taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
971B3C7E19345262A911D6469EE7F473.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571116	https://doi.org/10.3897/herpetozoa.39.e184954.figure4	Figure 4. Comparative characters between Charadrahyla viruela sp. nov. and C. nephila. A. Lateral view of head of a live individual of C. viruela from ca 1.2 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, which shows the truncate snout in lateral profile; the “ starry-eyed ” reticulated iris; and the characteristic white blotch on the lip posterior to the eyes; B. Lateral view of head of a live individual of C. nephila from 6.6 km S of La Esperanza, at Arroyo Buena Vista, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristically rounded snout in lateral profile; the solid, copper-colored iris with no reticulations; and the lack of a distinctive white blotch on the upper lip posterior to the eye; C. Ventral view of a preserved C. viruela (KU 86966) from 4.2 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic pale cream to whitish ventral coloration; the seven large tan blotches on the throat; the lack of immaculate white spotting on the throat and chest; and the dark brown banding instead of white striping on ventrolateral portions of the thighs and tibia; D. Ventral view of a preserved C. nephila (136678) from 11.9 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic darker tan to brown ventral coloration, lack of tan blotches on throat, presence of immaculate white spotting on throat, chest, and upper abdomen, and immaculate white striping on the ventrolateral portions of the thighs and tibia.	Figure 4. Comparative characters between Charadrahyla viruela sp. nov. and C. nephila. A. Lateral view of head of a live individual of C. viruela from ca 1.2 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, which shows the truncate snout in lateral profile; the “ starry-eyed ” reticulated iris; and the characteristic white blotch on the lip posterior to the eyes; B. Lateral view of head of a live individual of C. nephila from 6.6 km S of La Esperanza, at Arroyo Buena Vista, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristically rounded snout in lateral profile; the solid, copper-colored iris with no reticulations; and the lack of a distinctive white blotch on the upper lip posterior to the eye; C. Ventral view of a preserved C. viruela (KU 86966) from 4.2 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic pale cream to whitish ventral coloration; the seven large tan blotches on the throat; the lack of immaculate white spotting on the throat and chest; and the dark brown banding instead of white striping on ventrolateral portions of the thighs and tibia; D. Ventral view of a preserved C. nephila (136678) from 11.9 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic darker tan to brown ventral coloration, lack of tan blotches on throat, presence of immaculate white spotting on throat, chest, and upper abdomen, and immaculate white striping on the ventrolateral portions of the thighs and tibia.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
971B3C7E19345262A911D6469EE7F473.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571117	https://doi.org/10.3897/herpetozoa.39.e184954.figure5	Figure 5. Comparison of individuals of Charadrahyla species. We have included photos of live specimens whenever possible, but were unable to obtain photos of live individuals of two populations. A. Charadrahyla viruela sp. nov. (MZFC 37345, paratype) from Sierra Juárez, at Arroyo Tres Manantiales, La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 1,688 m asl; B. Charadrahyla viruela sp. nov. from Sierra Juárez, at La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 1,630 m asl, photo by Jesse Delia; C. Charadrahyla nephila from Sierra Juárez, in the upper reaches of Arroyo Tepetotutla, Municipio de San Felipe Usila, Oaxaca, at 2,600 m asl, photo by Rogelio Simón Salvador; D. Charadrahyla nephila Arroyo Grande at 16 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 2,150 m asl (CIG 1871); E, F. Charadrahyla nephila type-locality, 5.8 km SW of Totontepec, Municipio de Totontepec Villa de Morelos, Oaxaca, at 2,000 m asl; G, H. Charadrahyla chaneque type-locality, 6.2 km S of Rayón Mescalapa, Municipio de Rayón, Chiapas at 1,790 m asl.	Figure 5. Comparison of individuals of Charadrahyla species. We have included photos of live specimens whenever possible, but were unable to obtain photos of live individuals of two populations. A. Charadrahyla viruela sp. nov. (MZFC 37345, paratype) from Sierra Juárez, at Arroyo Tres Manantiales, La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 1,688 m asl; B. Charadrahyla viruela sp. nov. from Sierra Juárez, at La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 1,630 m asl, photo by Jesse Delia; C. Charadrahyla nephila from Sierra Juárez, in the upper reaches of Arroyo Tepetotutla, Municipio de San Felipe Usila, Oaxaca, at 2,600 m asl, photo by Rogelio Simón Salvador; D. Charadrahyla nephila Arroyo Grande at 16 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, at 2,150 m asl (CIG 1871); E, F. Charadrahyla nephila type-locality, 5.8 km SW of Totontepec, Municipio de Totontepec Villa de Morelos, Oaxaca, at 2,000 m asl; G, H. Charadrahyla chaneque type-locality, 6.2 km S of Rayón Mescalapa, Municipio de Rayón, Chiapas at 1,790 m asl.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571114	https://doi.org/10.3897/herpetozoa.39.e184954.figure2	Figure 2. Charadrahyla viruela sp. nov. Holotype (INIRENA 2950). Live specimen (A) and preserved in dorsolateral (B) and ventrolateral (C) views.	Figure 2. Charadrahyla viruela sp. nov. Holotype (INIRENA 2950). Live specimen (A) and preserved in dorsolateral (B) and ventrolateral (C) views.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571115	https://doi.org/10.3897/herpetozoa.39.e184954.figure3	Figure 3. Paratypes of Charadrahyla viruela sp. nov. in preservative, showing variation amongst specimens from different localities, all in the Sierra Juárez, Oaxaca in dorsal (A, C, E, G, I) and ventral (B, D, F, H, J) view.	Figure 3. Paratypes of Charadrahyla viruela sp. nov. in preservative, showing variation amongst specimens from different localities, all in the Sierra Juárez, Oaxaca in dorsal (A, C, E, G, I) and ventral (B, D, F, H, J) view.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571116	https://doi.org/10.3897/herpetozoa.39.e184954.figure4	Figure 4. Comparative characters between Charadrahyla viruela sp. nov. and C. nephila. A. Lateral view of head of a live individual of C. viruela from ca 1.2 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, which shows the truncate snout in lateral profile; the “ starry-eyed ” reticulated iris; and the characteristic white blotch on the lip posterior to the eyes; B. Lateral view of head of a live individual of C. nephila from 6.6 km S of La Esperanza, at Arroyo Buena Vista, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristically rounded snout in lateral profile; the solid, copper-colored iris with no reticulations; and the lack of a distinctive white blotch on the upper lip posterior to the eye; C. Ventral view of a preserved C. viruela (KU 86966) from 4.2 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic pale cream to whitish ventral coloration; the seven large tan blotches on the throat; the lack of immaculate white spotting on the throat and chest; and the dark brown banding instead of white striping on ventrolateral portions of the thighs and tibia; D. Ventral view of a preserved C. nephila (136678) from 11.9 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic darker tan to brown ventral coloration, lack of tan blotches on throat, presence of immaculate white spotting on throat, chest, and upper abdomen, and immaculate white striping on the ventrolateral portions of the thighs and tibia.	Figure 4. Comparative characters between Charadrahyla viruela sp. nov. and C. nephila. A. Lateral view of head of a live individual of C. viruela from ca 1.2 km S of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, which shows the truncate snout in lateral profile; the “ starry-eyed ” reticulated iris; and the characteristic white blotch on the lip posterior to the eyes; B. Lateral view of head of a live individual of C. nephila from 6.6 km S of La Esperanza, at Arroyo Buena Vista, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristically rounded snout in lateral profile; the solid, copper-colored iris with no reticulations; and the lack of a distinctive white blotch on the upper lip posterior to the eye; C. Ventral view of a preserved C. viruela (KU 86966) from 4.2 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic pale cream to whitish ventral coloration; the seven large tan blotches on the throat; the lack of immaculate white spotting on the throat and chest; and the dark brown banding instead of white striping on ventrolateral portions of the thighs and tibia; D. Ventral view of a preserved C. nephila (136678) from 11.9 km S of Vista Hermosa, Municipio de Santiago Comaltepec, Oaxaca which shows the characteristic darker tan to brown ventral coloration, lack of tan blotches on throat, presence of immaculate white spotting on throat, chest, and upper abdomen, and immaculate white striping on the ventrolateral portions of the thighs and tibia.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571118	https://doi.org/10.3897/herpetozoa.39.e184954.figure6	Figure 6. Ventral aspect of hand A. and foot B. of Charadrahyla viruela sp. nov. INIRENA 2950, Highway 175 between Vista Hermosa and La Esperanza, 1.9 km ENE of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, Mexico. The black bar represents 10 mm. Illustrations by Iván Ahumada-Carrillo.	Figure 6. Ventral aspect of hand A. and foot B. of Charadrahyla viruela sp. nov. INIRENA 2950, Highway 175 between Vista Hermosa and La Esperanza, 1.9 km ENE of La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, Mexico. The black bar represents 10 mm. Illustrations by Iván Ahumada-Carrillo.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571119	https://doi.org/10.3897/herpetozoa.39.e184954.figure7	Figure 7. Distribution map of species of Charadrahyla in southeastern Mexico. Diamonds represent type localities. See materials and methods for source of records.	Figure 7. Distribution map of species of Charadrahyla in southeastern Mexico. Diamonds represent type localities. See materials and methods for source of records.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
15CCDCC6CB7857ADA717EE3C5B1D1458.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1571120	https://doi.org/10.3897/herpetozoa.39.e184954.figure8	Figure 8. Photo of the locality and habitat at the type-locality of Charadrahyla viruela sp. nov. near La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, Mexico.	Figure 8. Photo of the locality and habitat at the type-locality of Charadrahyla viruela sp. nov. near La Esperanza, Municipio de Santiago Comaltepec, Oaxaca, Mexico.	2026-03-27	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.		Zenodo	biologists	Canseco-Márquez, Luis;Grünwald, Christoph I.;Reyes-Velasco, Jacobo;Montaño-Ruvalcaba, Carlos;Campbell, Jonathan A.			
