identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AE1212727AA75F939F2904CA8B873455.text	AE1212727AA75F939F2904CA8B873455.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus dayangmerindu Aulia & Nurinsiyah 2026	<div><p>Chamalycaeus dayangmerindu Aulia &amp; Nurinsiyah sp. nov.</p><p>Fig. 5 c</p><p>Material examined.</p><p>Holotype. Indonesia • Shell H = 2.7 mm; D = 3.9 mm; HA = 1.4 mm; DA = 1.4 mm; whorl 3; South Sumatra, Ogan Komering Ulu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92067&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92067/lat -4.067333)">Padang Bindu (Plot 11)</a>; 4°04.04'S, 103°55.24'E; alt. 124 m; May-June 2021; Nurinsiyah leg.; MZB Gst. 24.132 . Paratypes. Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24131/40 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24132/1 .</p><p>Description (n = 11).</p><p>Shell minute size with H = 2–2.7 mm (mean 2.3 mm); D = 3.1–4.1 mm (mean 3.5 mm); HA = 1–1.4 mm (mean 1.2 mm); DA = 1–1.2 mm (mean 1.1 mm); 3–3.5 whorls. Shell dextral; depressed conical shaped, wider than tall, with depressed apex; white color and glossy; body whorl rounded, separated by a deep suture; protoconch smooth; the teleoconch with fine spiral lines, without radial ribs; constriction ~ 0.5 mm from aperture; followed by short bulging sutural tube, with 7–9 radial micro-tunnels; region termed R 3 by Páll-Gergely et al. (2017) (which usually bears a swelling in most species of Alycaeinae) absent (i. e., the aperture is situated in the immediate vicinity of the constriction); aperture almost perfectly round, without wrinkles; peristome thickened, not reflected; the boundary between inner and outer peristomes visible but not conspicuous; umbilicus wide.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species is so far only recorded from Padang Bindu Karst, South Sumatra.</p><p>Remarks.</p><p>Although spiral striation generally absent in Dicharax (Páll-Gergely et al. 2020), this character is obvious in D. candrakirana from Sempu Island, Java (Nurinsiyah and Hausdorf 2017). However, that species has a well-developed R 3 with low swelling. The most similar species are Chamalycaeus microconus Möllendorff, 1887 and Chamalycaeus mixtus Zilch, 1957 from Peninsular Malaysia, due to the absence of R 3. However, the former differs from the new species due to its strong radial ribs, and the latter has practically no constriction and its peristome is not expanded.</p><p>Etymology.</p><p>The species name Dayang Merindu was inspired by Princess Dayang Merindu, a figure from local folklore. According to the legend, Princess Dayang Merindu ignored the greeting of a man, as she was already married. Her action was then perceived as arrogance, and the man cursed her into stone. The stone associated with this tale is located inside Gua Putri, a karst cave formation in the Padang Bindu area.</p></div>	https://treatment.plazi.org/id/AE1212727AA75F939F2904CA8B873455	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
60FD0277B3DE5FFB8B0386B4475BFE4D.text	60FD0277B3DE5FFB8B0386B4475BFE4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclophorus perdix subsp. tuba (G. B. Sowerby I 1842)	<div><p>Cyclophorus perdix tuba (Sowerby, 1842)</p><p>Fig. 4 a</p><p>Cyclophorus tuba Sowerby, 1842: 122, pl. 27, figs 129, 130. von Martens 1867: 133, pl. 3, figs 2–4.</p><p>Cyclophorus perdix tuba – van Benthem Jutting 1959: 69 –71.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 2)</a>; 4°04.00'S, 103°55.79'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23952/2 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23953/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93433&amp;materialsCitation.latitude=-4.0723333" title="Search Plazi for locations around (long 103.93433/lat -4.0723333)">Padang Bindu (Plot 16)</a>; 4°04.34'S, 103°56.06'E; alt. 139 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23954/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9315&amp;materialsCitation.latitude=-4.0668335" title="Search Plazi for locations around (long 103.9315/lat -4.0668335)">Padang Bindu (Plot 4)</a>; 4°04.01'S, 103°55.89'E; alt. 111 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23955/2 .</p><p>Description (n = 5).</p><p>Shell medium size with H = 12.5–22.1 mm (mean 18.2 mm); D = 17.8–32.2 mm (mean 27.2 mm); HA = 7.4–21.6 mm (mean 14.1 mm); DA = 8.9–17.6 mm (mean 14.9 mm); 4.2–5 whorls. Shell conical, with a low tower and very large body whorl, not too thick, and not shiny; rotation dextral; slightly transparent on the inside; pale brown accompanied by a dark brown and white pattern in the form of lines on the suture edges; the pattern fades slightly in the middle of the body whorl exactly ¼ after the keel; a spiral dark brown pattern appears at the bottom of the body whorl; umbilicus wide; keel in the middle of the body whorl; aperture almost round, slightly slanted; peristome expands upwards.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species is widely distributed in Sumatra.</p><p>Remarks.</p><p>In general, in the species and subspecies of Cyclophorus perdix almost all shell morphology is similar, namely the shell shape, keel on the body whorl, and aperture shape (Novianti et al. 2023). The difference lies on the taper level of the keel, the position of the keel, the width of the aperture, and the degree of reflection of the aperture. Apart from that, a quite distinctive difference is in the spiral pattern of the shell.</p></div>	https://treatment.plazi.org/id/60FD0277B3DE5FFB8B0386B4475BFE4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
1D8DBA5ED69658E9AAD267E59C3EC034.text	1D8DBA5ED69658E9AAD267E59C3EC034.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina liwaensis Aldrich 1898	<div><p>Diplommatina liwaensis Aldrich, 1898</p><p>Fig. 7 a</p><p>Diplommatina liwaensis Aldrich, 1898: 4, pl. 1, figs 3, 4.</p><p>Diplommatina liwaensis – van Benthem Jutting 1959: 83. Maassen 2002 b: 169, figs 13, 14. Marwoto 2016: 18.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 10)</a>; 4°04.11'S, 103°55.34'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZBGst. 24148/4 • South Sumara, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24113/204 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24112/7 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 2)</a>; 4°04.00'S, 103°55.79'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24111/8 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93467&amp;materialsCitation.latitude=-4.0705" title="Search Plazi for locations around (long 103.93467/lat -4.0705)">Padang Bindu (Plot 3)</a>; 4°04.23'S, 103°56.08'E; alt. 114 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24146/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24108/11 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24110/5 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24109/6 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9245&amp;materialsCitation.latitude=-4.0675" title="Search Plazi for locations around (long 103.9245/lat -4.0675)">Suka Merindu (Plot 6)</a>; 4°04.05'S, 103°55.47'E; alt. 123 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24147/5 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 1)</a>; 4°04.00'S, 103°55.79'E; alt. 146 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24118/1 .</p><p>Description (n = 34).</p><p>Shell small size with H = 5.8–9.33 mm (mean 6.7 mm); D = 2.6–4.1 mm (mean 3.2 mm); HA = 1.5–2.1 mm (mean 1.8 mm); DA = 1.7–2.3 mm (mean 2.1 mm); 6.5–8.5 whorls. Shell spindle shape, thin, with a conical protoconch then narrowed at the last whorl; penultimate whorl largest, separated by deep sutures; smooth protoconch; teleoconch with smooth, oblique, dense radial ribs with 26–40 ribs on the penultimate whorl and 26–39 on the body whorl; color pale brown to white, slightly transparent; narrowing of the ventral part of the body whorl; aperture rounded, outer lip widened; peristome is wide; columellar lamella small; inner palatal indistinct; umbilicus closed.</p><p>Remarks.</p><p>This species is similar to Diplommatina auriculata Möllendorff, 1897 from Java based on the characteristics of its medium-sized shell, protruding columellar lamella teeth, and close-set ribs. D. liwaensis has a shell ~ 4 mm high with close ribs and small columellar lamella teeth, while D. liwaensis has a shell height of 5–9 mm and quite large, protruding, columellar teeth. Apart from the columellar teeth (columellar lamella), the palatal radials differ. Diplommatina liwaensis does not have palatal radial lines. A slight angle at the columellar tip towards the basal part in D. auriculata is not found in D. liwaensis .</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species was previously recorded in North and West Sumatra (Maassen 2002 b). The distribution in South Sumatra is a new record. The species has never been recorded outside Sumatra, and is thus endemic to the island.</p></div>	https://treatment.plazi.org/id/1D8DBA5ED69658E9AAD267E59C3EC034	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
254C85A310FC5E28B70CE8F2DB414DC7.text	254C85A310FC5E28B70CE8F2DB414DC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina wilhelminae Maassen 2002	<div><p>Diplommatina wilhelminae Maassen, 2002</p><p>Fig. 7 b</p><p>Diplommatina wilhelminae Maassen, 2002 b: 175, figs 29, 30.</p><p>Diplommatina wilhelminae – Marwoto 2016: 18.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24199/3 .</p><p>Description (n = 3).</p><p>Shell small size with H = 5.4–5.56 mm (mean 5.5 mm); D = width 2.6– 2.6 mm (mean 2.6 mm); HA = 1.1–1.2 mm (mean 1.2 mm); DA = 1.1–1.2 mm (mean 1.1 mm); 6–6.5 whorls. Shell spindle shaped, with a conical protoconch and narrow body whorl; penultimate whorl approximately the same width as the last one, separated by a deep suture; shell dextral; protoconch smooth; teleoconch with straight radial ribs, but slightly oblique in body whorl with 12–14 ribs on the penultimate and 14–16 ribs on the body whorl), distinct, rather sharp; distinct spiral striae present; quarter last whorl slightly up towards the suture; elongated palatalis absent; umbilicus closed; peristome double, widened, palatal side tortuous; outer peristome widened.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species was previously recorded in West Sumatra (Maassen 2002 b). The distribution in South Sumatra is a new record. The species has never been recorded outside Sumatra and is thus endemic to the island.</p></div>	https://treatment.plazi.org/id/254C85A310FC5E28B70CE8F2DB414DC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
E3AE170B50C550CFA8D3E5F04FA1956D.text	E3AE170B50C550CFA8D3E5F04FA1956D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Japonia ciliocincta (E. von Martens 1865)	<div><p>Japonia ciliocincta (E. von Martens, 1865)</p><p>Figs 4 b, 9 a</p><p>Cyclophorus ciliocinctus E. von Martens, 1865: 52 .</p><p>Lagochilus ciliocinctum – van Benthem Jutting 1959: 61. Marwoto 2016: 17.</p><p>Japonia ciliocinctum – Vermeulen and Whitten 1998: 44, fig. 21.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23956/10 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23957/2 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93433&amp;materialsCitation.latitude=-4.0723333" title="Search Plazi for locations around (long 103.93433/lat -4.0723333)">Padang Bindu (Plot 16)</a>; 4°04.34'S, 103°56.06'E; alt. 139 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23958/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23959/4 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23960/11, MZB Gst. 24101/19 ) • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24153/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9245&amp;materialsCitation.latitude=-4.0675" title="Search Plazi for locations around (long 103.9245/lat -4.0675)">Suka Merindu (Plot 6)</a>; 4°04.05'S, 103°55.47'E; alt. 123 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24152/3 .</p><p>Description (n = 28).</p><p>Shell small size with H = 8.1–12.6 mm (mean 10.7 mm); D = 8.2–13 mm (mean 10.7 mm); HA = 3.6–5.8 mm (mean 5 mm); DA = 3.9–6.6 mm (mean 5.6 mm); 5–6.5 whorls. Operculum height 5.7 mm, width 4.9 mm. Shell conical, almost the same length as width, with the top slightly pointed; dextral; regularly the diameter of the whorl increases, with large body whorl, rather thin; white to pale yellowish brown, with few to many dark brown oblique lines, the spacing between the lines is quite loose; protoconch often dark purple; the outer shell layer is thin, brownish, form of a series spiral spines, falls off easily and leaves small holes; peristome double, whitish, outer edge often brown; aperture almost diagonal, continuous with a gap at the top of the upper right aperture; operculum oval, flat with exterior sculpture appressed radially spiral lamellae.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species is distributed in Sumatra, Java, and Nusa Tenggara. The distribution in South Sumatra is a new record.</p><p>Remarks.</p><p>Several Padang Bindu specimens have a different pattern from the type specimen: thin lines that are tightly oblique and indistinct. Meanwhile, the type specimen of Japonia ciliocincta has dark and oblique stripes with gaps between them. However, it cannot be determined if these differences constitute species differences because there are no other distinctive differences. Thus, further research is needed to confirm this.</p></div>	https://treatment.plazi.org/id/E3AE170B50C550CFA8D3E5F04FA1956D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
F282CC4D747052D0B23B3469A6766B9E.text	F282CC4D747052D0B23B3469A6766B9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagocheilus ciliferus (Mousson 1849)	<div><p>Lagocheilus ciliferus (Mousson, 1849)</p><p>Fig. 4 c</p><p>Cyclostoma ciliferus Mousson, 1849: 56–57, pl. 7, fig. 3; pl. 20, fig. 8.</p><p>Lagochilus ciliferus – van Benthem Jutting 1948: 557, fig. 15. Marwoto 2016: 17.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 10)</a>; 4°04.11'S, 103°55.34'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24105/21 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24103/16 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24104/29 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24102/2 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24106/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24107/1 .</p><p>Description (n = 17).</p><p>Shell small size with H = 8.0– 10.1 mm (mean 9.3 mm); D = 9.3–11.8 mm (mean 10.7 mm); HA = 3.7–5.2 mm (mean 4.7 mm); DA = 4–5.8 mm (mean 5 mm); 5–6 whorls. Shell conical, with pointed apex, body whorl with largest circle; keel in center of body whorl; suture clear and deep; dextral; quite thin; surrounded by spiral lines, no radial lines; monochromatic white to reddish brown; not shiny; umbilicus quite wide; aperture rounded, oblique, only slightly pointed at the top right; peristome double.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species is distributed in Sumatra and Java. The specimens found in South Sumatra constitute a new record.</p><p>Remarks.</p><p>Lagocheilus ciliferus can be differentiated from Japonia ciliocincta from the pattern on the shell, as well as by the shape of the shell, which has a prominent body whorl that tends to widen and has a keel but does not have radial ribs throughout the shell.</p></div>	https://treatment.plazi.org/id/F282CC4D747052D0B23B3469A6766B9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
8B9926ACE8DC54499B686558CCC95ED3.text	8B9926ACE8DC54499B686558CCC95ED3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Opisthostoma platycephalum van Benthem Jutting 1952	<div><p>Opisthostoma platycephalum van Benthem Jutting, 1952</p><p>Fig. 7 d</p><p>Opisthostoma (Opisthostoma) platycephalum van Benthem Jutting, 1952: 26, fig. 11.</p><p>Opisthostoma platycephalum — Maassen 2001: 35; 2002 b: 178, figs 43, 44. Aulia et al. 2024: 43 –50, fig. 2.</p><p>Plectostoma platycephalum — Marwoto 2016: 18.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 2)</a>; 4°04.00'S, 103°55.79'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23866/26 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23867/39 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23868/25 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23869/2 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23870/3 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 23871/16 .</p><p>Description (n = 45).</p><p>Shell minute size with H = 1.2–1.9 mm (mean 1.6 mm); D = 1.7–2.6 mm (mean 2.3 mm); Diameter without aperture = 1.2–1.8 mm (mean 1.5 mm); HA = 0.7–1.1 mm (mean 0.9 mm); DA = 0.7–1 mm (mean 0.8 mm); 3.5 whorls. Shell dextral; apical spire whorl with a depressed conical shape; top whorls in one plane, somewhat oblique on the axis of the shell; whitish color; protoconch smooth; second whorl with smooth rather close ribs (9–20 ribs per mm or total 18–26 ribs) slightly obliquely; following whorl (body whorl) has more distant ribs (4–8 ribs per mm or total 12–18 ribs); last whorl trumpet-shaped, towards the aperture, directed upward, but not deviating transversely, separated by a deep suture; shell height mm, shell width mm (without aperture) or (with aperture). Round aperture, aperture width mm, aperture height mm. Peristome continuous, double; umbilicus open, deep, very narrow.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species was previously recorded in Peninsular Malaysia, Aceh, and North Sumatra (Maassen 2002 b). The distribution in South Sumatra is a new record (Aulia et al. 2024).</p><p>Remarks.</p><p>All specimens of O. platycephalum from Padang Bindu Karst, South Sumatra are larger than those described by van Benthem Jutting (1952). Holotype: shell height 0.6 mm, shell diameter 1.2 mm, aperture diameter 0.4 mm; paratype with shell height 0.6–0.7 mm, shell diameter 1.1–1.2 mm, and aperture diameter 0.4 mm. According to van Benthem Jutting (1952), O. platycephalum has a shell that is not transparent, whereas a few O. platycephalum specimens from the Padang Bindu Karst are slightly transparent. Specimens from the recent collections probably fresher compared to the specimen examined by van Benthem Jutting because old shells tend to become opaque due to weathering. Based on the overall characteristics of shell morphology, the Opisthostoma samples found in Padang Bindu were identified as O. platycephalum . We follow the updated classification in MolluscaBase (2025) to include the species to the genus Opisthostoma .</p></div>	https://treatment.plazi.org/id/8B9926ACE8DC54499B686558CCC95ED3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
B48487D9E6635D4BBD95C860F7A50F9C.text	B48487D9E6635D4BBD95C860F7A50F9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pincerna crenilabris subsp. crenilabris (Mollendorff 1897)	<div><p>Pincerna crenilabris crenilabris (Möllendorff, 1897)</p><p>Figs 6, 9 c</p><p>Alycaeus (Orthalycaeus) crenilabris Möllendorff, 1897: 93 .</p><p>Pincerna crenilabris — Páll-Gergely et al. 2020: 173.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 10)</a>; 4°04.11'S, 103°55.34'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24144/4 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24137/15 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9315&amp;materialsCitation.latitude=-4.0668335" title="Search Plazi for locations around (long 103.9315/lat -4.0668335)">Padang Bindu (Plot 4)</a>; 4°04.01'S, 103°55.89'E; alt. 111 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24136/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24135/40 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24133/13, MZB Gst. 24134/19 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 2)</a>; 4°04.00'S, 103°55.79'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24143/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9245&amp;materialsCitation.latitude=-4.0675" title="Search Plazi for locations around (long 103.9245/lat -4.0675)">Suka Merindu (Plot 6)</a>; 4°04.05'S, 103°55.47'E; alt. 123 m; May-June 2021; A. S. Nurinsiyah leg.; MZB 24145/4 .</p><p>Description (n = 49).</p><p>Shells micro size with H = 3.8–5 mm (mean 4.4 mm); D = 3.8–4.8 mm (mean 4.2 mm); HA = 1.5–2 mm (mean 1.7 mm); DA = 1.4–2.7 mm (mean 2.3 mm); 4–5 whorls. Operculum width 1.4–1.5 mm, height 1.4–1.5 mm, concavity / thickness 0.4–0.5 mm. Shell dextral; almost globular with a depressed apex; pale pink or yellowish brown color, not transparent; protoconch smooth, R 1 with ~ 2.25 whorls, with widely-spaced, strong ribs (~ 20–22 ribs on the last half whorl of R 1) and fine spiral striation between the ribs, R 2 short, of ~ 1 / 8 whorl, with ~ 12 ribs, rib density slightly higher than that of R 1, constriction between R 2 and R 3 shallow, R 3 slightly shorter than a 1 / 4 whorl, ribs more widely-spaced on R 3 than on R 1, aperture oblique to shell axis in lateral view, aperture mostly rounded with slight upper incision (at the parieto-palatal corner), and the palatal part of the inner peristome may be slightly edentate, inner and outer peristomes clearly separated, inner peristome thickened and slightly protruding, outer peristome strongly expanding, having a projection on the basal part, and interestingly, turns anteriorly (above the inner peristome) at the upper palatal region, umbilicus rounded, open, very narrow, nearly half covered by peristome; operculum concave, inner surface with a tiny central nipple, outer surface multispiral, without elevated lamella.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The species was previously recorded in western Java (see Páll-Gergely et al. 2020 and references therein). The distribution in Padang Bindu karst is a new record.</p><p>Remarks.</p><p>Comparing typical Pincerna crenilabris (see photos of the holotype, Fig. 6 a), and the shells collected by us in Sumatra, we could observe some differences in rib density. Namely, on the body whorl of the Sumatran specimens, 35 ribs were counted on the body whorl before the constriction. In contrast, on the photographed holotype, there are ~ 45 ribs on the last whorl behind the constriction. Nevertheless, we interpret these differences as parts of the intrasubspecific variability.</p><p>Two subspecies of Pincerna crenilabris were described by van Benthem Jutting (1959), namely Alycaeus crenilabris laevis and Alycaeus crenilabris latecostatus . Since both names were primary homonyms, Páll-Gergely et al. (2020) proposed replacement names for both taxa (i. e., P. crenilabris korintjiensis Páll-Gergely, 2020 and P. crenilabris juttingae Páll-Gergely, 2020, respectively). We examined the photos of the holotype of Pincerna (?) crenilabris korintjiensis (ZMA. MOLL 135672, Fig. 5 d) (described from “ Kajo Aro Estate, Mt. Korintji ”) and confirmed that this is a valid Chamalycaeus species, which differs from P. crenilabris in the more conical shells shape ( P. crenilabris is rather globular) and the much denser R 2 ribs. Pincerna crenilabris juttingae is distributed in Brastagi, North Sumatra. No illustrations of this taxon have ever been published. Based on its shell characters (Fig. 6 c), P. crenilabris juttingae differs from the nominotypical subspecies by the smooth R 3, and the absence of the basal projection of the outer peristome, which justifies its distinction as a subspecies.</p></div>	https://treatment.plazi.org/id/B48487D9E6635D4BBD95C860F7A50F9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
A052882F48075DD29CDD74613296C9F9.text	A052882F48075DD29CDD74613296C9F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectostoma kitteli Maassen 2002	<div><p>Plectostoma kitteli Maassen, 2002</p><p>Fig. 7 c</p><p>Plectostoma kitteli Maassen, 2002: 176, figs 35, 36.</p><p>Plectostoma kitteli – Liew et al. 2014: 83 –85, fig. 4 A – F. Marwoto 2016: 18.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 22114/2320 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 22115/16 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93467&amp;materialsCitation.latitude=-4.0705" title="Search Plazi for locations around (long 103.93467/lat -4.0705)">Padang Bindu (Plot 3)</a>; 4°04.23'S, 103°56.08'E; alt. 114 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24116/23 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst 24117/36 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92667&amp;materialsCitation.latitude=-4.0721664" title="Search Plazi for locations around (long 103.92667/lat -4.0721664)">Padang Bindu (Plot 15)</a>; 4°04.33'S, 103°55.60'E; alt. 205 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24121/1 .</p><p>Description (n = 45).</p><p>Shell micro size with H with tube 3.5–5.3 mm (mean 3.9 mm); shell height without tube 2.3–3.9 mm (mean 2.7 mm); D with tube 3.3–4.8 (mean 3.7 mm); DA from inner peristome 1.1–1.4 mm (mean 1.3 mm); DA from outer peristome 1.2–1.8 mm (mean 1.4 mm); HA from inner peristome 1.1–1.5 mm (mean 1.3 mm); HA from outer peristome 1.5–2.1 mm (mean 1.7 mm); 7 whorls. Shell dextral; conical, sinistroid, with slightly convex sides; apex slightly oblique; width of each whorl increases regularly; brownish yellow color, not transparent; protoconch smooth; teleoconch with sharp and distinct radial ribs (7 or 8 ribs per mm on the penultimate whorl). Spiral striae absent; umbilicus open, with diameter 0.1 mm; peristome double, outer peristome widened; inner peristome stands out clearly, slightly widened.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution in Sumatra.</p><p>The species was previously recorded in West Sumatra by Maassen (2002 b). The distribution in South Sumatra is a new record. The species was never recorded outside Sumatra and is thus endemic to the island.</p><p>Remarks.</p><p>All specimens of Plectostoma kitteli from the Padang Bindu Karst are larger in width and height than those collected by Mr. Kittel in West Sumatra. We initially suspect the species to be a new species for Plectostoma . However, after examining the species morphology, range of morphometry as well as the ribbing, we concluded that this is P. kitteli . Genetic analysis, which is beyond the scope of the current research, is recommended to solve the systematics issue.</p></div>	https://treatment.plazi.org/id/A052882F48075DD29CDD74613296C9F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
D317C14533CC5251A9B9CF5BDEF0A270.text	D317C14533CC5251A9B9CF5BDEF0A270.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupina turgidula Dohrn 1881	<div><p>Pupina turgidula Dohrn, 1881</p><p>Fig. 8 a, b</p><p>Pupina turgidula Dohrn, 1881: 66 .</p><p>Pupina (Eupupina) treubi — O. Boettger, 1890: 157, pl. VI, fig. 8 a, b.</p><p>Pupina treubi — van Benthem Jutting, 1948: 381, figs 34, 36. Maassen 2002 a: pl. 30, fig. 5. Heryanto 2001: 767, 2008: 362. Nurinsiyah et al. 2023: 52, figs 1 e – g, 2 d.</p><p>Pupina verbeeki — van Benthem Jutting 1948: 582, figs 34 E, 37.</p><p>Pupina turgidula — van Benthem Jutting 1959: 75. Maassen 2002 a: 282, fig. 6.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24119/2, 24120/1 .</p><p>Description (n = 3).</p><p>Shell small size with H = 8.6–9.1 mm (mean 8.9 mm); D = 5.9–6.3 mm (mean 6.2 mm); HA = 3.2–3.3 mm (mean 3.3 mm); DA = 3.1–3.2 mm (mean 3.1 mm); 4.75–5 whorls. Shell dextral; conical, rather thin, with an apex pointed but not sharp; pale white to yellowish white, glossy, slightly transparent; body whorl largest with deep suture; spiral striae absent; umbilicus closed; aperture round, nearly vertical, with two short teeth near or on the parietal near the peristome.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution in Sumatra.</p><p>The species was previously recorded in Central Sumatra and Nias (van Benthem Jutting 1959), West Sumatra (Maassen 2002 a), and Java (Nurinsiyah et al. 2023). The distribution in South Sumatra is a new record.</p><p>Remarks.</p><p>The specimens collected from the karst areas of South Sumatra exhibit two morphological forms (Fig. 8 a, b) that closely resemble Pupina treubi Boettger, 1890 and Pupina turgidula Dohrn, 1881 as illustrated by Maassen (2002 a). The South Sumatran samples also show strong shell similarity to the lectotype of P. treubi and the lectotype of P. verbeeki (Möllendorff, 1897) illustrated in Nurinsiyah et al. (2023). In that study, P. verbeeki was synonymized with P. treubi, and the authors concluded that all three nominal taxa represented the same species. This interpretation is further supported by our current material, in which all three morphotypes were collected from the same sampling plot. Following this conclusion, P. turgidula should be regarded as the valid name for this species.</p><p>The type specimen of P. turgidula from Mount Singgalang, West Sumatra, was destroyed during World War II, and our attempts to locate any remaining type material in the SMF and NHMW were unsuccessful. To conclusively resolve the taxonomic distinction between these taxa, fieldwork at the type locality is required. Additional specimens and further analyses, including molecular phylogenetic approaches, are necessary to robustly justify this taxonomic decision.</p></div>	https://treatment.plazi.org/id/D317C14533CC5251A9B9CF5BDEF0A270	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
24405F918B6E55B3B28B7652CD168F04.text	24405F918B6E55B3B28B7652CD168F04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stomacosmethis jagori (E. von Martens 1860)	<div><p>Stomacosmethis cf. jagori (E. von Martens, 1860)</p><p>Figs 5 a, 9 b</p><p>Alycaeus jagori E. von Martens, 1860: 208 .</p><p>Stomacosmethis jagori – Páll-Gergely et al. 2020: 191.</p><p>Material examined.</p><p>Indonesia • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 10)</a>; 4°04.11'S, 103°55.34'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24141/78 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92233&amp;materialsCitation.latitude=-4.0685" title="Search Plazi for locations around (long 103.92233/lat -4.0685)">Suka Merindu (Plot 11)</a>; 4°04.11'S, 103°55.34'E; alt. 124 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24138/450 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92333&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92333/lat -4.0676665)">Suka Merindu (Plot 8)</a>; 4°04.06'S, 103°55.40'E; alt. 134 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24128/78 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93433&amp;materialsCitation.latitude=-4.0723333" title="Search Plazi for locations around (long 103.93433/lat -4.0723333)">Padang Bindu (Plot 16)</a>; 4°04.34'S, 103°56.06'E; alt. 139 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24129/10 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92983&amp;materialsCitation.latitude=-4.0666666" title="Search Plazi for locations around (long 103.92983/lat -4.0666666)">Padang Bindu (Plot 2)</a>; 4°04.00'S, 103°55.79'E; alt. 158 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24125/11 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.9315&amp;materialsCitation.latitude=-4.0668335" title="Search Plazi for locations around (long 103.9315/lat -4.0668335)">Padang Bindu (Plot 4)</a>; 4°04.01'S, 103°55.89'E; alt. 111 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24126/7 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92533&amp;materialsCitation.latitude=-4.0676665" title="Search Plazi for locations around (long 103.92533/lat -4.0676665)">Suka Merindu (Plot 7)</a>; 4°04.06'S, 103°55.52'E; alt. 126 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24123/26 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.067333" title="Search Plazi for locations around (long 103.92267/lat -4.067333)">Suka Merindu (Plot 5)</a>; 4°04.04'S, 103°55.36'E; alt. 150 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24124/3, MZB Gst. 24139/23 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92834&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92834/lat -4.070667)">Padang Bindu (Plot 13)</a>; 4°04.24'S, 103°55.70'E; alt. 170 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24122/16 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93467&amp;materialsCitation.latitude=-4.0705" title="Search Plazi for locations around (long 103.93467/lat -4.0705)">Padang Bindu (Plot 3)</a> 4°04.23'S, 103°56.08'E; alt. 114 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24151/5 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92433&amp;materialsCitation.latitude=-4.0693336" title="Search Plazi for locations around (long 103.92433/lat -4.0693336)">Suka Merindu (Plot 12)</a>; 4°04.16'S, 103°55.46'E; alt. 154 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24149/2, MZB Gst. 24127/12 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92267&amp;materialsCitation.latitude=-4.0681667" title="Search Plazi for locations around (long 103.92267/lat -4.0681667)">Suka Merindu (Plot 9)</a>; 4°04.09'S, 103°55.36'E; alt. 122 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24142/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92667&amp;materialsCitation.latitude=-4.0721664" title="Search Plazi for locations around (long 103.92667/lat -4.0721664)">Padang Bindu (Plot 15)</a>; 4°04.33'S, 103°55.60'E; alt. 205 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24150/1 • South Sumatra, Ogan Komering Ulu, Semidang Aji, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.92917&amp;materialsCitation.latitude=-4.070667" title="Search Plazi for locations around (long 103.92917/lat -4.070667)">Padang Bindu (Plot 14)</a>; 4°04.24'S, 103°55.75'E; alt. 154 m; May-June 2021; A. S. Nurinsiyah leg.; MZB Gst. 24140/27 .</p><p>Description (n = 98).</p><p>Shell small size with H = 6.1–11.6 mm (mean 8.8 mm); D = 7.7–12.6 mm (mean 9.2 mm); HA = 3.4–5.6 mm (mean 4.2 mm); DA = 3.6–5.6 mm (mean 4.3 mm); 5–6 whorls; operculum height 2.7–3.6 mm, operculum width 2.6–3 mm. operculum concavity / thickness 0.61–0.78 mm. Shell dextral; conical shaped, with depressed apex; pale lime yellow to white, sub-transparent, the spire (protoconch) darker than the last whorl, smooth; teleoconch with fine and close radial ribs; spiral striae present; last whorl very large; constriction ~ 2–3 mm from the aperture; a small breathing tube (sutural tube) present; micro-tunnel under sutural tube; aperture almost perfectly round, slightly oblique at the top; peristome continuous, thick, and double; umbilicus open very small; operculum round, with concave rounded surface, convex inner surface, multispiral with central nucleus.</p><p>Habitat.</p><p>Limestone rocks.</p><p>Distribution.</p><p>The specimens collected in Sumatra are provisionally identified as Stomacosmethis jagori (see Remarks). The distribution in South Sumatra is a new record.</p><p>Remarks.</p><p>The genus Stomacosmethis has not been revised yet, and it is considered challenging, because the shell characters are relatively few. Namely, with a few exceptions, the shell shape and sculpture are relatively uniform across the genus, and the sutural tube (the length of which is an important character in other genera of this subfamily) is always very short. Several new species described by Foon and Liew (2017) are treated as subspecies in a later review (Páll-Gergely et al. 2020). Currently, 26 species are considered valid (Páll-Gergely et al. 2020; MolluscaBase 2025). The specimens we found in Sumatra are most similar to Stomacosmethis jagori (Fig. 5 b), which was originally described from Java, and was subsequently reported from Sulawesi and Borneo (van Benthem Jutting 1948; Hendriks et al. 2019), although the identity of the populations from Borneo are questionable (Páll-Gergely et al. 2020).</p><p>We note, however, that shells of Stomacosmethis kelantanensis clementsi (Foon &amp; Liew, 2017) (originally described as Alycaeus clementsi) from Peninsular Malaysia are also similar to our shells from Sumatra. However, they are probably not closely related due to the large geographic distance between them. Minor differences are observed between the Stomacosmethis kelantanensis clementsi shells and the ones we collected in both shell size and the sculpture of the radial ribs, with the Padang Bindu population displaying relatively larger shells and smoother radial ribs.</p></div>	https://treatment.plazi.org/id/24405F918B6E55B3B28B7652CD168F04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Aulia, Latifah Nurul;Nurinsiyah, Ayu Savitri;Mujiono, Nova;Páll-Gergely, Barna;Ambarwati, Reni	Aulia, Latifah Nurul, Nurinsiyah, Ayu Savitri, Mujiono, Nova, Páll-Gergely, Barna, Ambarwati, Reni (2026): Operculate land snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the description of a new species, Chamalycaeus dayangmerindu. ZooKeys 1272: 1-31, DOI: 10.3897/zookeys.1272.179378
