF5F9AC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26116D72DDFBF9FFFA8CF8BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26116D72DDFBF9FFFA8CF8BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26116D72DDFBF9FFFA8CF8BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26116D72DDFBF9FFFA8CF8BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26116D72DDFBF9FFFA8CF8BC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881510/files/figure.png	https://doi.org/10.5281/zenodo.18881510	Fig. 101. Map of the distribution of all samples examined for B. sibiricus (Fabricius, 1781) s. lat., showing the distributions of the red-tailed taxon (colour pattern) sibiricus s. str. (orange spots) and the black-tailed taxon (colour pattern) flaviventris Friese, 1905 (blue spots) on either side of the Mongolian deserts (data from the collections of the AB, EA, EB, IAR 2006‒2012, IZCAS, LMOB, NHMUK, PW, RJ, UUH, and VLA, all non-Russian specimens identified or checked by PW, together with Russian records identified by AB). The southern border of Mongolia runs through the middle of the desert region where no bumblebees have been found. The yellow area is classified as desert. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI. The background map is international boundaries on the ‘World Imagery’ from ESRI with credits to ESRI, Maxar, Earthstar Geographics, and the GIS User Community.	Fig. 101. Map of the distribution of all samples examined for B. sibiricus (Fabricius, 1781) s. lat., showing the distributions of the red-tailed taxon (colour pattern) sibiricus s. str. (orange spots) and the black-tailed taxon (colour pattern) flaviventris Friese, 1905 (blue spots) on either side of the Mongolian deserts (data from the collections of the AB, EA, EB, IAR 2006‒2012, IZCAS, LMOB, NHMUK, PW, RJ, UUH, and VLA, all non-Russian specimens identified or checked by PW, together with Russian records identified by AB). The southern border of Mongolia runs through the middle of the desert region where no bumblebees have been found. The yellow area is classified as desert. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI. The background map is international boundaries on the ‘World Imagery’ from ESRI with credits to ESRI, Maxar, Earthstar Geographics, and the GIS User Community.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261E6D70DDAEF8CFFB03FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261E6D70DDAEF8CFFB03FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261E6D70DDAEF8CFFB03FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261E6D70DDAEF8CFFB03FB39.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261C6D76DDD0FB48FEB1FD1E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261C6D76DDD0FB48FEB1FD1E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261C6D76DDD0FB48FEB1FD1E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261C6D76DDD0FB48FEB1FD1E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261A6D75DD85FD6EFEDFFAD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261A6D75DD85FD6EFEDFFAD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261A6D75DD85FD6EFEDFFAD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC261A6D75DD85FD6EFEDFFAD0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26196D74DDD1FAD3FCAAF9AE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26196D74DDD1FAD3FCAAF9AE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26196D74DDD1FAD3FCAAF9AE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26196D74DDD1FAD3FCAAF9AE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26186D49DD89F9F9FE23FAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26186D49DD89F9F9FE23FAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26186D49DD89F9F9FE23FAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26186D49DD89F9F9FE23FAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26256D4EDDEAFA08FE37FCE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26256D4EDDEAFA08FE37FCE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26256D4EDDEAFA08FE37FCE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26256D4EDDEAFA08FE37FCE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26226D43DD4CFC3BFD82F983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26226D43DD4CFC3BFD82F983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26226D43DD4CFC3BFD82F983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC26226D43DD4CFC3BFD82F983.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262F6D40DD85F98DFA96FE51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262F6D40DD85F98DFA96FE51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262F6D40DD85F98DFA96FE51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262F6D40DD85F98DFA96FE51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262C6D47DDAFFE51FE6CFA66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262C6D47DDAFFE51FE6CFA66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262C6D47DDAFFE51FE6CFA66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262C6D47DDAFFE51FE6CFA66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262B6D44DDF8F9A1FAF1FE73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881494/files/figure.png	https://doi.org/10.5281/zenodo.18881494	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	Fig. 1. Support for apparent species’ gene coalescents within the subgenus Sibiricobombus Vogt, 1911 from unique longer haplotypes for COI barcode-like sequences (with the exception of the single short sequence of the taxon obtusus Richards, 1951) including likely numts (nuclear copies of mitochondrial sequences), obtained from an evolutionary tree estimated with MrBayes and using maximum likelihood to fit Poisson-Tree-Process (PTP) models. Numbers at each node show the probability that all daughter sequences to the right are parts of a single unique species. Branches change from blue to red at the node with the best fit (maximum local probability) for the change from inter-species to intra-species branching PTP models (species’ coalescent nodes) (grey dots artificially extend branches to make the tree easier to read). Each sequence is labelled with: sequence length; a morphological taxon name; a code for the colour pattern consisting of three characters: Y[ellow]/W[hite]/R[ed]band anteriorly + B[anded]/ U[nbanded]between the wing bases + W[hite]/R[ed]/B[lack]-tailed metasoma; a code consisting of a sequence identifier from the project database and a specimen identifier from the online database; its country and (for larger countries) state or province of origin. The sequence AT3% score is shown to the right of each sequence label. Sequences marked with ‘*’ are proxy types for names (Table 2) and sequences marked with ‘(*)’ match haplotypes for proxy types when the proxy types are not included in this tree. To the right in grey is the interpretation as the 11 species of the subgenus from the integrative analysis (see the text).	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262B6D44DDF8F9A1FAF1FE73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881500/files/figure.png	https://doi.org/10.5281/zenodo.18881500	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	Fig. 7. BEAST estimate of dated relationships as an ultrametric tree (outgroup B. rufocinctus Cresson, 1863 not shown) from six genes (COI, 16S, opsin, Ef-1α, ArgK, PEPCK) for species of the subgenus Sibiricobombus Vogt, 1911, estimated with a GTR+Γ+I model over 100 million MCMC generations and a 5% burn-in. Numbers below nodes are support values. Numbers above nodes are estimates of the age of a node in Ma before the present, with grey node bars representing the 95% highest posterior density interval of the age estimates. Note added in proof: the estimate has been re-run to include a PEPCK sequence for B. morawitzi, which increases the Bayesian posterior support for the group of morawitzi + niveatus-group + asiaticus-group from 0.5 to 0.75.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262B6D44DDF8F9A1FAF1FE73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881502/files/figure.png	https://doi.org/10.5281/zenodo.18881502	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	Figs 8‒9. Geographical distribution of the sites from which Sibiricobombus Vogt, 1911 samples have been barcoded, mapped to indicate the approximate location and a part of the extent of occurrence (EOO) for each corresponding species as accepted in Fig. 7. Spherical projection with international boundaries as recognized by the UN shown as grey lines. Map projected in ArcGIS using the World_Shaded_Relief basemap © 2014 ESRI.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
FE5D87EC262B6D44DDF8F9A1FAF1FE73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18881508/files/figure.png	https://doi.org/10.5281/zenodo.18881508	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	Figs 90–100. Morphology of the male genitalia for species of the subgenus Sibiricobombus Vogt, 1911 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 90. B. sibiricus (Fabricius, 1781). 91. B. semenovi Morawitz, 1887 stat. rev. 92. B. oberti Morawitz, 1883 (left penis-valve head recurved hook missing). 93. B. morawitzi Radoszkowski, 1876. 94. B. sulfureus Friese, 1905 (left penisvalve head recurved hook missing). 95. B. niveatus Kriechbaumer, 1870. 96. B. obtusus Richards, 1951. 97. B. tescorum Williams nom. nov. et stat. rev. 98. B. longiceps Smith, 1878 stat. rev.. 99. B. falsificus Richards, 1930 stat. rev.. 100. B. asiaticus Morawitz, 1875. Images taken by L. Berridge. Scale bars = 1 mm.	2026-02-19	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin		Zenodo	biologists	Williams, Paul H.;An, Jiandong;Byvaltsev, Alexandr;Jonghe, Guillaume Ghisbain Roland De;Huang, Jiaxing;Mei, Maurizio;Monfared, Alireza;Orr, Michael C.;Raina, Rifat;Streinzer, Martin			
