identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F12B8A1C7E53FFBB132AFB80FC86964D.text	F12B8A1C7E53FFBB132AFB80FC86964D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diodora J. E. Gray 1821	<div><p>Genus Diodora J.E. Gray, 1821</p><p>Type species</p><p>Patella apertura Montagu, 1803 by monotypy, junior synonym of Diodora graeca (Linnaeus, 1758) . Present day, Western Europe.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Knight et al. 1960) to present day.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E53FFBB132AFB80FC86964D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E53FFBA139EFA5CFE0A908A.text	F12B8A1C7E53FFBA139EFA5CFE0A908A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diodora undetermined	<div><p>Diodora sp.</p><p>Fig. 5A–D</p><p>Diodora sp. – Merle &amp; Pacaud 2018a: 169–170, fig. 155a–b. — Merle 2023: 25.</p><p>Material examined (3 specs)</p><p>FRANCE – Marne • 1 spec., internal imprint; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 155a) and herein Fig. 5A; MNHN.F.A66743 • 1 spec., internal mold; same data as for preceding; Ch. d’Orbigny coll., ex n°GG. Gg2005/6003; figured in Merle &amp; Pacaud (2018a: fig. 155b) and herein Fig. 5B–C; MNHN.F.C02614 • 1 spec., part of an external imprint; same data as for preceding; MMM coll.; Fig. 5D; MNHN.F.C02608 .</p><p>Measurements</p><p>MNHN.F. A66743, L: 34 mm; W: 25 mm. MNHN.F.C02614, L: 16 mm; W: 13 mm. MNHN.F.C02608, L (incomplete): 11 mm.</p><p>Description</p><p>Conical and moderately elevated limpet shell. Outline rounded at anterior and posterior edges, almost straight at mid point. Anal aperture approximately located at mid part of shell length, elongated, 1.8 mm long and rounded in both extremes. On internal imprint, crenulations made by axial ribs more marked on anterior part. On external imprint, delicate major ribs with three or four minor ribs intercalated in between. Surface weakly squamose.</p><p>Remarks</p><p>Sohl (1992) described two species of Diodora from the Late Cretaceous carbonate-platform deposits of the Caribbean: D. decussata Sohl, 1992 and D. hazeli Sohl, 1992 . Diodora sp. from the Paris Basin differs from D. decussata by having coarser radial ribs, and from D. hazeli by lacking prominent radial ribs. In the North European Maastrichtian, no specimens of Diodora have been identified. The studied material can be compared to Danian Diodora described from the Paris Basin in France (see Pacaud 2004) and the Mons limestone in Belgium (see Glibert 1973). Among them, Diodora sp. is reminiscent of Diodora corneti (Rutot in Cossmann, 1919) in its moderately elevated shells, its elongated posterior aperture, its shape rounded in both extremes and its delicate major ribs with three or four minor ribs intercalated in between. However, Diodora sp. differs from D. corneti by having a narrower and less rounded posterior edge and a longer and wider anterior edge (see holotype IRSNB n° 3023 in Glibert 1973: pl. 1 fig. 7).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E53FFBA139EFA5CFE0A908A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E52FFBA1095FC19FAF397C8.text	F12B8A1C7E52FFBA1095FC19FAF397C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trochida	<div><p>Order Trochida authority not given see MolluscaBase</p><p>Taxonomic comment</p><p>Most of the species of Trochoidea Rafinesque, 1815 and Seguenzoidea A.E. Verrill, 1884 described in this study are new species, but usually do not have a preserved aperture and their protoconchs are not visible. Generic assignments have had to be made on the basis of comparisons of shape and sculpture, which are not the best diagnostic characters. For this reason, these assignments are considered provisional and are preceded by a question mark until more material becomes available to confirm or refute them.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E52FFBA1095FC19FAF397C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E52FFBC13E5FA99FB35929C.text	F12B8A1C7E52FFBC13E5FA99FB35929C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trochoidea Rafinesque 1815	<div><p>Trochoidea indet.</p><p>Fig. 5E</p><p>? Jujubinus sp. – Merle &amp; Pacaud 2018a: 169–170, fig. 155g. — Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; figured in Merle &amp; Pacaud (2018a: fig. 155g) and herein Fig. 5E; MNHN.F.A66745 .</p><p>Measurements</p><p>H (incomplete apex): 4.2 mm; D: 4 mm.</p><p>Description</p><p>Two preserved whorls (penultimate and last whorls). Shell of small size; conical shape with fairly flat whorls. Suture shallow. Extreme abapical part of last whorl near base slightly convex. Penultimate whorl smooth (sculpture not preserved). Last whorl with three major cords more prominent at base.</p><p>Remarks</p><p>With its conical shape with fairly flat whorls, the fragment of shell is more reminiscent of members of Jujubinus, than species of Tectus Montfort, 1810 (Montfort 1810: 186), Calliomphalus Cossmann, 1888 (Cossmann 1888: 52) or Agathodonta Cossmann, 1918 (Cossmann 1918: 193) described in the present paper. Nevertheless, this fragment does not allow to give a reliable generic assigment.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E52FFBC13E5FA99FB35929C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E54FFBC130DFD8FFB3D9079.text	F12B8A1C7E54FFBC130DFD8FFB3D9079.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calliostoma Swainson 1840	<div><p>Genus Calliostoma Swainson, 1840</p><p>Type species</p><p>Trochus conulus Linnaeus, 1758 by subsequent designation (Herrmannsen 1846: 154). Present day, NE Atlantic and Mediterranean.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous ( Calliostoma s. lat.) according to Knight et al. (1960) to present day.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E54FFBC130DFD8FFB3D9079	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E54FFBF136CFC48FDBA9071.text	F12B8A1C7E54FFBF136CFC48FDBA9071.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calliostoma monswavinarensis Merle 2026	<div><p>? Calliostoma monswavinarensis sp. nov.</p><p>urn:lsid:zoobank.org:act: B6DCCF53-A6C0-4494-BEFF-A7EEB4202E66</p><p>Figs 5F, 6A–B</p><p>Etymology</p><p>From the ancient name of the locality Mont Aimé (‘Mons Wavinarum’ in the 7 th century).</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations of Mont Aimé, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02609.</p><p>Measurements</p><p>H (incomplete apex): 7 mm, D: 7.5 mm.</p><p>Description</p><p>Medium-sized shell. Conical and pyramidal spire. Spiral angle 54°. Four teleoconch whorls preserved, nearly concave and enlarged at base. Suture shallow. From first to last whorl, two major cords M1 and M2 (Fig. 6A–B). M1 prominent, smooth, at base of whorl, making a keel and delineating the periphery (Fig. 6A–B). M2 close set the anterior suture and weaker than M1 (Fig. 6A–B). Aperture not preserved.</p><p>Comparisons</p><p>? Calliostoma monswavinarensis sp. nov. is reminiscent of Trochus lineatus Binkhorst, 1861 from the Maastrichtian of The Netherlands (see Binkhorst 1861: pl. 5a1 fig. 9a–b) in its prominent M1 delineating the periphery. However,? C. monswavinarensis has fewer threads adapically and displays a narrower shape.? C. monswavinarensis also resembles Tectus carinodentatus (Briart &amp; Cornet, 1887) from the Danian of Mons in having a prominent M1 forming a keel delineating the periphery, but it differs by lacking crenulations on M1, fewer threads adapically and a narrower shape.? Calliostoma monswavinarensis and Tectus gabrielis (d’Orbigny, 1850) from the Danian of the Paris Basin have a prominent M1, but? C. monswavinarensis differs by having fewer threads adapically and by having a prominent M2. The generic attribution of? C. monswavinarensis is difficult to give. It cannot be easily attributed to Tectus Montfort, 1810, whose sculpture is more developed, particularly the peripheral cord, which bears numerous crenulations. The type species of Calliostoma, Calliostoma conulus (Linnaeus, 1758), present day, Mediterranean Sea, has a prominent cord delineating the periphery, but its early whorls bear four beaded cords. Moreover, species of Calliostoma are usually more ornamented than? C. monswavinarensis . A narrow shape, a strong peripherical cord at the base of whorls is also found in Jujubinus Monterosato, 1884, but as in Calliostoma, the species of Jujubinus have numerous other cords. In the absence of apertural characters, it is difficult to choose between Calliostoma, Tectus or Jujubinus . This is why Calliostoma is provisionally proposed with a question mark for this new species.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E54FFBF136CFC48FDBA9071	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E57FFBF1323FC13FDA79546.text	F12B8A1C7E57FFBF1323FC13FDA79546.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sparnollonia Merle 2026	<div><p>Genus Sparnollonia gen. nov.</p><p>urn:lsid:zoobank.org:act: 0F0B6F64-CADB-43C6-8111-F0CAA38FDAAF</p><p>Type species</p><p>Sparnollonia bicarinata gen et sp. nov., by present designation.</p><p>Included species</p><p>Sparnollonia bicarinata gen et sp. nov., Paris Basin, France, Maastrichtian.</p><p>Etymology</p><p>Name based from the ‘Sparnacus’ (old name of Epernay, France) and ‘Collonia’ using the letters ‘sparn-’ as prefix and ‘-ollonia’ as suffix. Genus female.</p><p>Diagnosis</p><p>Shell small with low spire. Whorls bicarinate. Spiral sculpture with three prominent major cords (M1–M3), M1: peripheric cord. Several threads between M1 and adapical suture and between M3 and umbilicus. Outer lip thin, slightly prosocline. Columella with a weak tubercle. Ombilicus very narrow. Columellar lip slightly adherent.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E57FFBF1323FC13FDA79546	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E57FFA01364F945FDBA9139.text	F12B8A1C7E57FFA01364F945FDBA9139.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sparnollonia bicarinata Merle 2026	<div><p>Sparnollonia bicarinata gen. et sp. nov.</p><p>urn:lsid:zoobank.org:act: B681154D-2AA0-45FD-A107-09FC558C3F8B</p><p>Figs 5G–K, 6C–D</p><p>? Afrollonia sp. – Merle &amp; Pacaud 2018a: 169–170, fig. 155i–j. — Merle 2023: 25.</p><p>Etymology</p><p>From ‘ bicarinatus ’ (Latin adjective) referring to the double carina formed by the major cords M1 and M2.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont); sandy limestone; MMM coll.; Figs 5G, 6D; MNHN.F.C02610.</p><p>Paratypes (13 specs) FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02611 • 1 spec.; same data as for holotype; MMM coll.; Figs 5I, 6C; MNHN.F.C02612 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02613 • 1 spec.; same data as for holotype; MMM coll.; Fig. 5H; MNHN.F.C02616 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02617 • 1 spec., same data as for holotype; MMM coll.; Fig. 5J; MNHN.F.C02618 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02619 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02620 • 1 spec., same data as for holotype; MMM coll.; Fig. 5K; MNHN.F.C02621 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02622 • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 155I); MNHN.F.A66746 • 1 spec.; same data as for preceding; Ch. d’Orbigny coll. (ex n°GG.gg2005/5999); figured in Merle &amp; Pacaud (2018a: fig. 155j); MNHN.F.C02615 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02623 .</p><p>Other examined material (5 specs)</p><p>FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02624 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02626 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02625 • 1 spec.; Bergères-les Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02627 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02628 .</p><p>Measurements</p><p>Holotype MNHN.F.C02610, H: 4.1 mm, D: 4.4 mm. Paratype MNHN.F. A66746, H: 4.6 mm, D: 4.7 mm. Paratype MNHN.F.C02615, H: 3.8 mm, D: 3.7 mm. Paratype MNHN.F.C02611, H: 4.2 mm, D: 4.8 mm. Paratype MNHN.F.C02612, H: 4.5 mm, D: 4.6 mm. Paratype MNHN.F.C02613, H 2.9 mm, D: 3.8 mm. Paratype MNHN.F.C02616, H: 4.2 mm, D: 4.2 mm. Paratype MNHN.F.C02617, H: 4.1 mm, D: incomplete. Paratype MNHN.F.C02618, H: incomplete, D: 4.9 mm. Paratype MNHN.F.C02620, H: incomplete, D: 3.9 mm. Paratype MNHN.F.C02621, H: incomplete, D: 4.6 mm. Paratype MNHN.F.C02622, H: 4.5 mm, D: 4.1 mm. Paratype MNHN.F.C02623, H 4.0 mm, D: 3.9 mm.</p><p>Description</p><p>Shell small, with low spire. Spiral angle 45°. Four teleoconch whorls; first and second whorls flat, third and fourth whorls conical, bicarinate. Suture shallow on three first whorls and moderately deep on last whorl. Spiral sculpture on first one not preserved. On second whorl, M1 smooth and prominent delineating the periphery. On third whorl, four to seven threads between adapical suture and M1; M1 and M2 smooth, prominent. Between M1 and M2, occasionnally one to three threads. Last whorl five to eight threads between adapical suture and M1; M1 and M2 prominent (Fig. 6C–D). Between M1 and M2, occasionnally one to three threads. On base M3 smooth, prominent yet slightly smaller than M2 and four weaker cords abapically. Base flat delineated by M3 (Fig. 6C–D). Aperture rounded. Outer lip thin, slightly prosocline. Columella with a weak tubercle. Columellar lip slightly adherent. Ombilicus very narrow.</p><p>Comparisons</p><p>This species was first attributed by Merle &amp; Pacaud (2018a) to? Afrollonia Adegoke, 1977 (type species: Turbo nigeriensis Adegoke, 1977 by original designation; Danian, Nigeria, see Fig. 5L), because of shell similarities (shell shape and sculpture) with Afrollonia vilcassina Pacaud, 2017 from the Danian of the Paris Basin. Afrollonia vilcassina differs in having more numerous threads between adapical suture and M1, by M1 and M2 being less prominent (Fig. 6E–F) and by having small axial ribs on first teleoconch whorls (see Pacaud 2017: pl. 2 fig. 1). Pacaud (2017: 187) stated that the base is deeply umbilicate and did not indicate a tubercle on the columella. In addition, the type species of Afrollonia differs from Sparnollonia bicarinata gen et sp. nov. in having a deep and moderate umbilicus, with a semilunar exterior outline and umbilical rim coarsely noded. Other Nigerian species of Afrollonia share these apertural characters which are frequently found in species of Margaritinae . The aperture of Sparnollonia bicarinata, is more reminiscent of that of the Collininae Otomphalus dumasi Cossmann, 1902 (Bartonian, Western France) in having a narrow umbilicus lacking a semilunar exterior outline. Moreover, its shell bears a low shape and two prominent M1–M2 as in S. bicarinata . Nevertheless, S. bicarinata differs from O. dumasi by having an M3, by lacking a columellar tubercle and by lacking a strong cord surrounding the umbilicus. Sparnollonia bicarinata and Otomphalus kleinoichingensis Traub, 1979 from the Thanetian (Oichinger Schichten) of Kleinoichinger Graben in Austria share M3, but M1–M2 are more prominent and wider apart in S. bicarinata . Cirsochilus Cossmann, 1888 (type species Delphinula striata Lamarck, 1804, accepted as Cirsochilus varius (Defrance, 1818), by original designation; Lutetian, Paris Basin) resembles S. bicarinata in its shape and its sculpture, but S. bicarinata differs in lacking a thickened outer lip and in having a tubercle on the columellar lip. Homalopoma P.P. Carpenter, 1864 (type species Turbo sanguineus Linnaeus, 1758 by monotypy; present day, Mediterranean Sea) has a more rounded shape, displays more numerous spiral cords, but some species bear a columellar tubercle as in S. bicarinata . Finally, Sparnollonia bicarinata cannot be classified without problems in apparently similar genera of Colloninae and for this reason we propose a new genus.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E57FFA01364F945FDBA9139	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E48FFA01327FD48FB059798.text	F12B8A1C7E48FFA01327FD48FB059798.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tectus Montfort 1810	<div><p>Genus Tectus Montfort, 1810</p><p>Type species</p><p>Tectus pagodalis Montfort, 1810, junior synonym of Trochus mauritianus Gmelin, 1791, type by original designation. Present day, Indian Ocean.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Knight et al. 1960) to present day.</p><p>Comments</p><p>The sculpture of fossil Tectus is quite variable. However, many species display at the base of the whorls a prominent peripheral cord (P4), often beaded, whilst the upper cords (P1–P3) are less developed. A study of numerous species of Tectus from the Lutetian of Normandy (Gain et al. 2019) shows that the P1 to P4 cords are frequently present. The three species described below also have flattened whorls, a flat base and bear the same cords (herein M1–M4, most likely corresponding to P1–P4). Due to these similarities, and pending observations of their aperture, they are provisionally assigned to Tectus .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E48FFA01327FD48FB059798	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E48FFA21326FB2EFD109019.text	F12B8A1C7E48FFA21326FB2EFD109019.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tectus acmantensis Merle 2026	<div><p>? Tectus acmantensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 20C5E291-1686-4758-B6D7-81D2DFFD2472</p><p>Figs 5M–O, 7A–C</p><p>Tectus montissanctipetri – Merle &amp; Pacaud 2018a: 169–170, fig. 155e–f. — Merle 2023: 25 (non Trochus montissanctipetri Binkhorst, 1861).</p><p>Etymology</p><p>From the ancient name of the locality Esmans (‘Acmantum’, 9 th century).</p><p>Type locality</p><p>Esmans (Bois d’Esmans), Seine-et-Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Esmans, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Seine-et-Marne • Esmans ( Bois d’Esmans); sandy limestone of Esmans; MNHN coll.; figured in Merle &amp; Pacaud (2018a: fig. 155f) and herein Figs 5M, 7A; MNHN.F.A66744.</p><p>Paratypes (2 specs) FRANCE – Seine-et-Marne • 1 spec.; same data as for holotype; Hébert coll.; Figs 5N, 7B; UPMC.PAL.2017.0.15.1a • 1 spec., historical imprint; same locality as for holotype; Hébert coll.; figured in Merle &amp; Pacaud (2018a: fig. 155e) and herein Figs 5O, 7C; UPMC.PAL.2017.0.15.1b .</p><p>Measurements</p><p>Holotype MNHN.F. A66744, H (incomplete apex): 19 mm, D: 14 mm. Paratype UPMC.PAL.2017.0.15.1a, H (incomplete apex): 19 mm, D: 14 mm. Paratype UPMC.PAL.2017.0.15.1b, H (incomplete apex): 18 mm, D: 16 mm.</p><p>Description</p><p>Medium-sized shell. Conical and pyramidal spire. Spiral angle between 40° and 46°. Six teleoconch whorls preserved, nearly concave, enlarged and very angulose at base. Suture incised, moderately deep. On paratype UPMC.PAL.2017.0.15.1bis, spiral sculpture of first preserved whorl consisting of three equally-sized cords (M1 to M3); fourth cord (M4) making periphery at base of whorl, prominent and protruding. Second to sixth whorls: first cord (M1) close set to adapical suture; M2 and M3 at the centre of whorl; M4 close set to anterior suture. M1 strongly beaded, M2 to M4 less beaded (Fig. 7A–C). On cords, between 12 and 16 fairly rounded, closely spaced beads and more prominent adapically. Base flat, apparently smooth. Aperture not preserved.</p><p>Comparisons</p><p>These three specimens of? Tectus acmantensis sp. nov. were first attributed to? Tectus montissanctipetri (Binkhorst, 1861) from the Maastrichtian of Limburg, because of a prominent peripheral cord (M1) at the base of the whorls and finer cords adapically. However, a more detailed comparison with the figure by Binkhorst (1861: pl. 5a1 fig. 10b–c and Fig. 7E herein) shows that the specimens from the Paris Basin bear narrower whorls, major cords (M1 to M3) with coarser beads and a protruding M4 that clearly delineates the periphery by forming an acute angle at the base of the whorls (Fig. 7A–C). Moreover,? T. montissanctipetri differs in having m1 and m2, threads between M2 and M3, a prominent M3 delineating the periphery and a smaller M4 close set the anterior suture. From the figure of Binkhorst (1861: pl. 3 fig. 6a–b),? Tectus detritus (Binkhorst, 1861) resembles? T. acmantensis in having M1–M3 above the peripheric cord M4. However, M2 is more prominent than M1 and M 3 in? T detritus, whereas they are subequal in? T. acmantensis . In addition,? T. detritus differs in having a strongly beaded M4, a smooth M6 and an M5 close set the anterior suture.</p><p>Stratigraphic range</p><p>Maastrichtian: Seine-et-Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E48FFA21326FB2EFD109019	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4AFFA4133FFCAFFDBA919C.text	F12B8A1C7E4AFFA4133FFCAFFDBA919C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tectus fraterculus Merle 2026	<div><p>? Tectus fraterculus sp. nov.</p><p>urn:lsid:zoobank.org:act: E0991DA6-AFD4-4EA4-91B6-AF0857E93343</p><p>Figs 7D, 8A–D</p><p>Etymology</p><p>From the Latin (‘ frater ’ meaning ‘brother’) and the suffix (‘ culus ’) to recall? T. acmantensis sp. nov. from the Maastrichtian of Bois d’Esmans (Seine-et-Marne) with which it shares several similarities.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont); sandy limestone; MMM coll.; Figs 7D, 8A; MNHN.F.C02629.</p><p>Paratype FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; Fig. 8B–D; MNHN.F.C02630 .</p><p>Measurements</p><p>Holotype: MNHN.F.C02629, H (incomplete apex): 9 mm, D: 9 mm. Paratype MNHN.F. C02630, H (incomplete apex): 5.5 mm, D: 7.5 mm.</p><p>Description</p><p>Small sized shell. Conical spire. Spiral angle 46°. Two teleoconch whorls preserved, almost straight, enlarged at base. Suture shallow. On holotype, spiral sculpture consisting of four cords (M1 to M4); M1 beaded, prominent and close set the suture; M2 beaded, weaker than M1; M3 weaker than M2 and M4 and with weak beads; M4 more prominent that M3, almost smooth and making periphery at base of whorl (Fig. 7D). On last whorl of holotype, 17–18 fairly rounded and closely spaced beads, on M1 and M2. Base flat, apparently smooth. Aperture not preserved.</p><p>Comparisons</p><p>? Tectus fraterculus sp. nov. is reminiscent of? T. acmantensis sp. nov. in its spiral sculpture of fourth major cords (M1 to M4). However, its size is smaller (H: 9 mm instead of 18–19 mm), its apical angle larger (46° instead of 40°) and its suture shallower. Moreover, there are some differences in the spiral sculpture: M4 is not protuding as it is in? T. acmantensis, the beads are more prominent on M1 and M2 and more numerous and larger (17–18 instead of 12–16 in? T. acmantensis).? Tectus fraterculus resembles? T. montissanctipetri, but differs in lacking m1 and m2, threads present between M2 and M3. It bears a weaker M3 delineating the periphery and a prominent M4 close set the anterior suture (see Fig. 7E 2).</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4AFFA4133FFCAFFDBA919C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4CFFA71339FD13FD1092D9.text	F12B8A1C7E4CFFA71339FD13FD1092D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tectus senarmonti Merle 2026	<div><p>? Tectus senarmonti sp. nov.</p><p>urn:lsid:zoobank.org:act: BB8F18D5-54BA-43D0-8FFB-D559E184AB05</p><p>Figs 8E–H, 9A</p><p>Etymology</p><p>Dedicated to Alexandre-Antoine Hureau, baron de Sénarmont who briefly described the sandy limestone of Bois d’Esmans.</p><p>Type locality</p><p>Esmans (Bois d’Esmans), Seine-et-Marne, France. The label of the sample indicates Montereau as the locality, but it is most likely the locality of Bois d’Esmans at Esmans.</p><p>Type horizon</p><p>Sandy limestone of Esmans, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Seine-et-Marne • Esmans ( Bois d’Esmans); sandy limestone; Munier-Chalmas coll.; SU.PAL.2017.0.2.404.</p><p>Measurements</p><p>SU.PAL.2017.0.2.404: H (incomplete base): 16 mm, probably 20 mm; D: 34 mm.</p><p>Description</p><p>Large-sized shell. Conical and pyramidal spire, with large base. Spiral angle 56°. Three teleoconch whorls preserved, nearly concave and enlarged at periphery. Periphery rather thick, slightly rounded. Suture shallow. Spiral sculpture consisting of five equally-sized spiral cords (M1 to M5) from the adapical suture of whorl until adapical top of periphery; periphery with two or three cords (M6–M7 or M6–M8) close set the abapical suture. M1–M5 delicately beaded, M6 to M8 less beaded (Fig. 9A). Between 16 and 20 small and spirally elongate beads on cords. Base flat, apparently smooth. Aperture not preserved.</p><p>Comparisons</p><p>? Tectus senarmonti sp. nov. has a much wider shape and a lower spire than? Tectus acmantentis sp. nov. It cannot be considered a gerontological specimen of? T. acmantensis, because it has a very different spiral sculpture. First,? T. senarmonti bears delicately beaded M1–M5, whereas? T. acmantensis bears only three strongly beaded major cords (M1–M3), and second,? T. senarmonti has two-three cords (M6– M8) making a rounded periphery whereas? T. acmantensis has a prominent M4 making a very angulose periphery at the base of the whorl.</p><p>Stratigraphic range</p><p>Maastrichtian: Seine-et-Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4CFFA71339FD13FD1092D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4FFFA71377FD60FB61972F.text	F12B8A1C7E4FFFA71377FD60FB61972F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agathodonta Cossmann 1918	<div><p>Genus Agathodonta Cossmann, 1918</p><p>Agnathodonta Wenz, 1938: 298, error of spelling for Agathodonta Cossmann (1918) .</p><p>Type species</p><p>Trochus dentigerus d’Orbigny, 1842 by original designation. Early Cretaceous (Hauterivian), Aube, France.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Hauterivian) according to d’Orbigny (1842) to?Maastrichtian (this paper).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4FFFA71377FD60FB61972F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4FFFA610F1FBFEFCFC97C4.text	F12B8A1C7E4FFFA610F1FBFEFCFC97C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agathodonta granosecincta (Binkhorst 1861) Merle 2026	<div><p>? Agathodonta granosecincta (Binkhorst, 1861) comb. nov.</p><p>Figs 8I, 9B</p><p>Turbo granose-cinctus Binkhorst, 1861: 46, pl. 5a1 fig. 6a–b.</p><p>Danilia strombecki – Merle &amp; Pacaud 2018a: 169–170, fig. 155h. — Merle 2023: 25 (non Turbo strombecki Binkhorst, 1861).</p><p>Type material (not examined)</p><p>Problably lost (John Jagt pers. com.).</p><p>Type locality</p><p>Maastricht or Fauquemont, cf. Binkhorst (1861: iv), The Netherlands; Meerssen Member, Maastrichtian.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 155h) and herein Figs 8I, 9B; MNHN.F.R63293 .</p><p>Measurements</p><p>H (incomplete apex): 13 mm, D: 12 mm.</p><p>Description</p><p>Medium-sized shell. Penultimate and last whorls preserved. Convex whorls delineated by a moderately deep incised suture. Base convex. Sculpture of spiral cords crossed by axial riblets, cords ornamented with small and rounded beads. On penultimate whorl, five major cords (M1–M5); two weak cords adapically (M1–M2), two prominent cords in centre of whorl (M3–M4) and a weaker cord close set abapical suture (M5), see Fig. 9B. Cords ornamented with around 20 beads. On last whorl, most adapical cord prominent (M1), three weaker cords abapically (m1+M2+m2), three prominent cords in the centre of whorl (M3–M5) and around five weaker cords until base (M6–M10), see Fig. 9B. All spiral cords ornamented with around 30 small and rounded beads. Axial riblets almost orthocline and connected to the beads. Aperture not preserved.</p><p>Remarks</p><p>This specimen from the Paris Basin was previously attributed to Turbo strombecki Binkhorst, 1861 from the Maastrichtian of Limburg (The Netherlands) by Merle &amp; Pacaud (2018a). The beaded spiral sculpture is quite similar, but the holotype of Turbo strombecki has less convex whorls, a higher spire, and according to Binkhorst’s illustration (Binkhorst 1861: pl. 5a fig. 6b) its sculpture lacks minor cords in the adapical part of the whorls. The specimen from the Paris Basin is more strongly reminiscent of Turbo granosecinctus Binkhorst, 1861 (Binkhorst 1861: pl. 5a1 fig. 6a–b), from the Maastrichtian of Limburg (The Netherlands), in its very rounded whorls, its incised suture, its sculpture of spiral cords crossed by axial riblets, its cords ornamented with small and rounded beads and its prominent M1 (see Fig. 9B–C). Moreover, it displays minor (m1, m2 and m3) in the apical part of the whorl, as observed in the specimen from Maastricht. It therefore seems more appropriate to attribute it to T. granosecinctus . Merle &amp; Pacaud (2018a) included T. strombecki in the genus Danilia Brusina, 1865 (type species: Olivia otaviana Cantraine, 1835 by monotypy). However, species of Danilia display less rounded whorls and coarser beaded spiral cords. As stated by Herbert (2012: 399), strongly convex whorls and spiral cords finely and regularly beaded are found in the type species Agathodonta, A. dentigera (d’Orbigny, 1842), from the Hauterivian of France, and not in the other genera of Chilodontidae . This sculpture is also shared with T. granosecinctus . The type species of Agathodonta only differs by having varices on spire whorls and prosocline riblets. Therefore, T. granosecinctus is provisionally assigned to? Agathodonta pending a validation by the examination of the aperture.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France; Limburg, The Netherlands.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4FFFA610F1FBFEFCFC97C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4EFFA61304FACAFC9D96DD.text	F12B8A1C7E4EFFA61304FACAFC9D96DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clypeostoma Herbert 2012	<div><p>Genus Clypeostoma Herbert, 2012</p><p>Type species</p><p>Turcica salpinx Barnard, 1964 by origninal designation. Present day: South Africa.</p><p>Stratigraphic range of the genus</p><p>Present day, South Africa (fossil range unknown).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4EFFA61304FACAFC9D96DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E4EFFA91304F9D3FDBA96D4.text	F12B8A1C7E4EFFA91304F9D3FDBA96D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clypeostoma vertusensis Merle 2026	<div><p>? Clypeostoma vertusensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 4007445E-2C73-4388-971C-E0B4CECFD5A9</p><p>Figs 8J–K, 9D</p><p>Etymology</p><p>From Vertus (Marne), the name of the type locality.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont); sandy limestone; MMM coll.; Figs 8K, 9D; MNHN.F.C02631.</p><p>Paratype FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; Fig. 8J; MNHN.F.C02632 .</p><p>Measurements</p><p>Holotype, H (incomplete base): 8 mm, D: 7.1 mm. Paratype, H: 6.1 mm; D: 5 mm.</p><p>Description</p><p>Medium-sized shell. Conical spire, base large. Spiral angle between 62° (paratype) and 67° (holotype). Three preserved teleoconch whorls, slightly convex and almost straight. Periphery not thickned at base of whorls. Suture deep and canaliculate. Base convex. Spiral cords well developed and crossed by prosocline axial riblets. Cords ornamented with beads small and slightly pointy at their top. Space between cords deep. On spire whorls, five subegal major cords (M1–M5) and one minor cord (m1) between M1 and M2 (Fig. 9D). M5 delineating the periphery. Cords ornamented with around twenty beads. On last whorl, 13 cords from suture to base. Cords ornamented with around 25 small beads. Aperture not preserved.</p><p>Comparisons</p><p>The sculpture of? Clypeostoma vertusensis sp. nov. is reminiscent of that of? A. granosecincta comb. nov. in its cords of small beads crossed by axial riblets. However,? C. vertusensis differs from? A. granosecincta by having a conical spire with a weakly convex whorl, less numerous minor spiral cords (only m1 instead of m2–m3) and prosocline axial riblets (see Figs 8I–K, 9B–D).? Clypeostoma vertusensis is distinguished from Turbo rimosus Binkhorst, 1861 (Binkhorst 1861: pl. 5a1 fig. 5a–b) from the Maastrichtian of The Netherlands in having a narrower shape, coarser spiral ridges and prosocline axial riblets. Conical whorls with a canaliculate suture and spiral cords which, where they intersect prosoclinic riblets, form rows of small beads that are slightly pointed at their tips, a featured shared with? C. vertusensis and members of Clypeostoma (Herbert 2012) . However, it is difficult to be certain of this identification due to the lack of apertural characters and thus this genus assigment is provisionally proposed.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E4EFFA91304F9D3FDBA96D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E41FFA91378F994FE8D9422.text	F12B8A1C7E41FFA91378F994FE8D9422.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calliomphalus Cossmann 1888	<div><p>Genus Calliomphalus Cossmann, 1888</p><p>Type species</p><p>Turbo squamulosus Lamarck, 1804 by original designation. Middle Eocene (Lutetian), Paris Basin, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E41FFA91378F994FE8D9422	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E41FFA8130BF8E1FCB59250.text	F12B8A1C7E41FFA8130BF8E1FCB59250.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calliomphalus (Planolateralus) Sohl 1961	<div><p>Subgenus Planolateralus Sohl, 1961</p><p>Type species</p><p>Calliomphalus argenteus Wade, 1926, by original designation, Late Cretaceous (Maastrichtian), Ripley Formation, Mississippi (USA).</p><p>Stratigraphic range of the subgenus</p><p>Late Cretaceous (Maastrichtian) of Mississippi (USA) according to Wade (1926) to Paleocene (Danian) of Paris Basin according to Merle &amp; Pacaud (2018b).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E41FFA8130BF8E1FCB59250	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E40FFAB10A3FE57FDBA9033.text	F12B8A1C7E40FFAB10A3FE57FDBA9033.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calliomphalus (Planolateralus) montenati Merle 2026	<div><p>Calliomphalus (Planolateralus) montenati sp. nov.</p><p>urn:lsid:zoobank.org:act: 07EC388F-C859-4D33-9EBA-FC2A5CC87B47</p><p>Figs 8L–M, 9E–F</p><p>Calliomphalus sp. – Merle &amp; Pacaud 2018a: 169–170, fig. 155c–d. — Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to the French geologist Christian Montenat, who worked on the Maastrichtian of the Paris Basin.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; Ch. d’Orbigny coll., ex n° GG.gg2007/16527; figured in Merle &amp; Pacaud (2018a: fig. 155d) and herein Figs 8L, 9E; MNHN.F.C02633.</p><p>Paratype FRANCE – Marne • 1 spec., historical cast of the imprint; Vertus ( Faloise); Hébert coll.; figured in Merle &amp; Pacaud (2018a: fig. 155c) and herein Figs 8M, 9F; UPMC.PAL.2017.0.14.1 .</p><p>Measurements</p><p>Holotype, H 8.2 mm, D: 6.2 mm. Paratype UPMC.PAL.2017.0.14.1, H: 12.5 mm, D: 9 mm.</p><p>Description</p><p>Medium-sized shell with conical shape. Five teleoconch whorls becoming increasingly convex with growth. Last whorl 40% of total height. Suture slightly incised. Spiral cords ornamented with rounded beads. Based on preseverd part of holotype (Fig. 9E): first spire whorl, with poorly visible beads; second whorl: three beaded (M1 to M3) cords (between 10 and 13 small beads per cord) and abapically two larger beaded (M4–M5) cords (15–16 beads per cord). Second whorl (penultimate): three (M1–M3) beaded cords (between 9 and 12 small beads per cord), abapically two (M4–M5) larger cords (between 11 and 16 beads per cord) and one smaller cord (M6) close set to the suture; M4 larger than the others and delineating the periphery. Last whorls: three (M1–M3) small beaded cords (between 11 and 16 beads per cord), M1 slightly larger that M1–M2 and abapically four (M4–M7) larger beaded cords (between 15 and 25 beads per cord); M4 larger than the other major cord and delineating the periphery. Seventh cord (M7) delineating the base. Aperture not preserved.</p><p>Comparisons</p><p>A conical shape with weakly convex whorls, an incised suture, several (three to five) adapical cords of small beads and cords of more prominent beads placed abapically are typical of Calliomphalus (Planolateralus) . The type species, Calliomphaus (Planolateralus) argenteus, displays five cords of small beads adapically whereas C. (P.) montenati sp. nov. bears three and has two prominent cords of beads or hollow spines delineating the periphery. Moreover, its spire is lower, particularly in specimens from the Mexcala Formation, Mexico (Perrilliat et al. 2000). Calliomphalus (Planolateralus) montenati displays a lower spire and less elevated whorls than C. (P.) cypliensis Vincent, 1930, from the Danian of Ciply (Belgium). Its spire in C. (P.) montenati bears seven beaded cords instead of only four in C. (P.) cypliensis . In addition, the beads of C. (P.) cypliensis are arranged in oblique series, whereas they are not in C. (P.) montenati . Calliomphalus (Planolateralus) gravesii (d’Orbigny, 1850), from the Danian of the Paris Basin, differs in having cords of more or less subequal beads that increase in size towards the base of the whorls (see Pacaud 2018a: fig. 225b–f). Finally, Calliomphalus (Planolateralus) montenati is easily distinguished from C. (P.) firketi from the Danian of the Mons Basin (Briart &amp; Cornet 1887) by stronger cords (see Glibert 1973: fig. 22).</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E40FFAB10A3FE57FDBA9033	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E42FFAA130DFE1BFEE9903C.text	F12B8A1C7E42FFAA130DFE1BFEE9903C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinoclavis Swainson 1840	<div><p>Genus Rhinoclavis Swainson, 1840</p><p>Type species</p><p>Murex vertagus Linnaeus, 1767 by subsequent designation (Herrmannsen 1848). Present day, Indo-Pacific.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian) of Maastricht (see below) and Paris Basin (this work) to present day in tropical waters.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E42FFAA130DFE1BFEE9903C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E42FFAC1307FCF2FDBA9753.text	F12B8A1C7E42FFAC1307FCF2FDBA9753.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinoclavis parvisulcatus Merle 2026	<div><p>Rhinoclavis parvisulcatus sp. nov.</p><p>urn:lsid:zoobank.org:act: 9FB955B3-9A14-4DC4-A3E7-0E3D646491D2</p><p>Figs 10A–C, 11A–F</p><p>Rhinoclavis (Semivertagus) sp. – Merle &amp; Pacaud 2018a: 169, 171, fig. 156d–e. — Merle 2023: 25.</p><p>Etymology</p><p>From the latin adjectives ‘ parvus ’ (meaning ‘small’) and ‘ sulcatus ’ (meaning ‘grooved’) to refer the small spiral groove of the species.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; Figs 10C, 11A; MNHN.F.C02634.</p><p>Paratypes (9 specs) FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; Fig. 11D; MNHN.F.C02635 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02636 • 1 spec.; same data as for holotype; MMM coll.; Figs 10B, 11B; MNHN.F.C02637 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02638 • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156d); MNHN.F.A66752 • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156e1–e2b) and herein Fig. 11E–F; MNHN.F.A66753 • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156d); MNHN.F.C02639 • 1 spec.; Vertus ( Grand Mont); sandy limestone; MMM coll.; Figs 10A, 11C; MNHN.F. C02639 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02640 .</p><p>Measurements</p><p>Holotype: H (base incomplete), 11.2 mm, D: 4.6 mm. Paratype MNHN.F. C02635, H (base incomplete): 10.3 mm, D: 4.5 mm. Paratype MNHN.F.C02636, H (base incomplete): 9 mm, D: 3.5 mm. Paratype MNHN.F.C02637: H (base incomplete), 6.8 mm, D: 3.2 mm. Paratype MNHN.F.C02638, H: 10.7 mm, D: 3.6 mm. Paratype MNHN.F. A66753, H: 14.4 mm, D: 5.7 mm. Paratype MNHN.F.C02639, H: 10.6 mm, D: 4.1 mm. Paratype MNHN.F.C02640, H (base incomplete): 8 mm, D: 3.1 mm.</p><p>Description</p><p>Shell medium in size, fairly pupate with convex apex. Around height-nine moderately convex whorls. Apical angle between 32° and 44°. Suture linear, shallow. Height of last whorl: approximately of 37% of shell length (based on paratype MNHN.F.C02638). No axial sculpture. Spire whorls notched by one moderately small spiral groove located halfway up apical part of whorls (about 45% of the height of whorl starting from adapical suture). On last whorl, spiral groove located in adapical third of whorl. On spire and in adapical part of whorl located between spiral groove and adapical suture, spiral sculpture varying from flat threads to three very flat cords. On spire and in abapical part of whorl located between spiral groove and abapical suture, surface of four small shallow grooves. On last whorl, similar spiral sculpture in part located between spiral groove and the adapical suture; from spiral groove to base of whorl, sculpture varying from flat threads to six flat cords. Rhinoclavis -type aperture with smooth and well detached columellar lip.</p><p>Comparisons</p><p>Rhinoclavis parvisulcatus sp. nov. has a variable spiral sculpture ranging from flat threads to small flat spiral ridges (Fig. 10A–C). However, it is clearly characterised by a spiral groove located approximately halfway around the shell. This sculpture and the shape of its aperture are found in R. unisulcatus (Lamarck, 1804) from the Eocene of the Paris Basin. The first specimens of R. parvisulcatus illustrated by Merle &amp; Pacaud (2018a: fig. 156d–e) as Rhinoclavis sp. bear spiral threads, but additionnal specimens demonstrate that the species can be more ornamented with small flat cords (Fig. 10B–C). Among Paleocene species, R. (S.) parvisulcatus is comparable to R. urania d’Orbigny (1850) from the Danian of the Paris and Mons basins (Glibert 1973). Rhinoclavis parvisulcatus differs from R. urania in having a wider shape, a shallower spiral groove and a coarser spiral sculpture (see the syntypes of R. (S.) urania illustrated by Pacaud (2018a: 249, fig. 196g –h and herein Fig. 10D). In the Danian of Mons, Glibert (1973) cited four other species: R. gibbosus (Briart &amp; Cornet, 1873), R. abnormis (Briart &amp; Cornet, 1873), R. elisae (Briart &amp; Cornet, 1873) and R. edmondi (Briart &amp; Cornet, 1873) . These four species display a shorter shape than R. parvisulcatus and, except R. gibbosus, a spiral groove is not clearly visible. The genus is still not reported in the Late Cretaceous, but Kaunhowen (1898: 48, pl. 4 fig. 4aa) described Turritella egregia Kaunhowen, 1898 from the Bryozoen-Schichten of Maastricht. The shape of the illustrated specimen bears a convex apex, a spiral sculpture with flat threads and whorls bearing a spiral groove which are typical characters of Rhinoclavis and not those of Turritellidae . Kaunhowen (1898: 48) mentioned clearly this groove in his description: “Die Nähte sind und können leicht mit einer Furche verwechselt werden, welche von jedem Umgange hinten einen Gürtel abtrennt, dessen Höhe ungefähr ein Dritttheil bis die Hälfte von der einer jeden Windung ausmacht” [“The sutures are and can easily be confused with a groove that separates a belt from each whorl at the back, the height of which is approximately one-third to one-half of that of each whorl”]. Rhinoclavis egregius comb. nov. differs from R. parvisulcatus by having a weaker sculpture of flat threads and by a slenderer shape. It is the first record of the genus Rhinoclavis in the Maastrichtian.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E42FFAC1307FCF2FDBA9753	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E44FFAC1376FB13FD5796D9.text	F12B8A1C7E44FFAC1376FB13FD5796D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoaluco B. L. Clark 1946	<div><p>Genus Pseudoaluco B.L. Clark, 1946</p><p>Type species</p><p>Cerithium jussieui Mayer-Eymar, 1876 by original designation. Middle Eocene (Lutetian), Paris Basin, France.</p><p>Stratigraphic range of the genus</p><p>Maastrichtian (this paper) to Middle Eocene.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E44FFAC1376FB13FD5796D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E44FF9113FDF9EFFDBA90D1.text	F12B8A1C7E44FF9113FDF9EFFDBA90D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoaluco undetermined	<div><p>Pseudoaluco sp.</p><p>Figs 11G – 12A</p><p>Pseudoaluco sp. – Merle &amp; Pacaud 2018a: 171, fig. 156f. — Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156f); MNHN.F.A66755 .</p><p>Measurements</p><p>H (two last whorls preserved): 11 mm, D: 7 mm.</p><p>Description</p><p>Fragment of shell corresponding to two preserved teleoconch whorls in ventral face: penultimate and last whorls. Penultimate whorl slightly convex; last whorl slightly impressed beneath suture with convex base. Suture linear, shallow. Axial sculpture consisting of 10–12 low and slightly opisthocyrt riblets. On penultimate whorl, a prominent rib and on last whorl another prominent rib in abapertural side of shell. On penultimate whorl, four subequal beaded major cords (M1–M4); on last whorl, four subequal beaded major cords from suture to centre of whorl and abapically several weaker and poorly visible cords (Fig. 12A).Aperture ovate. Columella twisted. Columellar lip not visible. Siphonal canal medium, reflexed to left of shell axis. Posterior canal indistinct. Outer lip thin.</p><p>Remarks</p><p>In agreement with Merle &amp; Pacaud (2018a), this shell fragment is more related to Pseudoaluco than to other genera of Cerithioidea . It is comparable to P. carolinum (d’Orbigny, 1850) from the Danian of the Paris Basin. However, P. carolinum differs by having more cords (M1–M5, m1, m2–m3), more prominent ribs and by lacking beaded cords except on M1. Pseudolacuco dejaeri (Briart &amp; Cornet, 1873), from the Danian of Mons (Belgium), figured by Glibert (1973: pl. 6 fig. 10, spec. IRSNB n° 5356), displays a detached columellar lip and a posterior canal, lacking in Pseudoaluco sp. However, Pseudoaluco sp. is probably juvenile, which may explain why these characters are not yet present. The sculpture of Pseudoaluco sp. also resembles that of the Maastricht specimen of Cerithium pseudoclathratum d’Orbigny, 1850, illustrated by Kauenhowen (1897: pl. 11 fig. 12). However, in that illustration, the prominent ribs on the penultimate and last whorl of Pseudoaluco sp. are not visible and Kaunhowen (1898: 65) makes no mention of them in his description. An examination of the materiel of Kaunhowen would be necessary to explore this comparison in greater depth.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E44FF9113FDF9EFFDBA90D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E79FF91137EFBF0FBC69617.text	F12B8A1C7E79FF91137EFBF0FBC69617.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vicinocerithium E. Wood 1910	<div><p>Genus Vicinocerithium E. Wood, 1910</p><p>Type species</p><p>Vicinocerithium parallelum E. Wood, 1910 (junior synonym of Cerithium bouei Deshayes, 1833) by original designation. Middle Eocene (Bartonian), Paris Basin, France.</p><p>Stratigraphic range of the genus</p><p>?Late Cretaceous (Maastrichtian) of Paris Basin (this paper) to Middle Eocene.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E79FF91137EFBF0FBC69617	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E79FF931318FA94FB369099.text	F12B8A1C7E79FF931318FA94FB369099.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vicinocerithium heberti Merle 2026	<div><p>? Vicinocerithium heberti sp. nov.</p><p>urn:lsid:zoobank.org:act: D4BC61B9-13DB-4244-9B89-2B7BE551CE97</p><p>Figs 11H–J, 12B–C</p><p>Cerithium carolinum d’Orbigny, 1850: 129, n° 19, pars, only the syntype of Mont Aimé.</p><p>? Cerithium carolinum – Combes 1908: 14 (non Cerithium carolinum d’Orbigny, 1850).</p><p>Vicinocerithium sp. 1 – Merle &amp; Pacaud 2018a: 169–171 fig. 155k–m, 156a–b. — Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to Edmond Hébert, French geologist of the 19 th century who collected Maastrichtian material from this species in the Paris Basin.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156b) and herein Figs 11I, 12C; MNHN.F.A66750.</p><p>Paratypes (5 specs) FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 155k); MNHN.F.A66747 • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 155m); MNHN.F.A66748 • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156a) and herein Fig. 11H; MNHN.F.A66749 • 1 spec.; same data as for holotype; MMM coll.; Figs 11J, 12B; MNHN.F.C02641 • 1 spec., historical cast of the imprint; Vertus ( Faloise); Hébert coll.; figured in Merle &amp; Pacaud (2018a: fig. 155l); UPMC.PAL.2017.0.16.1 .</p><p>Other examined material (9 specs)</p><p>FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02642 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02643 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02644 • 1 spec.; Vertus ( Grand Mont); sandy limestone; MMM coll.; MNHN.F.C02645 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02646 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C0264 • 2 specs; same data as for preceding; MMM coll.; MNHN.F.C02647 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02648. Many fragments of this species are not listed in the Examined material .</p><p>Measurements</p><p>Holotype, H: 23 mm, D: 11 mm. Other material broken.</p><p>Description</p><p>Shell medium in size, elongate and turreted. Protoconch unknown. Around eleven slightly convex whorls. Apical angle 34°. Suture shallow. Height of last whorl: 39% of shell length (based on holotype). Axial sculpture consisting of moderately strong and rather straight ribs. 10–12 ribs on spire and 13–14 ribs on last whorl. On first five whorls, three major cords (M1–M3) thicker in abapical direction. On later whorls three major cords (M1–M3) and a minor cord (m1). Last whorl with five major cords (M1–M5), one minor cord m1 and occasionally m4 and m5 (Fig. 7B–C). Abapical M4–M5 weaker than the adapical major cords. Major cords swelling on axial ribs and forming acute beads at intersection between axial and spiral sculptures. Aperture ovate. Outer lip not preserved. Columellar lip smooth, detached.</p><p>Comparisons d’Orbigny (1850) described Cerithium carolinum from the Danian by introducing material from Bergères-lès-Vertus (Mont Aimé), Meudon (Hauts-de-Seine, Danian) and Montainville (Yvelines, Danian) in the type series. Comparison of specimens from Mont Aimé with those from the two cited Danian localities shows that these are two different species (Pacaud 2018a; Merle &amp; Pacaud 2018a, 2018b). In order to stabilise the nomenclature, a lectotype was designated from a Danian specimen of C. carolinum from Montainville (Pacaud 2018a: 240, 195e). Furthermore, the sculptural features C. carolinum are those of Pseudoaluco and therefore C. carolinum is attributed to this genus.? Vicinocerithium heberti sp. nov. bears similarities with Cirsocerithium Cossmann, 1906 (type species: Cerithium subspinosum Leymerie, 1842 by original designation; Albian, Paris Basin). Compared to C. subspinosum,? V. heberti has a more elongate spire, a less rounded aperture and lacks a prominent, spiny spiral cord. Cirsocerithium collignoni Kiel, 2006 from the Albian of Madagascar displays a more comparable sculpture lacking a prominent, spiny spiral cord, but it bears a shorter spire and a more rounded aperture than in? V. heberti . Cirsocerithium crassilabris (von Koenen, 1885) from the Danian of Copenhagen also differs from? V. heberti by having a shorter spire and a more rounded aperture. The sculpture of? Vicinocerithium heberti is more reminiscent of that of Cerithium inopinatum Deshayes, 1864 (Deshayes 1864: pl. 82 fig. 32) reported and illustrated from the Danian of Mons by Glibert (1973: 43, pl. 5 fig. 12). Moreover, both species share a turreted shape, moderately strong and rather straight axial ribs as well as a detached columellar lip.? Vicinocerithium heberti differs clearly by having four (M1–M4 and m1) instead of only three major cords on spire (M1–M3) in? V. inopinatum, and by having more axial ribs, 10–12 ribs on spire and 13–14 ribs on last whorl, 7–8 on spire instead of 12 on last whorl in? V. inopinatum . Glibert (1973) attributed? V. inopinatum to Vicinocerithium, but the type species of the genus Vicinocerithium parallelum E. Wood, 1910 and many members of the genus differ by having a prominent spiny cord located in the adapical part of the whorls. However, several Eocene species of the genus, as V. turbinoides (Deshayes, 1864) from the Ypresian of the Paris Basin, are less spiny. Finally,? V. heberti has an atypical sculpture for Vicinocerithium, but its shape is more similar to that of this genus than to that of Cirsocerithium . For this reason, we assign? V. heberti to Vicinocerithium, whilst indicating uncertainty by the use of a question mark.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne and?Seine-et-Marne, France. This species is one the most common from the Maastrichtian of the Paris Basin. It is found from the base of the section of Vertus (Grand Mont) to the top of section of Bergères-lès-Vertus (Mont Aimé). Combes (1908) reported Cerithium carolinum from the Bois d’Esmans (Seine-et-Marne), but it could be? Vicinocerithium heberti sp. nov.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E79FF931318FA94FB369099	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7BFF9213D6FC2FFBB19262.text	F12B8A1C7E7BFF9213D6FC2FFBB19262.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vicinocerithium undetermined	<div><p>? Vicinocerithium sp.</p><p>Figs 11K–L, 12D</p><p>Vicinocerithium sp. 2 – Merle &amp; Pacaud 2018a: 169, fig. 156c. — Merle 2023: 25.</p><p>Material examined (3 specs)</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156c) and herein Figs 11K, 12D; MNHN.F.A6675 • 1 spec.; same data as for preceding; MMM coll.; MNHN.F.C02650 • 1 spec.; Vertus ( Grand Mont); sandy limestone; MMM coll.; Fig. 11L; MNHN.F.C02651 .</p><p>Measurements</p><p>MNHN.F.A6675, H: 15.7 mm, D: 7 mm. Paratype MNHN.F.C02651, H: 10 mm, D: 4.3 mm. MNHN.F.C02650, H (three last whorls only): 12 mm, D: 8.3 mm.</p><p>Description</p><p>Shell medium in size, elongate and turreted. Around nine moderately convex whorls. Apical angle 32° (based on holotype). Suture moderately shallow. Height of last whorl: 36% of shell length (based on holotype). Axial sculpture consisting of low ribs becoming slightly stronger on penultimate and last whorls. Number of ribs increasing from 12 to 25 from second to antepenultimate whorl and falling to 16 ribs on penultimate whorl. On two first teleoconch whorls, three major cords (M1–M3), on third whorl appearance of minor cord m1 and on other spire whorls M1, m1, M2 and M3 (Fig. 12D). Cords weak forming spiral rows of small rounded beads. Aperture not preserved.</p><p>Remarks</p><p>? Vicinocerithium sp. is reminiscent of Pseudoaluco sp. in its low and slightly curved ribs and its four subequal beaded major cords. However,? Vicinocerithium sp. lacks varical nodes observed in Pseudoaluco sp. In addition, the last whorls of? Vicinocerithium sp. bear stronger varices recalling more those of? V. heberti sp. nov. ? Vicinocerithium sp. differs from? V. heberti in having weaker and more numerous axial ribs (12 to 25 on spire and 16 on penultimate whorl instead of 10–12 on spire and 13–14 on last whorl in? V. heberti) and weaker and more rounded beads on cords.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7BFF9213D6FC2FFBB19262	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7AFF921367FEA2FAAB91AF.text	F12B8A1C7E7AFF921367FEA2FAAB91AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Granulolabium Cossmann 1889	<div><p>Genus Granulolabium Cossmann, 1889</p><p>Type species</p><p>Granulolabium plicatum Bruguière, 1792 by original designation. Early Oligocene (Rupelian), Paris Basin, France.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous of Paris Basin (this work) to Serravallian of Paratethys (Harzhauser et al. 2023).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7AFF921367FEA2FAAB91AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7AFF951303FD7EFDBA9701.text	F12B8A1C7E7AFF951303FD7EFDBA9701.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Granulolabium cretaceum Merle 2026	<div><p>Granulolabium cretaceum sp. nov.</p><p>urn:lsid:zoobank.org:act: 9EBABB04-A051-4CD6-B984-64808CD34F44</p><p>Figs 11M–O, 12E–F</p><p>Granulolabium sp. – Merle 2023: 25.</p><p>Etymology</p><p>‘Cretaceum’: name given to recall the Cretaceous range of the species.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Mont Aimé, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; Fig. 11O; MNHN.F.C02654.</p><p>Paratypes (2 specs) FRANCE – Marne • 1 spec.; Vertus ( Grand Mont); sandy limestone; MMM coll.; Figs 11M, 12F; MNHN.F.C02652 • 1 spec.; same data as for preceding; MMM coll.; Figs 11N, 12E; MNHN.F.C02653 .</p><p>Measurements</p><p>Holotype, H: (incomplete apex): 7 mm, D: 3.9 mm. Paratype MNHN.F.C02652, H (incomplete apex): 6.8 mm, D: 2.4 mm. Paratype MNHN.F.C02653, H (incomplete apex): 7 mm, D: 3 mm.</p><p>Description</p><p>Shell small in size, slender and turreted. Protoconch unknown. Around nine nearly straight-sided whorls. Apical angle 25° (based on paratype MNHN.F.C02652). Suture shallow, weakly incised. Height of last whorl: 31% of shell length (on paratype MNHN.F.C02652). Axial sculpture not varicose, with no defined ribs. Four major cords (M1–M4) on spire. M1 and M2 more prominent, ornamented with subquadratic beads. M3 linaear poorly beaded. M4 smooth, smaller than other adapical major cords. M4 and abapical suture close set. In holotype, M5 and M6 present (Fig. 12E–F). Small threads between most prominent cords M1, M3 and M4. Aperture subquadrangular. Columellar lip adherent. Siphonal canal short.</p><p>Comparisons</p><p>These narrow shells, regular, not varicose and bearing major cords decorated of subquadratic beads, are typical of the genus Granulolabium . According to Lozouet (1986), Granulobium montense (Briart &amp; Cornet, 1873) from the Danian of Mons is the oldest record of the genus. Briart &amp; Cornet (1873) described two other species: Cerithium ovalituberosum and Cerithium sexlinum from the same beds. However, Granulobium montense displays a large variability discussed by Glibert (1973: 35) and Kowalke (2002). As first reviser, Glibert (1973) considered Cerithium ovalituberosum and Cerithium sexlinum to be junior subjective synonyms of Cerithium montense . Kowalke (2002) admitted, as Glibert (1973), that there is only one species of Granulolabium in the Danian of Mons, but did not took into account the nomenclature act of Glibert (1973) and used erroneously the binome Granulolabium ovalituberosum . Although G. montense displays a large shell variability, Glibert (1973: 35) noted that “la sculpture spirale comporte sur chaque tour quatre cordons spiraux principaux munis de nodosités longitudinales de grosseur variable [= M1 to M4]. ... Un cinquième cordon [= M5], un peu plus faible et lisse, longe la suture antérieure”. [English translation: “The spiral structure comprises, on each whorl, four main spiral cords with longitudinal nodules of varying sizes [= P1 to P4]. ... A fifth cord [= P5], slightly thinner and smooth, extends along the anterior suture”]. Granulolabium cretaceum sp. nov. differs from G. montense by having only four major M1 to M4 on the spire. In addition, M4 is close to the abapical suture in G. cretaceum, whereas M5 is close to abapical suture in G. montense . The occurrence of Granulolabium in the Maastrichtian is the oldest occurrence of the genus and indicates that it was already present in the Late Cretaceous.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7AFF951303FD7EFDBA9701	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7DFF95131CFB61FE4096C3.text	F12B8A1C7E7DFF95131CFB61FE4096C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haustator Montfort 1810	<div><p>Genus Haustator Montfort, 1810</p><p>Type species</p><p>Haustator gallicus Montfort, 1810 (junior synonym of Turritella imbricataria Lamarck, 1804) by original designation. Middle Eocene (Lutetian), Paris Basin, France.</p><p>Stratigraphic range of the genus</p><p>Early Cretaceous (Valanginian) of Poland (Bandel 1993; Schröder 1995) to Early Miocene of South West India (Harzhauser 2014)</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7DFF95131CFB61FE4096C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7DFF971315F9C3FDBA95F3.text	F12B8A1C7E7DFF971315F9C3FDBA95F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haustator bergierensis Merle 2026	<div><p>Haustator bergierensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 04B7CC19-0E2B-4A52-984A-00BD4EFBBBE1</p><p>Figs 13A–D, 14A–E</p><p>Haustator sp. – Merle &amp; Pacaud 2018a: 169, fig. 156g. — Merle 2023: 25.</p><p>Etymology</p><p>From ‘bergier’ the ancient name of the type locality Bergères-lès-Vertus.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Mont Aimé, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; Figs 13A, 14A; MNHN.F.C02655.</p><p>Paratypes (5 specs) FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 156g); MNHN.F.A66756 • 1 spec.; same data as for holotype; MMM coll.; Figs 13B, 14B; MNHN.F.C02656 • 1 spec.; same data as for holotype; MMM coll.; Figs 13C, 14C; MNHN.F.C02657 • 1 spec.; same data as for holotype; MMM coll.; Figs 13D, 14D; MNHN.F.C02658 • 1 spec.; same data as for holotype; MMM coll.; Fig. 14E; MNHN.F.C02659 .</p><p>Other examined material (11 specs)</p><p>FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; MNNH.F.C02660 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02661 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02662 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02663 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F. C02664 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02665 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02666 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02667 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02668 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02669 • 1 spec.; same data as for holotype; MMM coll.; MNHN.F.C02670. Many fragments are not listed in the examined material .</p><p>Measurements</p><p>Holotype, H (six whorls, apex and base incomplete): 10 mm, D: 4 mm. Paratype MNHN.F.C02656, H (four whorls, apex and base incomplete): 12 mm, D: 6 mm. Paratype MNHN.F.C02657, H (five whorls, apex and base incomplete): 12 mm, D: 5.5 mm. Paratype MNHN.F. C02658, H (eight whorls, apex subcomplete): 11.3 mm, D: 4 mm. Paratype MNHN.F. C02659, H (around 13 whorls, base incomplete): 10.5 mm, D: 3.5 mm. Paratype MNHN.F.A667560, H (two whorls, apex and base incomplete): 12 mm, D: 7 mm.</p><p>Description</p><p>Shell medium in size, slender, comprising up to 20 teleoconch whorls when complete. Protoconch not preserved. Whorl profile shifting from frustate to campanulate during ontogeny (terminology adopted from Allmon 1996). Spiral angle between 15° (juvenile, based on paratype MNHN.F.C02659) and 21° (adulte, based on holotype). On first ten teleoconch whorls M3 close set to abapical suture and becoming prominent during ontogeny. On later whorls, M1 (most adapical cord) and M2 (central) more or less weak, M1 being weaker than M2. Minor cords m1 and m2 present; m1 occasional (see Fig. 13B–C). Sutural ramp indistinct, flat. Suture moderately incised. Sinuses not preserved. Aperture subquadrate. Base flat.</p><p>Comparisons</p><p>Haustator bergierensis sp. nov. is reminiscent of three turritellid species from the Aachener Formation (Santonian of Germany) in its slender shape, its frustate to campanulate profile and in its three major cords (M1–M3): 1°) ‘ Turritella ’ omaliusi Müller, 1851 (Müller 1851: 31, pl. 4 fig. 12a–b) a junior synonym of Nodosella alternans (F.A. Roemer, 1841), 2°) ‘ Turritella humbolti ’ Müller, 1851 (Müller, 1851: 32, pl. 4 fig. 17) and 3°) ‘ Turritella ’ acutissima Müller, 1849 (Müller, 1849; 31, pl. 4 fig. 19a–b). The major cord M3 appears late in the ontogeny in H. bergieriensis, whereas it appears on the early teleoconch whorl in ‘ T. ’ omaliusi. Moreover, only M1 is prominent in H. bergieriensis, whereas M1 and M3 are prominent in ‘ T. ’ omaliusi (M2 being smaller). ‘ Turritella ’ omaliusi is recorded in the Maastrichtian of Maastricht (Binkhorst 1861: pl. 1 fig. 6). ‘ Turritella ’ humboldti differs in having M1 to M3 appearing later in the ontogeny than in H. bergierensis and M4 is not close set to the abapical suture on the last whorls. The major cords M1 to M3 of ‘ T. ’ acutissima appear earlier in the ontogeny than in H. bergierensis . The suture is always shallow in H. bergierensis, whereas that of ‘ T. ’ acutissima is deep and delineates the teleoconch whorls. In addition, M3 is clearly more prominent in H. bergieriensis, whereas M1 to M3 have a more or less similar relief in ‘ T. ’ acutissima. The shape of the whorls of Haustator plana (Binkhorst, 1861) from the Maastrichtian of Maastricht is reminiscent of that of Haustator bergierensis, but the latter is clearly distinguishable by its coarser spiral cords and a significantly shallower suture. The cords of Haustator plana are extremely fine and cannot be identified (see the text and the figure of Binkhorst 1861: 30, pl. 1 fig. 5). This is not the case with specimens of Haustator bergierensis, in which the M1–M3 cords are easily identifiable. The Cenozoic Haustator nysti (Briart &amp; Cornet, 1873) from the Danian from Mons (Belgium) shares with H. bergierensis a prominent M3 and weaker M1 and M2. However, this species differs from H. bergierensis in having a more prominent M3 appearing on the early teleoconch whorls and in having teleoconch whorls bearing an imbricate shape.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7DFF971315F9C3FDBA95F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7FFF961382F932FDFC91C6.text	F12B8A1C7E7FFF961382F932FDFC91C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haustator undetermined	<div><p>Haustator sp.</p><p>Fig. 14F</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus (Mont Aimé); sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02671 .</p><p>Measurements</p><p>H (four whorls, apex and base incomplete): 7 mm, D: 2.5 mm.</p><p>Description</p><p>Shell slender. Whorl profile subquadrate (terminology adopted from Allmon 1996). No spiral sculpture. No sutural ramp. Suture very shallow. Sinuses not preserved.</p><p>Remarks</p><p>This fragment of turritellid differs from H. bergierensis sp. nov. by having a subquadrate profile and in lacking a spiral sculpture. Chavan (1948) cited from the Mont Aimé Turritella cf. supracretacea, a species originally described from the Danian of Meudon. Unfortunately, Chavan did not illustrate any specimens. The two syntypes illustrated by Pacaud (2018a: fig. 193a–b) as Haustator supracretacea bear a weak sculpture as in Haustator sp.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7FFF961382F932FDFC91C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7EFF961304FCA1FB5897AE.text	F12B8A1C7E7EFF961304FCA1FB5897AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanile P. Fischer 1884	<div><p>Genus Campanile P. Fischer, 1884</p><p>Type species</p><p>Cerithium leve Quoy &amp; Gaimard, 1834 (non Perry, 1810 = Campanile symbolicum Iredale, 1917) by subsequent designation (Crosse 1888). Present day, Western Australia.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian) of Mexico (Kiel et al. 2000) to present day (Australia).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7EFF961304FCA1FB5897AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E7EFF98133EFB3FFAED917D.text	F12B8A1C7E7EFF98133EFB3FFAED917D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Campanile (Campanile) undetermined	<div><p>Campanile (Campanile) sp.</p><p>Fig. 14G</p><p>Campanile sp. – Merle &amp; Pacaud 2018a: 169, fig. 156h.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); unknown bed; MNHN coll., ex n°GG. gg2007/16527; figured in Merle &amp; Pacaud (2018a: fig. 156h); MNHN.F.C02672 .</p><p>Measurements</p><p>H (incomplete): 82 mm; D: 47 mm.</p><p>Description</p><p>Shell large and turreted. Internal mould displaying four spire whorls. Inside of whorls convex with two grooves corresponding to two columellar folds.</p><p>Comments</p><p>Hébert (1848: 394) reported a Campanile from Vertus (probably from Faloise), which he identified, albeit with some doubt, as Campanile giganteum (Lamarck, 1804) from the Lutetian of Paris Basin. Later, d’Orbigny (1850) reported in the type material of the Danian Campanile unisulcatum (d’Orbigny, 1850), a specimen coming from Vertus. However, this specimen has not been retrieved in his collection. In the collections of the MNHN, we found only a Maastrichtian specimen of Campanile from the locality of Bergères-lès-Vertus (Mont Aimé). However, it is an internal mould and it does not allow an identification at species level (see Fig 14G). Therefore it is not possible to validate this identification. Binkhorst (1861: pl. 6 fig. 1c) illustrated an internal mold of Campanile (s. str.) maximum (Binkhorst, 1861) from Maastricht, but it differs by having three grooves instead of two in the Campanile from Bergères-lès-Vertus (Mont Aimé).</p><p>Stratigraphic range</p><p>Combes (1908: 14–15) cited Cerithium uniplicatum d’Orbigny, 1850 which is now attributed to the genus Campanile (see Merle &amp; Pacaud 2018b). This identification of Combes (1908: 15) confirms unequivocally the presence of the genus Campanile at the Bois d’Esmans: “… la présence de moules internes de gros cérites, analogues à ceux du Cerithium giganteum lutétien ...”. English translation [“… the presence of internal moulds of large cerites, similar to those of the Lutetian Cerithium giganteum … ”].</p></div>	https://treatment.plazi.org/id/F12B8A1C7E7EFF98133EFB3FFAED917D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E70FF981312FCE8FE5F977F.text	F12B8A1C7E70FF981312FCE8FE5F977F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sveltella Cossmann 1889	<div><p>Genus Sveltella Cossmann, 1889</p><p>Type species</p><p>Cancellaria quantula Deshayes, 1864 by original designation (junior synonym of Cancellaria biplicata Lamarck, 1803). Middle Eocene (Lutetian), Paris basin, France.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian) of Paris Basin (this work) to Late Oligocene (Chattian) of Aquitaine Basin (Lozouet 1999).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E70FF981312FCE8FE5F977F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E70FF9B133EFB4CFDBA97E6.text	F12B8A1C7E70FF9B133EFB4CFDBA97E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sveltella binkhorsti Merle 2026	<div><p>Sveltella binkhorsti sp. nov.</p><p>urn:lsid:zoobank.org:act: F58C53D6-A0DF-47B1-9343-9A2A808E84FC</p><p>Fig. 14H–I</p><p>Caveola sp. – Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to the palaeontologist Jan Theodor Binkhorst van den Binkhorst who described the molluscs fauna from the Maastrichtian of Limburg.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont), sandy limestone; MMM coll.; Fig. 14H; MNHN.F.C02675.</p><p>Paratype FRANCE – Marne • 1 spec.; same data as for holotype; MMM coll.; Fig. 14I; MNHN.F.C02676 .</p><p>Measurements</p><p>Holotype MNHN.F.C02675, H: 9.5 mm, D: 4.7 mm. Paratype MNHN.F.C02676, H: 9 mm, D: 4 mm.</p><p>Description</p><p>Shell slender, subfusiform small in size. Protoconch missing. Spiral angle 32°. Five spire whorls. Two early whorls poorly preserved. Antepenultimate and penultimate whorls high, subcarinate, slightly constricted adapically. Narrow subsutural ramp. Suture distinct, moderately deep. Last whorl moderately subcarinate, equals 53% of total shell height. Siphonal canal short. On antepenultimate and penultimate whorls four–five strong, varicose ribs in dorsal view (= one half whorl) tending to fade near sutural ramp. Five thin spiral cords covering delicately the ribs and more prominent between them. On last whorl, five lower ribs. Nine–ten thin spiral cords of equal size undulating across the ribs. Siphnal canal short.</p><p>Comparisons</p><p>The shell shape of Sveltella binkhorsti sp. nov. is reminiscent of that of Caveola producta Stephenson, 1941 from the Maastrichtian of Texas (USA), type species of Caveola Stephenson, 1941 (by original designation) and attributed to the family Mataxidae Bandel &amp; Dockery, 2012 and to the superfamily Tonnoidea Suter, 1913 . Nevertheless, the sculpture of S. binkhorsti differs from that of Caveola in having higher and less numerous ribs (5–6 instead of 9–11 on spire and 5 instead of 15–16) crossed by weaker spiral cords. Sveltella has a similar shape and its sculpture resembles more that of S. binkhorsti . Sveltella loevstroemi Schnetler &amp; Petit, 2010 from the Danian of Nuusuaq (Greenland) and S. binkhorsti share 5–6 varicose ribs on the spire and undulating cords on the ribs. Furthermore, in both species, the ribs become less strong on the last whorl. Sveltella binkhorsti is distinguished from S. loevstroemi by having a slightly higher spire whorls, fewer ribs (6–7 on spire and 8 on last whorls) and weaker cords. Sveltella binkhorsti differs from Sveltella tobiasseni Schnetler &amp; Petit, 2010 from the Danian of Nuusuaq by having less coarse spiral cords and more rounded whorls. The shape of S. quantula from the Lutetian of the Paris Basin is also quite similar to that of S. binkhorsti . Howewer, S. binkhorsti displays higher and less convex whorls and its spiral sculpture consists of coarser and fewer cords.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E70FF9B133EFB4CFDBA97E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E73FF9B131FFAA2FB4695AD.text	F12B8A1C7E73FF9B131FFAA2FB4695AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fusinus Rafinesque 1815	<div><p>Genus Fusinus Rafinesque, 1815</p><p>Type species</p><p>Murex colus Linnaeus, 1758 by typification of replaced name. Present day, Indian Ocean and Indo-West-Pacific.</p><p>Stratigraphic range of the genus</p><p>?Late Cretaceous (Campanian) of The Netherlands (this work) to present day, warm seas.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E73FF9B131FFAA2FB4695AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E73FF9A1326F963FDBA9445.text	F12B8A1C7E73FF9A1326F963FDBA9445.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fusinus bouchardae Merle 2026	<div><p>Fusinus bouchardae sp. nov.</p><p>urn:lsid:zoobank.org:act: 157C1A91-01D9-4B04-9FF4-25A5B8AA0940</p><p>Fig. 14J–K</p><p>Dolicholatirus sp. – Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to Blanche Bouchard for her help on the computerisation of the mollusc collection of Vertus (Grand Mont) and Bergères-lès-Vertus (Mont Aimé).</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02677.</p><p>Measurements</p><p>Holotype, H (three last whorls observed): 12.7 mm, D: 5 mm.</p><p>Description</p><p>Shell medium in size, fusiform, rather elongated, measuring approximately 15 mm. Spire whorls convex, slightly constricted adapically by a narrow subsutural band. Spiral angle around 41°. Suture distinct, but not canaliculate. On antepenultimate and penultimate whorls four varicose and rounded ribs in dorsal view (= one half whorl) tending to be more prominent abapically. Seven to eight thin spiral cords of equal size, covering delicately the ribs and more prominent between the ribs. Last whorl elongate. Siphonal canal moderately long occupying 53% of length of last whorl. Five varicose ribs in dorsal view (= one half whorl) tending to fade near siphonal canal. Eleven thin spiral cords of equal size on convex part of whorl. On siphonal canal, cords more distinct on adapical part.</p><p>Comparisons</p><p>Fusinus bouchardae sp. nov. is reminiscent of “ Fusus ” gracilis (J. Böhm, 1885), not Fusus gracilis Da Costa, 1778 or “ Fusus ” scala Holzapfel, 1888 from the Campanian Grünsand of Vaals (The Netherlands) in its fusiform shape, its varicose ribs crossed by thin spiral cords of equal size and in its whorls slightly constricted adapically (see Holzapfel 1888: 103, pl. 11 figs 1–2). However, “ Fusus ” gracilis has more (6–7 for one half whorl) and close-set axial ribs and “ Fusus ” scala displays flatter ribs than those of Fusinus bouchardae . There is no similar shell in the works of Binkhorst (1861) and Holzapfel (1888) on the Late Cretaceous mollusc fauna of Limburg. The shape and form of Fusinus schnetleri Hansen, 2019 from the lower Danian of Denmark closely resemble those of F. bouchardae, but the latter differs in having more elongated whorls and a less pronounced subsutural band. From the point of view of genus assignment, Merle (2023) previously attributed this species to Dolicholatirus Bellardi, 1884 (family Dolicholatiridae Kantor, Fedosov, Kosyan, Puillandre, Sorokin, Kano, R. Clark &amp; Bouchet, 2021). However Fusinus bouchardae . differs from members of Dolicholatirus (type species: Fusus bronnii Michelotti, 1847 by subsequent designation Cossmann (1901); Late Miocene, Italy) in having a shorter shape and finer spiral cords. The shape and the sculpture of “ Fusus ” gracilis, “ Fusus ” scala, Fusinus schnetleri and Fusinus bouchardae bear a closer resemblance to those of the species of Fusinus . For this reason, it seems more appropriate to provisionally assign F. bouchardae to Fusinus . This assignment is provisional and requires further verification using new material to determine whether the columella of this species is smooth, as is the case with species of Fusinus .</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E73FF9A1326F963FDBA9445	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E75FF9D1379FE87FB3D914C.text	F12B8A1C7E75FF9D1379FE87FB3D914C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudopisania Cossmann 1897	<div><p>Genus Pseudopisania Cossmann, 1897</p><p>Type species</p><p>Mitrella (Pseudopisania) plateaui (Cossmann, 1889) by original designation. Early Eocene (Ypresian), Paris Basin, France.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian) of Paris Basin (this work) to Early Eocene (Europe).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E75FF9D1379FE87FB3D914C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E75FF9C1319FD43FDBA9276.text	F12B8A1C7E75FF9C1319FD43FDBA9276.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudopisania cretacea Merle 2026	<div><p>? Pseudopisania cretacea sp. nov.</p><p>urn:lsid:zoobank.org:act: 53A885F1-9D43-4918-919A-3C97EC9FF8DB</p><p>Fig. 14L–M</p><p>Etymology</p><p>From ‘ cretaceous ’, name given to recall the Cretaceous range of the species.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02684.</p><p>Measurements</p><p>H: 10 mm, D: 4 mm.</p><p>Description</p><p>Shell medium in size, fusiform, elongate. Protoconch missing. Spiral angle 35°. Six spire whorls, slightly rounded. Last whorl of 50% of total shell height. Suture linear, slightly impressed. No axial sculpture. On penultimate whorl, one weak thread below suture. On last whorl, two weak threads below suture and several weak basal cords on siphonal canal. Outer lip thickened by terminal varix. Siphonal canal moderately long (in dorsal face) of 36% of height of last whorl and slightly recurved.</p><p>Comparisons</p><p>The fusiform shape of Pseudopisania cretacea sp. nov., its slightly rounded whorls with a moderately long siphonal canal, spiral threads below the suture and on the siphonal canal recall some mitrids such as Mitra subglabra (Ravn, 1933) from the Danian of Denmark (Ravn 1933: 65 and figured specimen in Lauridsen &amp; Schnetler 2014: 97, fig. 153), or columbellids such as Pseudopisania plateaui (Cossmann, 1889) from the Thanetian of the Paris Basin (Cossmann 1889: 142 and figured specimen in Cossmann &amp; Pissarro 1911: pl. 37 fig. 179-18). The outer lip thickened by a terminal varix of P. cretacea lacks in Mitridae, but is common in Columbellidae . Among the Columbellidae, P. cretacea resembles P. plateaui which only differs in having more pronounced threads on the last whorl. The assignment of P. cretacea to Pseudopisania is, however, provisional and requires examination of the aperture to validate it.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E75FF9C1319FD43FDBA9276	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E74FF9C131FFE11FEFE903A.text	F12B8A1C7E74FF9C131FFE11FEFE903A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Popenoeum Squires 1989	<div><p>Genus Popenoeum Squires, 1989</p><p>Type species</p><p>Popenoeum maritimum Squires, Zinsmeister &amp; Paredes-Mejia, 1989 by original designation. Paleocene (Danian), southern California, USA.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian) of North Africa, Madagascar and northern Europe to Early Eocene (see Pacaud 1998).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E74FF9C131FFE11FEFE903A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E74FF9F136BFC89FA809047.text	F12B8A1C7E74FF9F136BFC89FA809047.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Popenoeum ambiguum (Binkhorst 1861)	<div><p>Popenoeum ambiguum (Binkhorst, 1861)</p><p>Fig. 14N–O</p><p>Pyrula ambigua Binkhorst, 1861: 6, pl. 1 fig. 9.</p><p>Pseudoliva ambigua – Kaunhowen 1898: 78, pl. 7 figs 3–4. — Pervinquière 1912: 64, pl. 5 figs 1–2a–c.</p><p>Sulcobuccinum ambiguum – Vermeij 1998: 81.</p><p>Popenoeum ambiguum – Pacaud 1998: 3, figs 1–9b, 12. — Merle &amp; Pacaud 2018a: 169, fig. 156h; 2018b: 322, fig. 230g –h. — Merle 2023: 25.</p><p>For a detailed synonymic list, see Pacaud (1998: 3).</p><p>Type material (not examined)</p><p>Two syntypes (see Binkhorst 1861: 6).</p><p>Type locality</p><p>Maastricht or Fauquemont, cf. Binkhorst (1861: iv), The Netherlands; Meerssen Member, Maastrichtian.</p><p>Material examined (4 specs)</p><p>FRANCE – Marne • 1 spec., external part of an outer lip showing the pseudolivid spiral groove; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured specimen of Merle &amp; Pacaud (2018a: fig. 156i); MNHN.F.A66757 • 1 spec., spire only; Vertus ( Grand Mont); sandy limestone; MMM coll.; MNHN.F.C02686 • 1 spec., subcomplete; same data as for preceding; MMM coll.; Fig. 14N–O; MNHN.F.C02685 • 1 spec., juv.; same data as for preceding; MMM coll.; MNHN.F.C02687 .</p><p>Measurements</p><p>MNHN.F.C02685, H: 11 mm, D: 6 mm.</p><p>Description</p><p>Shell medium in size and globose. Protoconch missing. Spiral angle 54°. Four spire whorls, rather high, moderately convex and separated by a canaliculate suture. Last whorl large, convex, of 73% of total shell height. On antepenultimate and penultimate whorls, 8–9 distinct and flexuous axial ribs (in dorsal view = one half whorl) reaching adapical suture, without bearing spines on shoulder. On last whorl, eight ribs (in dorsal view = one half whorl) reaching dorsal groove and disappearing abapically. Spiral sculpture lacking on spire and on last whorl consisting of around ten small threads extended from dorsal groove to base. Dorsal pseudolivid groove shallow. Outer lip not thickened. an canal short.</p><p>Comparisons</p><p>Merle &amp; Pacaud (2018a) identified this species from a fragment of an outer lip bearing a pseudolivid shallow dorsal groove found at Bergères-lès-Vertus (Mont Aimé) in 2014. The 2017 fieldtrip led to the discovery of three new specimens, one of which was fairly complete: MNHN.F.C02685 (Fig. 14N–O). It is reminiscent of the specimen from Maastricht illustrated by Kaunhowen (1898: pl. 7 figs 3–4) in having rather high spire whorls, flexuous axial ribs reaching the adapical suture, without bearing spines on the shoulder, and in having a poorly developed spiral sculpture. According to Pacaud (1998), P. ambiguum has a wide geographic range and is also present in the Danian.</p><p>Stratigraphic range</p><p>Maastrichtian: France (Marne), The Netherlands, North Africa and Madagascar to Danian–Ypresian: Europe, Western and Eastern Tethys, West Africa and Caribbean, for details see Pacaud (1998).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E74FF9F136BFC89FA809047	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E77FF9F1318FBDEFE06960B.text	F12B8A1C7E77FF9F1318FBDEFE06960B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Volutomorpha Gabb 1877	<div><p>Genus Volutomorpha Gabb, 1877</p><p>Type species</p><p>Volutilithes conradi Gabb, 1860 by original designation. Late Cretaceous, New Jersey, USA.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Turonian) of Egypt (El Qot 2006) to Maastrichtian (North America, West Africa, Europe, Tethys…).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E77FF9F1318FBDEFE06960B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E77FF9E13E5FA9AFEE992FA.text	F12B8A1C7E77FF9E13E5FA9AFEE992FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Volutomorpha undetermined	<div><p>Volutomorpha sp.</p><p>Fig. 14P</p><p>Volutocorbis sp. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02682 .</p><p>Measurements</p><p>H: 7 mm, D: 3.5 mm.</p><p>Description</p><p>Shell small, biconic. Protoconch not preserved. Four teleoconch whorls including last whorl. Last whorl of 79% of total length. Apical angle 66°. Spire moderately high. Spire whorls slightly convex; last whorl elongate, slightly convex in its adapical part. On two first whorls no sculpture, on penultimate and last whorls six–seven low and orthocline axial ribs (in dorsal view = one half whorl). On last whorl, several small cords from suture to first adapical part of last whorl. Siphonal canal slightly elongate.</p><p>Remarks</p><p>This small shell corresponds to a juvenile individual and is reminiscent of Volutocorbis Dall, 1890 or Volutomorpha in its shape and sculpture. The early sculpture of Paleocene/Early Eocene Volutocorbis (see Merle et al. 2014) is, however, stronger than in this specimen from the Paris Basin. Young specimens of Volutomorpha, a genus present in the Late Cretaceous, are more similar in having a poorly developed early sculpture.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E77FF9E13E5FA9AFEE992FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E76FF811348FADAFEF39144.text	F12B8A1C7E76FF811348FADAFEF39144.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undetermined undetermined	<div><p>Caenogastropoda indet. 2 (?calyptraeid shape)</p><p>Fig. 15B</p><p>Calyptraea sp. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02674 .</p><p>Measurements</p><p>D: 14.8 mm.</p><p>Description</p><p>Fragment of external part of last whorl; spire broken. Hat shell shape. Last whorl convex, moderately high. Suture line between penultimate whorl and last whorl linear. Outline of base subcircular. Shell surface smooth, but with a few small tips more or less spirally aligned.</p><p>Remarks</p><p>This fragment of the last whorl is reminiscent of the calyptraeids in its hat shell shape, its subcircular outline and its moderately high last whorl. Its shell surface with small lines more or less spirally aligned is found in the genus Sigapatella R.P. Lesson, 1831, particularly in Sigapatella aperta (Solander in Brander, 1766) from the middle Eocene (Bartonian) of the Paris Basin, see Cossman &amp; Pissarro (1911: pl. 12 fig. 73.1). The oldest member of the genus is Sigapatella montensis (Cossmann, 1919), see Pacaud (2018a: 29). Cossmann (1919: 70) stated “…il est orné de rugosités irrégulières, plus où moins alignées sur des lignes d’accroissement trè obliques” [English translation: “…it is decorated with irregular roughnesses, more or less aligned with very oblique growth lines”], as it is in? Sigapatella sp. According to Marshall (2003), “ Bandel &amp; Riedel (1994) interpreted Sigapatella as a synonym of Calyptraea Lamarck, 1799, but S. novaezelandiae and other species discussed herein differ from C. chinensis in that the septum edge is broadly and evenly concave instead of sigmoidal”. As the septum is not preserved in the collected fragment, it is not possible to determine with certainty whether it corresponds to Sigapatella .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E76FF811348FADAFEF39144	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E76FF9E134EFD8AFCD897CB.text	F12B8A1C7E76FF9E134EFD8AFCD897CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undetermined undetermined L. R. Cox 1960	<div><p>Caenogastropoda indet. 1 (?ampullinid shape)</p><p>Fig. 15A</p><p>Ampullinidae indet. – Merle &amp; Pacaud 2018a: 166. — Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02673 .</p><p>Measurements</p><p>H (incomplete): 12 mm, D (incomplete): 12 mm.</p><p>Description</p><p>Fragment of shell bearing three teleoconch whorls and internal part of last whorl. Whorls subcarinate to slightly convex. Surface apparently smooth. Basal internal part making an inverse cone with slightly concave sides.</p><p>Remarks</p><p>Although this fragment is significantly altered, it resembles a fragment of the spire of an Ampullinidae Cossmann, 1919 or maybe a Naticidae Guilding, 1834 .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E76FF9E134EFD8AFCD897CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E69FF811350FD4BFAF29677.text	F12B8A1C7E69FF811350FD4BFAF29677.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undetermined undetermined	<div><p>Caenogastropoda indet. 3 (?volutid shape)</p><p>Fig. 15C</p><p>Aporrhaidae indet. – Merle &amp; Pacaud 2018a: 169, 171, fig. 156j. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; figured specimen of Merle &amp; Pacaud (2018a, fig. 156j); MNHN.F.A66758 .</p><p>Measurements</p><p>H (spire missing): 7 mm, D (outer lip missing): 4 mm.</p><p>Description</p><p>Fragment of penultimate and last whorls in ventral side. Penultimate whorl convex and smooth. Last whorl conical, elongate with carina located in adapical quarter of whorl. Surface smooth. Suture linear. Inner lip slightly erect, making moderatery deep groove from adapical part of whorl to base.</p><p>Remarks</p><p>This fragment of shell is slightly reminiscent of a volutid in the conical and elongate shape of its last whorl, its carina and its inner lip. However, it is impossible to specify which genus it might belong to.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E69FF811350FD4BFAF29677	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E69FF80109BFAB6FEE2929F.text	F12B8A1C7E69FF80109BFAB6FEE2929F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenogastropoda L. R. Cox 1960	<div><p>Caenogastropoda indet. 4 (?volutid/cancellariid shape)</p><p>Fig. 15D</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02683 .</p><p>Measurements</p><p>H (apex and base incomplete), 4.7 mm, D: 4 mm.</p><p>Description</p><p>Two whorls preserved. Penultimate whorl moderately convex. On penultimate whorl, six low and orthocline ribs (in dorsal view = one half whorl) crossed by six subequal cords, tending to be nodulose when crossing the ribs. On last whorl, eight ribs (in dorsal view = one half whorl) crosssed by eight cords nodulose at intersection with ribs. In adapical part of whorl, subsutural cord clearly distinct from shoulder cord. Shoulder cord slightly spiny delineating small carina.</p><p>Remarks</p><p>This fragment of a small shell displays a stronger sculpture than Volutomorpha sp. It resembles Volutomorpha corrugata (Binkhorst, 1861) in having a shoulder cord delineaeting a small carina (see Binkhorst 1861: pl. 5 fig. 1a–b), but it has more numerous ribs. Its cancellate sculpture also recalls that of cancellariids.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E69FF80109BFAB6FEE2929F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E68FF8310ACFE2EFDA9916B.text	F12B8A1C7E68FF8310ACFE2EFDA9916B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undetermined undetermined	<div><p>Caenogastropoda indet. 5 (?turrid/aporrhaid shape)</p><p>Fig. 15E</p><p>Turridae indet . – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02678 .</p><p>Measurements</p><p>H (base missing): 14 mm, D: 8 mm.</p><p>Description</p><p>Shell medium in size, fusiform, rather elongated. Seven rather angular whorls. Spiral angle of 39°. Suture deep and linear. Four to five (in dorsal view = one half whorl) varicose protruding ribs, marking the periphery of whorls. On spire, subsutural ramp extending to middle of whorl with one or two small threads.</p><p>Remarks</p><p>The sculpture of varicose protruding ribs, marking the periphery of the whorls, is found in some Turridae or Aporrhaidae J.E. Gray, 1850 .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E68FF8310ACFE2EFDA9916B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6BFF8310ACFDB9FB67964D.text	F12B8A1C7E6BFF8310ACFDB9FB67964D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenogastropoda L. R. Cox 1960	<div><p>Caenogastropoda indet. 6 (?turrid/aporrhaid shape)</p><p>Fig. 15F</p><p>Material examined</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02679 .</p><p>Measurements</p><p>H (incomplete apex and siphonal canal broken): 10 mm, D: 4.9 mm.</p><p>Description</p><p>Shell medium in size, fusiform, moderately elongated. Five subcarinate whorls preserved. On spire, subsutural ramp extending to middle of whorl. Spiral angle of 33°. Suture shallow and linear. Five–six ribs on spire and eight on last whorl (in dorsal view = one half whorl). Ribs low, slightly nodulose on periphery; abapically to periphery, small pleurotomoid sinus and prosocyrt outline until base; adapically to periphery, ribs connected to a subsutural spiral row of small nodules, well visible on last whorl. Convex part of whorl and base of whorl lacking spiral sculpture.</p><p>Remarks</p><p>This shell called Caenogastropoda indet. 6 differs from that of Caenogastropoda indet. 5 in having a shorter shape, lower ribs, a subsutural row of small nodules and in lacking a spiral sculpture.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6BFF8310ACFDB9FB67964D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6BFF8210ACFA5CFDBB9138.text	F12B8A1C7E6BFF8210ACFA5CFDBB9138.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caenogastropoda L. R. Cox 1960	<div><p>Caenogastropoda indet. 7 (?turrid/aporrhaid shape)</p><p>Fig. 15G</p><p>Material examined (2 specs)</p><p>FRANCE – Marne • 1 spec.; Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02680 • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with carditid imprints; MMM coll.; MNHN.F.C02681 .</p><p>Measurements</p><p>MNHN.F.C02680, H (apex and base incomplete): 9 mm, D: 6 mm. MNHN.F.C02681, H (apex and base incomplete): 11.1 mm, D: 5.9 mm.</p><p>Description</p><p>Shell small, fusiform, moderately elongated. Four subcarinate whorls preserved including part of last whorl. On spire, subsutural ramp extending to middle part of whorl. Spiral angle of 37°. Suture shallow and linear. Five-six ribs on spire and six on last whorl (in dorsal view = one half whorl). Ribs low, slightly nodulose on periphery; abapically to periphery, ribs with prosocyrt outline until base; adapically to periphery, ribs connected to suture by oblique striae, well visible on last whorl. Spiral sculpture with delicate threads of subsutural ramp and convex part of whorl.</p><p>Remarks</p><p>These two other specimens resemble Caenogastropoda indet. 6, but differ in having a spiral sculpture of delicate threads on their whorls and in lacking a subsutural row of small nodules. Their characters likely correspond to another species.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6BFF8210ACFA5CFDBB9138	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6AFF82132DFCC3FC01970D.text	F12B8A1C7E6AFF82132DFCC3FC01970D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nucula Lamarck 1799	<div><p>Genus Nucula Lamarck, 1799</p><p>Type species</p><p>Arca nucleus Linnaeus, 1758 type by monotypy. Present day, North east Atlantic, Mediterranean and West Africa.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous to present day, cosmopolite (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6AFF82132DFCC3FC01970D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6AFF8513D3FB9CFDBA9607.text	F12B8A1C7E6AFF8513D3FB9CFDBA9607.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nucula bauti Merle 2026	<div><p>Nucula bauti sp. nov.</p><p>urn:lsid:zoobank.org:act: CBFA3580-162E-47E4-AEF5-C015E2A55BA0</p><p>Fig. 16A–D</p><p>Nucula sp. – Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to Jean-Paul Baut who actively participated in the fieltrip in July 2017.</p><p>Type locality</p><p>Vertus (Grand Mont), Marne, France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • RV, IF; Vertus ( Grand Mont), sandy limestone; MMM coll.; Fig. 16A; MNHN.F.C02688.</p><p>Paratypes (4 specs) FRANCE – Marne • 1 spec., LV, IF; same data as for holotype; MMM coll.; Fig. 16B; MNHN.F.C02689 • 1 spec., EF in anatomical connection and RV visible; same data as for holotype; MMM coll.; Fig. 16C; MNHN.F.C02701 • 1 spec., EF in anatomical connection and RV visible; same data as for holotype; MMM coll.; Fig. 16D; MNHN.F.C02702 • 1 spec., internal mould; same data as for holotype; MMM coll.; MNHN.F.C02703 .</p><p>Measurements</p><p>Holotype, DVL: 5.8 mm; APL: 8 mm. MNHN.F.C02701, DVL, 6 mm; APL: 7 mm. Paratype MNHN.F.C02702, DVL: 7.8 mm; APL: 10.2 mm.</p><p>Description</p><p>Subequilateral shell, anterior area rather elongate. Dorsal margin moderately curved, slightly convex; anterior margin rounded. Ventral margin wide and convex; postero-ventral margin angular and dorsal margin almost straight, slightly concave near beak. Fine crenulations on ventral edge. Posterior muscle scar ovoid; anterior scar not visible. Pallial line apparently without sinus. Posterior hinge plate short bearing nine strong teeth with rectangular cross-section projecting vertically from hinge. Anterior hinge plate slightly convex with 15–18 teeth arranged similarly to those of anterior part of hinge. Chondrophore elongated situated in line with anterior hinge teeth.</p><p>Comparisons</p><p>In Nucula bauti sp. nov., an elongated chondrophore situated in line with the anterior hinge teeth is reminiscent of that of the Liassic species Nucula subovalis Goldfuss, 1837 (Goldfuss 1837: pl. 25 fig. 19). Nucula subovalis seems differ by having more teeth (24–25) on the anterior hinge plate and on the posterior hinge plate (11). Nucula bauti displays a less rounded shape and more teeth than the Danish Maastrichtian species Nucula (Nucula) aff. truncata Nilsson, 1827 presented by Heinberg (1989: fig. 1). Nucula truncata is also known in the Maastrichtian of Poland (Abdel Gawad 1986: pl. 25 figs 1–2), but the figured specimens are two poorly preserved internal moulds and don’t allow a reliable comparison. Nucula montensis Cossmann, 1908, from the Danian of Mons (Belgium), resembles Nucula bauti in its elongate chondrophore and in the number of teeth (8–9) on the posterior hinge. However, N. bauti differs by having fewer teeth (more than 15–18 instead of 25) on the anterior hinge which are less close set near the umbo, by a ventral margin more rounded and by a less angular postero-ventral margin (see Glibert &amp; van de Poel 1973: pl. 1 fig. 1). Despite the limited number of specimens available, the characters of N. bauti differ sufficiently from stratigraphically close species for it to be considered new.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6AFF8513D3FB9CFDBA9607	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6DFF85133FF9DDFBC8959F.text	F12B8A1C7E6DFF85133FF9DDFBC8959F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arca Linnaeus 1758	<div><p>Genus Arca Linnaeus, 1758</p><p>Type species</p><p>Arca noae Linnaeus, 1758 by subsequent designation Opinion 189 (ICZN 1945). Present day, North east Atlantic (North Sea to Portugal), Mediterranean and West Africa until Angola.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6DFF85133FF9DDFBC8959F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6DFF851328F92EFBD294F3.text	F12B8A1C7E6DFF851328F92EFBD294F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arca Linnaeus 1758	<div><p>Subgenus Arca Linnaeus, 1758</p><p>Note</p><p>MolluscaBase recognises several subgenera within the genus Arca . See MolluscaBase - Arca Linnaeus, 1758</p><p>Stratigraphic range of the subgenus</p><p>Late Cretaceous to present day, cosmopolitan in warm seas (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6DFF851328F92EFBD294F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6CFF861302FEC5FDBA90E6.text	F12B8A1C7E6CFF861302FEC5FDBA90E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arca submontensis Merle 2026	<div><p>Arca (s. str.) submontensis sp. nov.</p><p>urn:lsid:zoobank.org:act: AEEFF7B0-A754-43A3-A785-004236B69C19</p><p>Fig. 16E–H</p><p>Arca (s.s.) sp. – Merle &amp; Pacaud 2018a: 172, fig. 157a. — Merle 2023: 25.</p><p>? Arca montensis – Chavan 1948: 566 (non Arca montensis Cossmann, 1908).</p><p>Etymology</p><p>Name formed by the Latin prefix ‘ sub ’ (‘under’) and ‘ montensis ’ to show the affinities with Arca montensis Cossmann, 1908 from the Danian of Mons.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Maastrichtian.</p><p>Type material</p><p>Holotype FRANCE – Marne • RV, EF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; Fig. 16E–F; MNHN.F.C02690.</p><p>Paratypes (2 specs) FRANCE – Marne • 1 spec., RV, IF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 157a) and herein Fig. 16G; MNHN.F.A66760 • 1 spec., LV, EF; Vertus ( Grand Mont); sandy limestone; Fig. 16H; MNHN.F.C02691 .</p><p>Measurements</p><p>Holotype, DVL: 5.5 mm; APL: 13.5 mm. Paratype MNHN.F.C02691, DVL: 7 mm, APL: 16.5 mm.</p><p>Description</p><p>Shell medium in size, convex, angular dorsally and inaequilateral. Anterior margin curved and smoothly passing into ventral margin, but meets straight hinge line at a nearly 90º angle. Ventral margin curved near anterior margin, slightly concave in central part and slightly convex near posterior margin. Posterior margin angular, almost perpendicular to hinge and delineated by transverse ridge starting from umbo flexure. Beaks high, inclined anteriorly. Shell surface covered with around 50 fine riblets on antero-ventral margin, 5–6 finer riblets crossed by small growth lamellae on posterior ridge and 14–15 riblets on posterior margin. Riblets giving cancellate sculpture in combination with commarginal lines. Ligamental area wide, assymmetrical and low triangular in shape. Teeth small in middle part, but clearly elongate at posterior margin.</p><p>Comparisons</p><p>From the bed with shell concentrations of Mont Aimé, Chavan (1948) cited, but without figure, Arca montensis Cossmann, 1908, a species known from the Montian of Mons. Merle &amp; Pacaud (2018a) presented from the same bed one valve in internal face, which was difficult to identify and called it Arca (s. str.) sp. New field research conducted in 2017 led to the discovery of other, better-preserved specimens, allowing for more accurate descriptions and precise comparisons with other species of Arca . Arca (s. str.) submontensis sp. nov. clearly differs from Arca (s. str.) ernae Heinberg, 1978, from Danish Maastrichtian in having a larger size, a lower ligamentary area, a less prominent posterior ridge and weaker ribs.</p><p>The external morphology of Barbatia hennigi Heinberg, 1978 sensu Abdel Gawad (1986: pl. 26 fig. 4) from the Maastrichtian of Poland is somewhat reminiscent of that of A. (s. str.) submontensis sp. nov., but the hinge of the type specimen of B. heninigi from the Danish Maastrichtian is not straight like that of Arca and is very similar to that of a Barbatia E. Gray, 1842 . The shape, the size and the riblets of Arca (s. str.) montensis Cossmann, 1908 from the Danian recall more Arca (s. str.) submontensis and these two species seem closely related. The antero-ventral margin of A. (s. str.) montensis bears around 50–60 riblets in specimens from the Danian of Mons (see Glibert &amp; van de Poel 1973: pl. 1 fig. 5) and from the Danian of the Paris Basin (see Pacaud 2018a: fig. 231f) as it does in Arca (s. str.) submontensis . The posterior margin is also very similar with around 15 riblets in the Danian from the Paris Basin and 10–11 in the Danian from Mons Basin. However, a space with 5–6 finer riblets crossed by small growth lamellae on posterior ridge is missing in A. (s. str.) montensis . In addition, the anterior margin is oblique in A. (s. str.) submontensis, whereas it is almost perpendicular to the line of the hinge in A. (s. str.) montensis . In addition, the posterior margin is is almost perpendicular to the line of the hinge in A. (s. str.) submontensis whereas it is slightly oblique in A. (s. str.) montensis .</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6CFF861302FEC5FDBA90E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6EFF86132DFC25FC3F9622.text	F12B8A1C7E6EFF86132DFC25FC3F9622.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Barbatia E. Gray 1842	<div><p>Genus Barbatia E. Gray, 1842</p><p>Type species</p><p>Arca barbata Linnaeus, 1758 by subsequent designation (Gray 1847). Present day, Northeastern Atlantic (Portugal to Morocco) and Mediterranean Sea.</p><p>Stratigraphic range of the genus</p><p>Triassic, Jurassic to present day, cosmopolite (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6EFF86132DFC25FC3F9622	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E6EFF89139AFAE0FE0491E6.text	F12B8A1C7E6EFF89139AFAE0FE0491E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Barbatia undetermined	<div><p>Barbatia sp.</p><p>Fig. 16I</p><p>Barbatia sp. – Merle &amp; Pacaud 2018a: 172, fig. 157d. — Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., LV, EF; Bergères-lès-Vertus ( Mont Aimé); MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 157d); MNHN.F.A66762 .</p><p>Measurements</p><p>DVL: 15 mm; APL (posterior magin broken): 23.5 mm.</p><p>Description</p><p>Shell medium in size large, low and with convex outline. Anterior margin rounded and smoothly passing into ventral margin. Ventral margin rounded near anterior margin. Posterior margin not visible. Shell surface covered with around 10 weak riblets on anterior margin and 30 on ventral margin. Near posterior margin no visible riblets. Riblets crossed by commarginal lines and more weaker on dorsal area than on ventral area.</p><p>Remarks</p><p>No new specimens of this arcid have been collected since the paper of Merle &amp; Pacaud (2018a). Among the Maastrichtian arcids from Marne, Barbatia sp. cannot be confused with Arca (s. str.) submontensis sp. nov. which displays more prominent riblets and a very angular posterior margin (Fig. 16C–F). It is also easily distinguished from Cucullearca supracretacea (d’Orbigny, 1850), which bears stronger ribs (Fig. 16H–N) and a posterior carina. Barbatia sp. differs from B. forchammeri (Lundgren, 1888) from the Maastrichtian of Stevns Klint (Denmark) by having weaker ribs (see Heinberg 1978: 107, fig. 3). Compared to Barbatia gravesi (d’Orbigny, 1850) from the Danian of the Paris Basin (see the syntype in Pacaud 2018a: fig. 199e), Barbatia sp. has higher ribs that are significantly fewer, crossed by lamellar concentric ribs. Barbatia sp. has ribs that are significantly less numerous than in Barbatia dejeari Glibert &amp; van de Poel (1973) from the Danian of Mons. Barbatia rutoti Cossmann, 1908 from the same locality resembles more Barbatia sp. in having riblets that are weaker on the dorsal area than on the ventral area.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E6EFF89139AFAE0FE0491E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E61FF891306FD2BFB419739.text	F12B8A1C7E61FF891306FD2BFB419739.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cucullaearca Conrad 1865	<div><p>Genus Cucullaearca Conrad, 1865</p><p>Type species</p><p>Byssoarca lima Conrad, 1847, accepted as Cucullaearca cuculloides (Conrad, 1833), by subsequent designation (Stoliczka 1871). Late Eocene (Priabonian) to Early Oligocene (Rupelian), Mississippi (USA).</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous to present day, cosmopolite in warm seas (Cox et al. 1969; El Qot 2006).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E61FF891306FD2BFB419739	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E61FF881348FB8FFD4897A7.text	F12B8A1C7E61FF881348FB8FFD4897A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cucullaearca supracretacea (d'Orbigny 1850)	<div><p>Cucullaearca supracretacea (d’Orbigny, 1850)</p><p>Fig. 16J–P</p><p>Arca supracretacea d’Orbigny, 1850: 131, n°38.</p><p>Cucullaearca supracretacea – Pacaud et al. 2000: 872, fig. 2.12. — Merle &amp; Pacaud 2018a: 172, fig. 157b–c; 2018b: 320, fig. 231j–m. — Pacaud 2018a: 253, 256, fig. 199a–b. — Merle 2023: 25.</p><p>Type material</p><p>Syntypes FRANCE – Yvelines • 1 spec., VG, EF; Montainville ( Pont de Falaise); Danian; A. d’Orbigny coll.; figured in Pacaud (2018a: 256, fig. 199a); MNHN.F.R63244 . – Val d’Oise • 1 spec., VG, EF; Vigny; Danian; A. d’Orbigny coll.; figured in Pacaud (2018a: 256, fig. 199b); MNHN.F.A53498 .</p><p>Type locality</p><p>Montainville (Pont de Falaise), Danian coral bed; Yvelines, France.</p><p>Other material (7 specs)</p><p>FRANCE – Marne • 1 spec., historical cast, LV, EF; Vertus ( Faloise); Ch. d’Orbigny coll., ex n°GG. gg2005/6523; figured in Merle &amp; Pacaud (2018a: fig. 157b) and herein Fig 16J; MNHN.F.C02692 • 1 spec., fragment of RV, EF; Vertus (Grand Mont); sandy limestone; MMM coll.; Fig. 16M; MNHN.F.C02694 • 1 spec., RV, EF; same data as for preceding; MMM coll.; Fig. 16 N-O; MNHN.F.C02695 • 1 spec., LV, EF; same data as for preceding; MMM coll.; Fig. 16P; MNHN.F.C02696 • 1 spec., LV, EF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: fig. 157c) and herein Fig. 16K; MNHN.F.A66761 • 1 spec., LV, EF; same data as for preceding; MMM coll.; Fig. 16L; MNHN.F.C02693 • 1 spec., LV, EV; same data as for preceding; MMM coll.; MNHN.F.C02697 .</p><p>Measurements</p><p>MNHN.F.C02692 (LV), DVL: 21 mm; APL: 12 mm. MNHN.F.C02693 (LV), DVL: 18 mm; APL: 9.5 mm. MNHN.F.C02695 (RV), DVL: 14 mm; APL: 7.5 mm. MNHN.F.C02697 (RV), APL: 25 mm, DVL: 15.2 mm. Other specimens incomplete.</p><p>Description</p><p>Shell medium in size, subtriangular and inaequilateral. Anterior margin curved delineated by a small ridge and smoothly passing into ventral margin. Ventral margin curved near anterior margin, and rather straight until posterior margin. Posterior margin angular, oblique to hinge and delineated by transverse ridge starting from the umbo flexure. Beaks high, inclined anteriorly. Shell surface covered with around 7–8 riblets on anterior margin, around 30–40 on ventral margin and 6–8 on posterior margin. Riblets more prominent on posterior and anterior margins than on ventral margin and giving cancellate sculpture in combination with commarginal lines. Ligamental area wide, assymmetrical and low in shape.</p><p>Remarks</p><p>Merle &amp; Pacaud (2018a) presented two specimens and attributed them to Cucullaearca supracretacea from the Danian of the Paris Basin. New specimens collected in 2017 have enabled its description to be completed. The radial sculpture of the Maastrichtian specimens is very similar to that of the Danian specimens (particularly specimen MNHN.F. A53498 figured by Pacaud (2018a: fig. 199b)) and displays the same pattern with riblets more prominent, fewer on posterior and anterior margins than on ventral margin and a prominent transverse ridge starting from the umbo flexure. These similarities, based on more significant material, validate the specific attribution made by Merle &amp; Pacaud (2018a).</p><p>Stratigraphic range</p><p>Maastrichtian to Danian, Paris Basin, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E61FF881348FB8FFD4897A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E60FF88132CFAFCFC09956E.text	F12B8A1C7E60FF88132CFAFCFC09956E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenoides Morch 1853	<div><p>Genus Ctenoides Mörch, 1853</p><p>Type species</p><p>Ostrea scabra Born, 1778 by subsequent designation (Stoliczka 1871). Present day, North Carolina (USA), Florida (USA), Caribbean and Venezuela to North Brazil.</p><p>Stratigraphic range of the genus</p><p>Late Jurassic to present day, cosomopolitan (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E60FF88132CFAFCFC09956E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E60FF8B1329F9BDFCFC94FE.text	F12B8A1C7E60FF8B1329F9BDFCFC94FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenoides vogeli Dhondt 1983	<div><p>Ctenoides vogeli Dhondt, 1983</p><p>Fig. 17A–C</p><p>Inoceramus nobilis Münster in Goldfuss, 1835: 117, pl. 113 fig. 3 (non Inoceramus nobilis Münster in Goldfuss 1835: 109, pl. 109 fig. 4).</p><p>? Ctenoides vogeli Dhondt, 1983: 3, pl. 1 figs 1, 4 (new name for Inoceramus nobilis Münster in Goldfuss 1835: pl. 113, fig. 3).</p><p>Lima nobilis – Bosquet 1860: 384. — Vogel 1895: fig. 12.</p><p>Acesta tecta – Merle &amp; Pacaud 2018a: 174, fig. 158c–d. — Merle 2023: 25 (non Lima tecta Goldfuss, 1835).</p><p>Non Lima tecta – Munier-Chalmas 1897: 83.</p><p>Non Lima texta – Combes 1908: 14. Error of spelling for Lima tecta Goldfuss, 1835 .</p><p>Type material (not examined)</p><p>As the type material of Inoceramus nobilis was destroyed during the 1939–45 war, Dhondt (1983: 3) designated a neotype n° ST13550 (Naturalis, Leiden) using a specimen illustrated by Vogel (1895: 183, fig. 12). In addition, Dhondt (1983) stated that Goldfuss (1835) considered two different species under the binome Inoceramus nobilis, one from the German Liassic of Falkenhagen (Goldfuss 1835: 109, pl. 9 fig. 4) and the other from the Maastrichtian of Sint Pietersberg (Goldfuss 1835: 117, pl. 113 fig. 3). Dhondt (1983) designated clearly a neotype for Inoceramus nobilis from the Maastrichtian, species for which she gave a new name Ctenoides vogeli . Therefore, this neotype should rather be considered the holotype of Ctenoides vogeli in order to fully meet with the recommandation 73A (designation of holotype) of the ICZN (1999).</p><p>Type locality</p><p>Sint Pietersberg near Maastricht, Limburg, The Netherlands.</p><p>Type horizon</p><p>“ E creta tophina ” (“Tuffkreide”), Late Maastrichtian.</p><p>Material examined (4 specs)</p><p>FRANCE – Marne • 1 spec., RV, IF; Vertus ( probably Faloise); MNHN coll.; figured in Merle &amp; Pacaud (2018a: fig. 158c) and herein Fig. 17B; MNHN.F.A66773 • 1 spec., IF, RV; same data as for preceding; Hébert coll.; Fig. 17C; SU.PAL.2017.0.13.290 • 1 spec., IF, LV; Vertus ( Faloise); Hébert coll.; SU.PAL.2017.0.13.291 • 1 spec., EF,? RV; same data as for preceding; Hébert coll.; figured in Merle &amp; Pacaud (2018a: fig. 158d) and herein Fig.17A; UPMC.PAL.2017.0.18 .</p><p>Measurements</p><p>MNHN.F. A66773, DVL: 77 mm, APL: 56 mm. UPMC.PAL.2017.0.18, DVL: 62 mm, APL: 48 mm. Other specimens incomplete.</p><p>Description</p><p>Moderately large shell, slightly opisthocline, large compressed and biauricular. Umbonal angle 64–70°. Lamellar concentric sculpture more developed at end of growth. Around 25 concentric lamellae more or less aequally spaced. No radial sculpture. Auriculae and hinge poorly preserved. No prismatic shell structure.</p><p>Remarks</p><p>Munier-Chalmas (1897) and Combes (1908) also identified this species, as Lima tecta Goldfuss, 1835 at the Bois d’Esmans. Merle &amp; Pacaud (2018a) used this name following the label given by Hébert in the UPMC collection. According to Dhondt (1983), Ctenoides vogeli has many similarities (particularly concentric lamellae) to Cretaceous limids generally classified as Ctenoides, such as Lima tecta Goldfuss, 1835 . However, Ctenoides tecta displays numerous and delicate radial ribs, whereas Ctenoides vogeli is devoid of radial ribs and bears only concentric lamellae (see Dhondt 1983: pl. 1). As the studied specimens have only concentric lamellae and are fairly identical to Ctenoides vogeli regarding the other characters of the shell, their attribution to this species rather than to? C. tecta is more justified.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France; Limburg, The Netherlands.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E60FF8B1329F9BDFCFC94FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E62FF8A1370F9C3FBA49408.text	F12B8A1C7E62FF8A1370F9C3FBA49408.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lecompteus Van de Poel 1959	<div><p>Genus Lecompteus Van de Poel, 1959</p><p>Type species</p><p>Mytilus ornatus Münster, 1837 by original designation. Late Cretaceous (Maastrichtian), Northern Europe.</p><p>Stratigraphic range of the genus</p><p>Jurassic to Late Cretaceous (Maastrichtian), India, Europe (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E62FF8A1370F9C3FBA49408	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E62FF8A133AFE78FB9891F5.text	F12B8A1C7E62FF8A133AFE78FB9891F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Septifer Recluz 1848	<div><p>Genus Septifer Récluz, 1848</p><p>Type species</p><p>Mytilus bilocularis Linnaeus, 1758 by subsequent designation Stoliczka (1871: 366). Present day, Indo-Pacific from East and South Africa to Japan, Australia, New Caledonia and eastern Polynesia.</p><p>Stratigraphic range of the genus</p><p>Triassic to preseent day, tropical and subtropical seas (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E62FF8A133AFE78FB9891F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E62FF8D10BEF89FFE8E948E.text	F12B8A1C7E62FF8D10BEF89FFE8E948E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lecompteus ornatus (Munster)	<div><p>Lecompteus ornatus (Münster in Goldfuss, 1837)</p><p>Fig. 17E–F</p><p>Mytilus ornatus Münster in Goldfuss, 1837: 171, pl. 129 fig. 8.</p><p>Modiola radiatula – Vogel 1895: 30, pl. 2 fig. 6–8 (non Mytilus (Brachiodontes) radiatulus Stoliczka, 1871).</p><p>Lecompteus ornatus – van de Poel 1959: 13.</p><p>Lecompteus aff. radiatulus – Jagt, van Rijsselt &amp; van Rijsselt 2014: 192, fig. 4. — Merle &amp; Pacaud 2018a: 172, fig. 157f–g.</p><p>Type material (not examined)</p><p>Not found, one specimen illustrated in Goldfuss (1837: pl. 129 fig. 8).</p><p>Type locality</p><p>Haldem (Germany)</p><p>Type horizon</p><p>Upper Campanian of Haldem subformation.</p><p>Material examined (2 specs)</p><p>FRANCE – Marne • 1 spec., RV, EF; Vertus ( Faloise); MNHN coll.; figured in Merle &amp; Pacaud (2018a: 172, fig. 157f) and herein Fig. 17E; MNHN.F.A66764 • 1 spec., LV, EF; same data as for preceding; MNHN coll.; figured in Merle &amp; Pacaud (2018a: 172, fig. 157g) and herein Fig. 17F; MNHN.F.A66765 .</p><p>Measurements</p><p>MNHN.F. A66764, APL (anterior area not preserved): 32 mm, DVL: 19.5 mm. MNHN.F. A66765, APL: 30.5 mm, DVL: 13.5 mm.</p><p>Description</p><p>Middle sized shell of mytiliform shape, slightly twisted anteriorly. Umbonal angle 19°. Inverse V-shaped ribs across entire shell. Around 20 ribs more prominent in dorsal area of shell. Internal face unknown.</p><p>Remarks</p><p>This mytiliform shell is reminiscent of Lecompteus ornatus from the upper Campanian of Westphalia in its typical inverse V-shaped ribs covering the entire shell. In addition, Vogel (1895) illustrated a comparable shell from the Maastrichtian of Sint Pietersberg (Limburg) that he identified as Modiola radiatula Stoliczka, 1871, originally described from the Late Cretaceous of India. More recently, Jagt et al. (2014: figs 3a, 4) illustrated well two well-preserved specimens from the Heidelberg Cement Group quarry (Maastrichtian, Meerssen Member) as Lecompteus aff. radiatulus . These shells, particularly the one in Figure 4, are so similar to those from Marne that there is no hesitation in considering them conspecific. Lecompteus radiatula (Stoliczka, 1871) comes from the Campanian-Maastrichtian of India (Ariyalur group [= Arrialoor group in Stoliczka], see Arkaprava et al. 2023). The figures given by Stoliczka (1871: pl. 23 figs 4, 6–7) illustrate shells that are very comparable to the European shells and it is difficult to know whether they belong to one or two different species. However, if they belong to the same species, Mytilus ornatus is senior and has the priority over Mytilus (Brachiodontes) radiatulus (Stoliczka, 1871) . For this reason, the name Lecompteus ornatus is preferred to Lecompteus radiatulus for the European shells.</p><p>Stratigraphic range</p><p>Campanian: North Rhine-Westphalia, Germany. Maastrichtian: Limburg, The Netherlands; Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E62FF8D10BEF89FFE8E948E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E67FF8F1306FA13FCE895FC.text	F12B8A1C7E67FF8F1306FA13FCE895FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isognomon Lightfoot 1786	<div><p>Genus Isognomon Lightfoot, 1786</p><p>Type species</p><p>Ostrea perna Linnaeus, 1767 . Present day, Indo-West Pacific.</p><p>Stratigraphic range of the genus</p><p>Late Triassic to present day, cosmopolite (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E67FF8F1306FA13FCE895FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E67FF8F13E0FEA1FBFB9676.text	F12B8A1C7E67FF8F13E0FEA1FBFB9676.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ostreoidea Rafinesque 1815	<div><p>? Ostreoidea indet.</p><p>Fig. 17G</p><p>Ostreoidea indet. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., LV, EF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02699 .</p><p>Measurements</p><p>H: 21 mm; D: 21.5 mm.</p><p>Description</p><p>Shell partially epigenised. Convex shell with triangular outline dorsally and rounded ventrally. Dorso-posterior area straight slightly concave, dorsal-anterior straight slightly convex. Five fairly spaced concentric lamellae bearing several weak irregular radial folds.</p><p>Remarks</p><p>This shell cannot be confused with specimens of Chama Linnaeus, 1758 which are present in the same beds but display denser concentric lamellae and lack radial folds. In addition, the aragonite shells of Chama are usually dissolved and only calcite shells are preserved in being more or less epigenised. Its concentric lamellae also resemble those of a specimen of Amphidonte obliquata (Pulteney, 1813) from the Cenomanian of Algeria (see Benyoucef et al. 2022: fig. 6a2), a species belonging to the family Gryphaeidae Vialov, 1936 . Chavan (1948) reported another species of oyster, Pycnodonte semiplana (now attributed to the genus Hyotissa Stenzel, 1971), from Vertus on the basis of the material housed in Hébert’s collection. However, Hyotissa semiplana bears stronger radial ribs. Moreover, the material of Hébert’s collection is poorly preserved and it is difficult to validate this identification. While these shells cannot be identified at family rank, they are reminiscent of those of Ostreoidea .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E67FF8F13E0FEA1FBFB9676	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E67FF8E138BF8CCFE0790BC.text	F12B8A1C7E67FF8E138BF8CCFE0790BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isognomon undetermined	<div><p>Isognomon sp.</p><p>Fig. 17H–I</p><p>Isognomon sp. – Merle &amp; Pacaud 2018a: 172, fig. 157e. — Merle 2023: 25.</p><p>Material examined (2 specs)</p><p>FRANCE – Marne • 1 spec., LV, IF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: 172, fig. 157E) and herein Fig. 17H; MNHN.F.A66763 • 1 spec., LV, IF; same data as for preceding; MMM coll.; Fig. 17I; MNHN.F.C02700 .</p><p>Measurements</p><p>MNHN.F. A66763, H (hinge fragment): 7 mm. MNHN.F.C02700, H (hinge fragment): 10 mm.</p><p>Description</p><p>Umbo acute. Hinge incomplete. Ligamental area bearing two (MNHN.F.C02700) to four (MNHN.F. A66763) ligamental grooves. Grooves rectangular, moderately deep. External part of umbo smooth.</p><p>Remarks</p><p>The material is too fragmentary to allow identification at the species level, but the hinge is typical of Isognomon . This genus is present in the European Mesozoic (see Goldfuss 1833: 104, pl. 107) as well as in the northwestern European Danian (France and Belgium) with Isognomon inaequicrenatus (Cossmann, 1908) .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E67FF8E138BF8CCFE0790BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E66FF8E1310FB42FCCB96A3.text	F12B8A1C7E66FF8E1310FB42FCCB96A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lyropecten Conrad 1862	<div><p>Genus Lyropecten Conrad, 1862</p><p>Type species</p><p>Pallium estrellatum Conrad, 1856 by subsequent designation (Dall 1898). Pliocene, California (USA).</p><p>Stratigraphic range of the genus</p><p>Early Cretaceous, Europe (Dhondt 1972) to present day.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E66FF8E1310FB42FCCB96A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E66FFF110BDFA63FCC9969E.text	F12B8A1C7E66FFF110BDFA63FCC9969E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lyropecten ternatus (Munster 1833)	<div><p>Lyropecten ternatus (Münster in Goldfuss, 1833)</p><p>Fig. 17J–K</p><p>Pecten ternatus Münster in Goldfuss, 1833: 52, pl. 9, fig. 13.</p><p>Pecten cicatrisatus Münster in Goldfuss, 1833: 56, pl. 93 fig. 6a–b (junior subjective synonym).</p><p>Lyropecten (Aequipecten?) ternatus – Dhondt 1972: 42, pl. II fig. 3a–b, pl. III fig. la–d.</p><p>Lyropecten ternatus – Niebuhr, Schneider &amp; Wilmsen 2014: 121, fig. 8b, d, g–h.</p><p>Chlamys (Lyropecten) cicatrisata – Merle &amp; Pacaud 2018a: 169, 173, fig. 157k–l.</p><p>For a complete synonymic list see Dhondt (1972: 42) and https://www.molluscabase.org/aphia. php?p=taxdetails&amp;id=1667106</p><p>Type material (not examined)</p><p>Holotype or syntype, Bayerische Staatssammlung für Paläontologie und Geologie according to Dhondt (1972: 48).</p><p>Type locality</p><p>Schandau (Saxony).</p><p>Type horizon</p><p>Quader sandstein of Schandau (?Turonian), Germany.</p><p>Material examined (5 specs)</p><p>FRANCE – Marne • 1 spec., RV, IF; Vertus ( Faloise); MNHN coll.; figured in Merle &amp; Pacaud (2018a: 173, fig. 157k) and herein Fig. 17J; MNHN.F.A66768 • 1 spec., RV, IF; same data as for preceding; MNHN coll.; figured in Merle &amp; Pacaud (2018a: 173, fig. 157l) and herein Fig. 17K; MNHN.F.A66769 • 3 specs (internal valves, 2 specs incomplete, 1 spec. subcomplete); same locality; Hébert coll.; SU.PAL.2017.0.13.292.1 to SU.PAL.2017.0.13.292.3 .</p><p>Measurements</p><p>MNHN.F. A66768, H (anterior part incomplete): 32 mm, D: 28.5 mm. MNHN.F. A66769, H: 39 mm, D: 35 mm. UPMC.PAL, H: 30 mm, D: 31 mm.</p><p>Description</p><p>Right valve (internal face), shell rather wide. Umbonal angle 72°. 7–8 broad ribs with plane spaces between ribs. Auricles unequal and small. Anterior auricle: winglike and elongated; posterior auricle: triangular; acute to right angled. Byssal sinus deep.</p><p>Remarks</p><p>Merle &amp; Pacaud (2018a) identified these specimens from Vertus as Chlamys (Lyropecten) cicatrisata (Goldfuss, 1833), a species originally described from Maastrichtian of Sint Pietersberg (Limburg). Although they consist solely of shells showing their internal faces, their shape and their 7–8 ribs are fairly typical of Pecten cicatrisatus . Following Dhondt (1972) and Niebuhr et al. (2014), Pecten cicatrisatus is considered to be a junior subjective synonym of Pecten ternatus .</p><p>Stratigraphic range</p><p>This species has a large European stratigraphic range from the Cenomanian to the Maastrichtian (Dhondt 1972). In the Maastrichtian, it has been recorded in Belgium, The Netherlands, Bulgaria and France (Manche), see Dhondt (1972). The Paris Basin is added.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E66FFF110BDFA63FCC9969E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E19FFF1130EF9EFFAA69439.text	F12B8A1C7E19FFF1130EF9EFFAA69439.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lyriochlamys Sobetski 1977	<div><p>Genus Lyriochlamys Sobetski, 1977</p><p>Type species</p><p>Pecten fissicosta Etheridge, 1881 by original designation. Late Cretaceous (Cenomanian), Great Britain.</p><p>Stratigraphic range of the genus</p><p>Late Triassic (Rhetian) to Late Cretaceous (Maastrichtian according Waller &amp; Marincovich 1992).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E19FFF1130EF9EFFAA69439	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E19FFF31365F888FE7D92B3.text	F12B8A1C7E19FFF31365F888FE7D92B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lyriochlamys faujasii (Defrance 1826)	<div><p>Lyriochlamys faujasii (Defrance, 1826)</p><p>Fig. 18A–D</p><p>Pecten faujasii Defrance, 1826: 265 .</p><p>Chlamys faujasii – Dhondt 1973a: 6, pl. 1 fig. 2a–e</p><p>Lyriochlamys faujasii – Waller &amp; Marincovich 1992: 223.</p><p>Chlamys dentata – Merle &amp; Pacaud 2018a: 169, fig. 157i–j (non Pecten dentatus S. Nilsson, 1827: 20).</p><p>For a complete synonymic list see Dhondt (1973a: 6–8).</p><p>Type material (not examined)</p><p>The type material is based on Faujas’s collection but this material is apparently lost (Dhondt 1973a: 8). However, Dhondt (1973a) stated that: “… it is not necessary to select a neotype: the figure in Faujas and the numerous topotypes leave no doubt as to the content of the species”.</p><p>Type locality</p><p>Sint Pietersberg (Limburg), Maastrichtian.</p><p>Material examined (10 specs)</p><p>FRANCE – Marne • 1 spec., LV, IF; Vertus ( Faloise); MNHN coll.; figured in Merle &amp; Pacaud (2018a: 173, fig. 157i) and herein Fig. 18A; MNHN.F.A66766 • 1 spec., LV, IF; same data as for preceding; Hébert coll.; Fig. 18B; SU.PAL.2017.0.13.293 • 1 spec., EF, subcomplete RV, same data as for preceding; Hébert coll.; Fig. 18D; SU.PAL.2017.0.13.294 • 1 spec., EF, subcomplete LV, same data as for preceding; Hébert coll.; SU.PAL.2017.0.13.295 • 3 specs, EF incomplete; same data as for preceding; Hébert coll.; SU.PAL.2017.0.13.296.1 to SU.PAL.2017.0.13.296.3 • 1 spec., LV, EF; same data as for preceding; MNHN coll.; figured in Merle &amp; Pacaud (2018a: 173, fig. 157j) and herein Fig. 18C; MNHN.F.A66767 • 1 spec., IF juv., incomplete; Vertus ( Grand Mont); basal biocalcarenite with pectinids; MMM coll.; MNHN.F.C02704 • 1 spec., IF incomplete; same data as for preceding; MMM coll.; MNHN.F.C02705 .</p><p>Measurements</p><p>MNHN.F. A66766, DVL: 41 mm, APL: 32.5 mm. MNHN.F. A66767, DVL: 34.2 mm, APL: 27.8. 5 mm. SU.PAL.2017.0.13.293, DVL: 38 mm, APL: 31.5 mm. U.PAL.2017.0.13.294, DVL: 43 mm, APL: 33 mm. SU.PAL.2017.0.13.295 (LV), DVL: 31 mm, APL: 26 mm.</p><p>Description</p><p>Middle sized shell. Valves elongated, prosocline and flattened. Left valve with undivided 37–41 ribs, bipartite on extremities of anterior and posterior areas of ventral margin. Right valves with 40 ribs. Ribs most undivided, but irregularly bipartite near center of ventral margin. Close set scales on ribs, denser near dorsal area. Anterior auricle triangular acute-angled. Posterior auricle not seen, probably smaller.</p><p>Remarks</p><p>These shells were initially identified as Pecten dentatus S. Nilsson, 1827 by Hébert (in collection) and by Chavan (1948). Dhondt (1973a) reviewed the Pectinidae of this group and concluded that the differences between Lyriochlamys dentata (S. Nilsson, 1827) and L. faujasii are subtle. However, she mentioned that L. dentata differs from L. faujasii in having a higher radial rib number and more developed spines rather than scales. The specimens from Marne have between 37 and 41 ribs, which is more in line with the variation seen in L. dentata . On the other hand, the specimens from Marne have more scales than spines, as in L. faujasii . According to Dhondt (1973a), L. dentata became extinct in the Campanian, whereas L. faujasii persisted in the upper Maastrichtian. It therefore seems preferable to use provisionally the binome L. faujasii for the Paris Basin material.</p><p>Stratigraphic range</p><p>This species has a large European stratigraphic range from the Turonian to the Maastrichtian (Dhondt 1973a). In the Maastrichtian, it has been recorded in Belgium and The Netherlands, see Dhondt (1973a). The Paris Basin is added.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E19FFF31365F888FE7D92B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1BFFF3133AFDCCFC449039.text	F12B8A1C7E1BFFF3133AFDCCFC449039.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neithea Drouet 1825	<div><p>Genus Neithea Drouët, 1825</p><p>Type species</p><p>Pecten aequicostatus Lamarck, 1819 by subsequent designation (Dhondt 1973b: 7). Early Cretaceous (Albian) to Late Cretaceous (Turonian), Western Europe.</p><p>Stratigraphic range of the genus</p><p>Early Cretaceous to Late Cretaceous, cosmopolite (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1BFFF3133AFDCCFC449039	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1BFFF51371FC88FEEF91AF.text	F12B8A1C7E1BFFF51371FC88FEEF91AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neithea regularis (von Schlotheim 1813)	<div><p>Neithea regularis (Schlotheim, 1813)</p><p>Fig. 18E–F</p><p>Pectinites regularis Schlotheim, 1813: 112 (refers to an illustration in Faujas de Saint-Fond 1799: pl. 23 fig. 2).</p><p>Pecten (Neithea) regularis – Pervinquière 1912: 138.</p><p>Neithea regularis – Torriani 1942: 88, pl. 6 fig. 1 a–b. — Dhondt 1973b: 20, pl 1 fig. 3, pl. 2 fig. 1 a–d. — Merle &amp; Pacaud 2018a: 169, fig. 157h.</p><p>For a detailed synonymic list, see Dhondt (1973b: 20–22).</p><p>Type material (not examined)</p><p>Syntype represented by Faujas de Saint-Fond (1799: pl. 23 fig. 2), see Dhondt (1973b). The material was originally deposited at the Muséum national d’Histoire naturelle, Paris, but is probably lost.</p><p>Type locality</p><p>Sint Pietersberg (Limburg), Maastrichtian.</p><p>Material examined (2 specs)</p><p>FRANCE – Seine-et-Marne • 1 spec., partial internal mould of a RV; Esmans ( Bois d’Esmans); Hébert coll.; figured in Merle &amp; Pacaud (2018a: 169, fig. 157h) and herein Fig. 18E; UPMC.PAL 2017.0.17 • 1 spec., LV, IF; same data as for preceding; Hébert coll.; Fig. 18F; SU.PAL.2017.0.13.297 .</p><p>Measurements</p><p>UPMC.PAL2017.0.17, DVL: 57 mm; APL: 45 mm. SU.PAL.2017.0.13.297, DVL (umbonal part incomplete): 28 mm, APL: 34 mm.</p><p>Description</p><p>Right valve convex, equilateral. Umbonal angle 80°. Trace of around 20 distinct low ribs, probably more (25–30). Parts of preserved shell showing smooth ribs. Left valve flattened with 26 ribs. Ribs narrow. Intercalary ribs undistinguishable.</p><p>Remarks</p><p>Although it is not very well preserved, this pectinid convex valve is reminiscent of Neithea regularis in its shape, and its low and smooth ribs (see Dhondt 1973b). Moreover, the number and the shape of the ribs of the left valve are similar to specimens from Sint Pietersberg (Limburg) described by Dhondt (1973b).</p><p>Stratigraphic range</p><p>This species has a wide European stratigraphic range, from the Cenomanian to the Maastrichtian (Dhondt 1973b). In the Maastrichtian, it has been recorded in Belgium, The Netherlands, Bulgaria and France (Calcaire à Baculites, Manche), see Dhondt (1973b). The Maastrichtian of the Paris Paris is a new record for this species.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1BFFF51371FC88FEEF91AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1DFFF51352FD7CFCD794C6.text	F12B8A1C7E1DFFF51352FD7CFCD794C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neithea quinquecostata (J. Sowerby 1814)	<div><p>Neithea quinquecostata (J. Sowerby, 1814)</p><p>Fig. 18G–H</p><p>Pecten quinquecostatus J. Sowerby, 1814: 122, pl. 56 figs 4–8.</p><p>Janira quinquecostata – d’Orbigny 1847: pl. 444 figs 1–5.</p><p>Vola quinquecostata – Stoliczka 1871: 437, pl. 31 figs 1–6, pl. 37 figs 4–9.</p><p>Neithea quinquecostata – Bayle 1878: pl. 122 figs 2–3. — Dhondt 1973b: 29, pl. 2 fig. 2 a–c. — Merle &amp; Pacaud 2018a: 167.</p><p>For a detailed synonymic list, see Dhondt (1973b: 29–32).</p><p>Type material (not examined)</p><p>Lectotype NHMUK n°43324 designated by Dhondt (1973b: 32).</p><p>Type locality</p><p>Chute (England), Greensand bed (lower Cenomanian).</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., RV, EF; Vertus ( Faloise); coll. Hébert; SU.PAL.2017.0.13.298 .</p><p>Measurements</p><p>SU.PAL.2017.0.13.298, DVL: 18.5 mm, APL: 16.5 mm.</p><p>Description</p><p>Right valve. Convex valve, equilateral. Umbonal angle 72°. Six prominent ribs. Between a pair of prominent ribs, 3- 4 intercalary ribs. Auricles triangular, slightly ornamented with fine riblets.</p><p>Remarks</p><p>Although the top of this right valve is damaged, its sculpture consisting of six ribs and 3–4 intercostal spaces is reminiscent of Neithea quinquecostata known from the Maastrichtian of Maastricht. Neithea striatocostata (Goldfuss, 1833) differs by less regular intercalary ribs (usually two). In addition, the ribs of N. striatocostata are always radially striated and this never occurs in N. quinquecostata .</p><p>Stratigraphic range</p><p>This species has a large European stratigraphic range from the Barremian to the Maastrichtian (Dhondt 1973b). In the Maastrichtian, it has been recorded in Belgium, The Netherlands and Bulgaria, see Dhondt (1973b). The Maastrichtian of the Paris is a new record.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1DFFF51352FD7CFCD794C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1CFFF41314FB09FB5696C6.text	F12B8A1C7E1CFFF41314FB09FB5696C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plicatula Lamarck 1801	<div><p>Genus Plicatula Lamarck, 1801</p><p>Type species</p><p>Spondylus plicatus Linnaeus, 1764 by subsequent designation (F.C. Schmidt 1818). Present day, Red Sea, eastern Africa, Chagos Islands, Japan, China, Philippines, Indonesia and Australia.</p><p>Stratigraphic range of the genus</p><p>Middle Triassic (Ladinian) to present day, subtropical and tropical seas (Cox et al. 1969)</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1CFFF41314FB09FB5696C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1CFFF413EAFE86FC27971D.text	F12B8A1C7E1CFFF413EAFE86FC27971D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spondylidae J. E. Gray 1826	<div><p>? Spondylidae indet.</p><p>Fig. 18I</p><p>Spondylidae indet. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., EF incomplete; Bergères-lès-Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; MNHN.F.C02706 .</p><p>Measurements</p><p>DVL (incomplete): 27 mm.</p><p>Description</p><p>Fragment of a convex shell (?cemented valve). Outline elongate dorso-ventrally with convex ventral margin. Surface with dense concentric lamellae (around 15 observed) crossed by delicate radial ribs between concentric lamellae.</p><p>Remarks</p><p>The shape and sculpture of this fragment are reminiscent of shells of Spondylidae . It does not appear to be a shell of Plicatula Lamarck, 1801, which is found in the same beds. The shells of Plicatula from Marne (see below) are flatter and bear a stronger radial sculpture.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1CFFF413EAFE86FC27971D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E1CFFF91335F9C5FDBA9143.text	F12B8A1C7E1CFFF91335F9C5FDBA9143.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plicatula pacaudi Merle 2026	<div><p>Plicatula pacaudi sp. nov.</p><p>urn:lsid:zoobank.org:act: BC5BECCA-0400-4833-A675-D631B3902036</p><p>Fig. 18J–K, 19A–D</p><p>Plicatula sp. – Merle &amp; Pacaud 2018a: 174, fig. 158a–b. — Merle 2023: 25.</p><p>Etymology</p><p>Species dedicated to Jean-Michel Pacaud for his malacological work on Cenozoic molluscs.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations, Maastrichtian.</p><p>Material examined</p><p>Holotype FRANCE – Marne • RV, EF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; Fig. 19A a 1 –Ba1; MNHN.F.C02707.</p><p>Paratypes (12 specs) FRANCE – Marne • 1 spec., RV, EF; same data as for holotype; MMM coll.; Fig. 19 Aa2–Ba2; MNHN.F.C03009 • 1 spec., RV, EF; same data as for holotype; MMM coll.; Fig. 19 Aa3; MNHN.F.C02987 • 1 spec., RV, EF; same data as for holotype; MMM coll.; Fig. 19A 4; MNHN.F.C02988 • 1 spec., RV, IF, incomplete; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 174, 158a) and herein Fig. 18J; MNHN.F.A66771 • 1 spec., RV, IF, incomplete; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 174, fig. 158b) and herein Fig. 18K; MNHN.F.A66772 • 1 spec.,?LV, EF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 172, 157m); MNHN.F.A66770 • 1 spec., RV, EF; same data as for holotype; MMM coll.; Fig. 19C; MNHN.F.C02989 • 1 spec., RV, IF; same data as for holotype; MMM coll.; Fig. 19D; MNHN.F.C02990 • 1 spec., RV, EF; same data as for holotype; MMM coll.; MNHN.F.C02991 • 1 spec., RV, IF; same data as for holotype; MMM coll.; MNHN.F.C02992 • 1 spec., RV, EF; same data as for holotype; MMM coll.; MNHN.F.C02993 • 1 spec., RV, EF; same data as for holotype; MMM coll.; MNHN.F.C02994 .</p><p>Measurements</p><p>Holotype MNHN.F.C02707, DVL: 18 mm, APL: 15 mm. Paratype MNHN.F.C03010, DVL: 20 mm, APL: 15 mm. Paratype MNHN.F.C02987, DVL: 17 mm, APL: 19.8 mm. Paratype MNHN.F.C02988, DLV: 20 mm, APL: 20 mm. Paratype MNHN.F. A66770, DVL: 19.7 mm, APL: 15.8 mm. Paratype MNHN.F.C02989, DVL: 20.2 mm, APL: 18 mm. Paratype MNHN.F.C02990, DVL: 20 mm, APL: 19.5 mm. Paratype MNHN.F.C02991, DVL: 18.8 mm, APL: 16.5 mm. Paratype MNHN.F.C02992, DVL: 14.5 mm, APL: 16.7 mm.</p><p>Description</p><p>Valves of medium size (DVL 20 mm), curved, ostreiform, moderately inaequilateral. Right valve convex, area of attachment in dorsal posterior beak region not visible. Numerous (between 45–55), very closely spaced, fine radial ribs with some intercalated in early growth stage; left valve flat, sculptured by numerous (around 50), closely spaced very fine radial ribs. At end of growth, radial ribs slightly tubular with numerous scales. Hinge well developed, cardinal area small with two teeth, lateral teeth larger, clearly diverging from the cardinal teeth.</p><p>Comparisons</p><p>Plicatula pacaudi sp. nov. can be easily distinguished from P. megaera (d’Orbigny 1850) from the Danian of the Paris Basin (see Merle &amp; Pacaud: 2018b: fig. 232f–g) in having significantly thinner and more numerous radial ribs. Plicatula megaera resembles P. cossmanni Cossmann, 1887 from the Ypresian and P. parisiensis Deshayes, 1858 from the Lutetian (both from the Paris Basin) in its few ribs, its hinge strongly developed, its cardinal area very small with two reduced cardinal teeth and its lateral teeth larger, clearly parallel to the cardinal teeth. This type of hinge is similar to that of the type species of the genus ( P. plicata), but differs from that of P. pacaudi, whose cardinal teeth are less reduced and whose lateral teeth are not parallel to the cardinal teeth, but divergent. A radial sculpture composed of numerous ribs more similar to that of P. pacaudi is found mainly in Cretaceous species such as P. gurgitis Pictet &amp; Roux, 1852 from the Albian of Folkestone (see Woods 1901: pl. 25 figs 13–21), P. batnensis Coquand, 1880 from the upper Cenomanian of southern Algeria (see Benyoucef et al. 2022: fig. 10c–g), P. hirsuta Coquand, 1880 from the upper Cenomanian of southern Algeria (see Pervinquière 1912: pl. 10 fig. 12) and P. ventilabrum Coquand, 1880 from the Coniacian of Tunisia (see Pervinquière 1912: pl.10 fig. 7). Plicatula simiensis Squires &amp; Saul, 1998 from the upper Paleocene of California also displays numerous (around 70) and fairly fine ribs and its shell outline also resembles that of P. pacaudi . These species, whose sculpture resembles that of P. pacaudi, have been illustrated without figure of their hinge, and therefore it is not known whether it is similar to that typical of P. plicata or P. pacaudi</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E1CFFF91335F9C5FDBA9143	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E11FFF910E9FCB3FABF97B8.text	F12B8A1C7E11FFF910E9FCB3FABF97B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wolfgangella Jagt, M. R. Cooper & Jagt-Yazykova 2022	<div><p>Genus Wolfgangella Jagt, M.R. Cooper &amp; Jagt-Yazykova, 2022</p><p>Type species</p><p>Wolfgangella neilpearti Jagt, M.R. Cooper &amp; Jagt-Yazykova, 2022 by original designation. Late Cretaceous (Maastrichtian), The Netherlands.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Maastrichtian), The Netherlands (Jagt et al. 2022) and Paris Basin (this work).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E11FFF910E9FCB3FABF97B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E11FFF81036FB0FFCEF9047.text	F12B8A1C7E11FFF81036FB0FFCEF9047.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wolfgangella neilpearti Jagt, M. R. Cooper & Jagt-Yazykova 2022	<div><p>Wolfgangella cf. neilpearti Jagt, M.R. Cooper &amp; Jagt-Yazykova, 2022</p><p>Fig. 19E–G</p><p>Type material of Wolfgangella neilpearti</p><p>Holotype (not examined) THE NETHERLANDS – Southern Limburg • LV, EF; Sint-Pietersberg area; Meerssen Member (see Jagt et al. 2022: 21 for the type locality); NHMM 00381a.</p><p>Paratype (not examined) THE NETHERLANDS – Southern Limburg • 1 spec., fragment of LV, EF and its internal mould; same data as for holotype; NHMM JJ 13544a/b .</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., LV, EF, Vertus ( Grand Mont); basal biocalacarenite with pectinids; MMM coll.; MNHN.F.C03010 .</p><p>Measurements</p><p>DVL: 20 mm; APL: 22 mm.</p><p>Description</p><p>Shell medium in size, trapezoidal, almost as long as high (DVL/APL = 0.90) and moderately inflated. Small, anteriorly positioned umbo. Convex anterior margin passing evenly into convex ventral margin. Posterior margin obliquely truncate, posterodorsal margin almost straight. Escutcheon carina lacking; escutcheon area discriminated by slight inflexion in commarginal ribs. On flank, 14–15 strong, nontuberculate, commarginal ribs about as wide as interspaces. Marginal carina not prominent, but forming a line marked by strong angulosity of ribs when passing on dorso-posterior margin. Ribs tighter on dorso-posterior margin than on ventral margin.</p><p>Remarks</p><p>This shell from Vertus has an outline and a shape with strong commarginal ribs that are very similar to those of the holotype of Wolfgangella neilpearti from the upper Maastrichtian of Maastricht, but it bears fewer ribs (14–15 ribs for DVL = 20 mm instead of 20 ribs for DVL = 14 mm). Therefore, this shell is identified as W. cf. neilpearti due to these differences. Wolgangella ignota Jagt et al., 2022 is another species from the upper Maastrichtian of Maastricht, but it differs from W. neilpearti and W. cf. neilpearti by having its smaller size (DVL = 10 mm) and by having beaded costellae on the escutcheon. Compared to W. cf. neilpearti, W. ignota displays 12–15 ribs, but its DVL = 10 mm instead of 20 mm.</p><p>Stratigraphic range</p><p>Maastrichtian:? Limburg, The Netherlands; Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E11FFF81036FB0FFCEF9047	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E10FFF81341FB95FBE29676.text	F12B8A1C7E10FFF81341FB95FBE29676.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baluchicardia Rutsch & Schenk 1940	<div><p>Genus Baluchicardia Rutsch &amp; Schenk, 1940</p><p>Type species</p><p>Cardita beaumonti d’Archiac &amp; Haime, 1854 by original designation. Paleocene (Danian), Pakistan.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous to Paleocene, India, Africa and North America (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E10FFF81341FB95FBE29676	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E10FFFA135AFAB5FDBA91F4.text	F12B8A1C7E10FFFA135AFAB5FDBA91F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Baluchicardia hebertiana (d'Orbigny 1850)	<div><p>Baluchicardia hebertiana (d’Orbigny, 1850)</p><p>Fig. 19H–L</p><p>Cardita hebertiana d’Orbigny, 1850: 130, n°32 (partim).</p><p>Cardium porulosum – Ch. d’Orbigny 1836: 284 (non Solander in Brander, 1766).</p><p>? Venericardia roedeli – Chavan 1948: 567 (non Ravn, 1939: 31).</p><p>Glyptoactis (Baluchicardia) hebertiana – Pacaud &amp; Merle 2016: pl. 1 figs 1–9. — Merle &amp; Pacaud 2018a: 165, 176, fig. 159a–d. — Pacaud 2018b: 251. — Merle 2023: 24, fig. 2a–b.</p><p>For a complete synonymic list, see Pacaud &amp; Merle (2016: 44).</p><p>Type locality</p><p>Vertus (Marne), France.</p><p>Type horizon</p><p>Sandy limestone of Vertus, Maastrichtian.</p><p>Type material</p><p>Lectotype FRANCE – Marne • LV, EF; Vertus; A. d’Orbigny coll.; designated by Pacaud &amp; Merle (2016: 44, 46, pl. 1 fig. 2) and herein Fig. 19H; MMM coll.; MNHN.F.R63240.</p><p>Paralectotypes (5 specs) FRANCE – Marne • 1 spec., LV, EF; Vertus; sandy limestone; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 3; MNHN.F.R63241 • 1 spec., RV, EF; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 4); MNHN.F.A46194 • 1 spec., LV, EF; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 5); MNHN.F.A46195 • 1 spec., LV, EF; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 1a–b); MNHN.F.A46243 • 1 spec., RV, EF juvenile; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 6) and herein Fig. 19J; MNHN.F.A57275. All paralectotypes from A. d’Orbigny coll.</p><p>Other examined material (5 specs)</p><p>FRANCE – Marne • 1 spec., LV, EF; Vertus ( Grand Mont), sandy limestone; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 9); MNHN.F.A57131 • 1 spec., LV, IF; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 8) and herein Fig. 19L; MNHN.F.A57132 • 1 spec., LV, EF; same data as for preceding; MMM coll.; figured in Pacaud &amp; Merle (2016: 46, pl. 1 fig. 7) and herein Fig. 19I; MNHN.F.A57133 • 1 spec., LV, IF; Bergères-lès-Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: 176, fig. 159d) and herein Fig. 19K; MNHN.F.A66778 • 1 spec., LV, EF; same data as for preceding; MMM coll.; figured in Merle (2023: 24, fig. 2a–b); MNHN.F.A77757. Many imprints have not been molded and are not counted here .</p><p>Measurements</p><p>Lectotype, DVL: around 19 mm, APL: around 21 mm. Paralectotype MNHN.F.R63241, DVL: around 23 mm, APL: around. 25 mm. Paralectotype MNHN.F.A46194, DVL: around 26 mm, APL: around 28 mm. Paralectotype MNHN.F. A46195, DVL: around 14 mm, APL: around 15 mm. Paralectotype MNHN.F.A46243, DVL: 26 mm, APL: 28 mm. Paralectotype MNHN.F.A57275, DVL: around 11 mm, APL: around 14 mm. Measurements from Pacaud &amp; Merle (2016: 44-45).</p><p>Description</p><p>Original description (d’Orbigny 1850: 130): “Espèce quadrangulaire, renflée, ornée d’environ 26 grosses côtes rayonnantes, saillantes, carénées, pourvues dessus d’une série de tubercules, et latéralement d’une saillie longitudinale”. [English translation: “Quadrangular species, bulging, with approximately 26 large, protruding, keeled ribs, featuring a series of tubercles on top and a longitudinal protrusion on the sides”.]</p><p>Descriptive complement: 20–27 ribs separated by deep interspaces. Ribs tripartite in larger specimens. Inner margin strongly crenulated (around 20 subtriangular crenulations).</p><p>Remarks d’Orbigny (1850) described Cardita hebertiana based on type material from Vertus and two Danian localities, Le Port-Marly (Yvelines) and Meudon (Hauts-de-Seine). Pacaud &amp; Merle (2016) showed that the type material included four different species. However, the diagnosis given by d’Orbigny (1850) corresponds perfectly to the specimens from Vertus, and a lectotype (MNHN.F.R63240) from this locality was thus designated. The Danian specimens represented the following species: Venericor duponti (Cossmann, 1908) from Meudon, Venericardia elisabethae Pacaud &amp; Merle, 2016 and Paraglans lydiae Pacaud &amp; Merle, 2016 from Port Marly. Chavan (1948) reported Venericardia roedeli Ravn, 1939 from the sandy limestone with shell concentrations of Mont Aimé, a species present in the Danian of Copenhagen (Denmark). This species is easily distinguished from Baluchicardia hebertiana by its much more rounded outline and its flatter, almost smooth ribs. Chavan (1948) apparently did not know the work of d’Orbigny (1850) and did not compare V. roedeli with B. hebertiana . As Chavan refers to the sandy limestone with shell concentration and carditid imprints of Mont Aimé where B. hebertiana is one of the most common species and the only Carditidae collected in this bed, it is very likely that what he identified as V. roedeli is in fact B. hebertiana . Baluchicardia hebertiana is a mollusc characteristic of the Maastrichtian of Marne. It occurs from the lower beds at the base of the Vertus section (Grand Mont) to the upper beds of the Mont Aimé section.</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E10FFFA135AFAB5FDBA91F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E12FFFA1365FCB3FBC3975D.text	F12B8A1C7E12FFFA1365FCB3FBC3975D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathytormus R. B. Stewart 1930	<div><p>Genus Bathytormus R.B. Stewart, 1930</p><p>Type species</p><p>Crassatella protexta Conrad, 1832 by original designation. Middle Eocene (Bartonian), Alabama (USA).</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous to present day, subtropical and tropical seas (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E12FFFA1365FCB3FBC3975D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E12FFFD1303FB6CFDBA96CE.text	F12B8A1C7E12FFFD1303FB6CFDBA96CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathytormus sparnacensis Merle 2026	<div><p>Bathytormus sparnacensis sp. nov.</p><p>urn:lsid:zoobank.org:act: BD7CB6BB-0A7C-47E9-8F11-9D27FB4A89DA</p><p>Fig. 20A–D</p><p>Bathytormus sp. – Merle &amp; Pacaud 2018a: 175, fig. 158e–h. — Merle 2023: 25.</p><p>Etymology</p><p>From Épernay (Latin name: ‘ Sparnacum ’), the city near the type locality.</p><p>Type locality</p><p>Bergères-lès-Vertus (Mont Aimé), Marne, France.</p><p>Type horizon</p><p>Sandy limestone with shell concentrations.</p><p>Type material</p><p>Holotype FRANCE – Marne • RV, EF; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: 175, fig. 158e) and herein Fig. 20A; MNHN.F.A66774.</p><p>Paratypes (3 specs) FRANCE – Marne • 1 spec., LV, EF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 175, fig. 158h) and herein Fig. 20C; MNHN.F.A66777 • 1 spec., RV, EF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 175, fig. 158f) and herein Fig. 20D; MNHN.F.A66775 • 1 spec., LV, EF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 175, fig. 158g) and herein Fig. 20B; MNHN.F.A66776 .</p><p>Measurements</p><p>Holotype, DVL: 12 mm, APL: 14.7 mm. Paratype MNHN.F. A66777, DVL: 22 mm, APL: 33.5 mm. Paratype MNHN.F. A66775, APL: around 18 mm, DVL: incomplete. Paratype MNHN.F. A66776, APL: 22 mm, DVL: 34 mm.</p><p>Description</p><p>Middle sized shell, moderately elongate posteriorly and not inflated. Antero-dorsal margin concave and postero-dorsal margin slightly straight; anterior and ventral margins rounded, posterior margin almost straight making large angle at transition with postero-dorsal margin and angulose when joining ventral margin. Posterior ridge weakly angulate near umbo, becoming smooth and connected to angulosity of ventral margin. On posterior ridge, seven undulating commarginal ribs in umbonal region and around ten more widely spaced and lamellose major commarginal ribs in the postero-ventral margin. On antero-ventral margin, around 20 undulating and small commarginal ribs corresponding to intercalated ribs between major ribs. Around two intercalated ribs between two major ribs.</p><p>Remarks</p><p>Bathytormus sparnacensis sp. nov. is comparable to B. pisolithicus (d’Orbigny, 1850) in its shape and in the organisation of its commarginal sculpture displaying intercalated ribs between major ribs on antero-ventral margin. However, the ribs of B. sparnacensis are more widely spaced and less numerous (around 17 on postero-ventral margin and 27 on antero-ventral margin instead of respectively 36 and 50 in B. pisolithicus). This sculptural difference had already been mentioned by Merle &amp; Pacaud (2018a: 165) who compared B. sparnacensis with the syntype of B. pisolithicus (MNHN.F. A53495) from the Danian of Meudon (Pacaud 2018a). Combes (1908) identified Crassatella pisolithica d’Orbigny, 1850 (= B. pisolithicus) from the Bois d’Esmans locality. Nevertheless, without material or description it is difficult to know which species this is, but it could be B. sparnacensis .</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E12FFFD1303FB6CFDBA96CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E15FFFF137BF99EFBDE9155.text	F12B8A1C7E15FFFF137BF99EFBDE9155.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euheterodonta Giribet & Distel 2003	<div><p>Euheterodonta indet. (tellinoid shape).</p><p>Fig. 20E</p><p>?Tellinoidea indet. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., internal mould, RV; Vertus ( Grand Mont), sandy limestone; MMM coll.; MNHN.F.C02995 .</p><p>Measurements</p><p>DVL: 5 mm, APL (incomplete): 10 mm.</p><p>Description</p><p>Shell, small and flat. Elongated and inequilateral outline with APL twice as large as DVL. Antero-dorsal margin straight. Dorso-posterior margin almost straight, but slightly concave near beak. Ventral margin moderately curved. Anterior and posterior margins incomplete.</p><p>Remarks</p><p>This small, flat, elongated shell with an inequilateral outline is somewhat similar to those of Tellinoidea Blainville, 1814 ( Psammobiidae J. Fleming, 1828 or Tellinidae Blainville, 1814). Its shape is also reminiscent of Nuculana Link, 1807, but the anterior area is shorter in this genus. The material is not sufficiently well preserved to allow for a more precise taxonomic attribution.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E15FFFF137BF99EFBDE9155	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E17FFFF1355FD55FD7F96D4.text	F12B8A1C7E17FFFF1355FD55FD7F96D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undetermined undetermined	<div><p>Euheterodonta indet. (? Recticardo shape)</p><p>Fig. 20F</p><p>Recticardo sp. – Merle &amp; Pacaud 2018a: 176, fig. 159k.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., LV, internal mould; Vertus ( Grand Mont), sandy limestone; MMM coll.; figured in Merle &amp; Pacaud (2018a: 176, fig. 159k) and herein Fig. 20; MNHN.F.A66784 .</p><p>Measurements</p><p>DVL: 32.8 mm, APL: 34 mm.</p><p>Description</p><p>Internal moulds showing right sides. Flattened shell, moderately large, subquadrangular. Dorso-posterior margin slightly convex, oblique. Posterior margin convex perpendicular to dorso-ventral axis. Ventral margin rounded. Anterior margin rather straight dorsally, rounded ventrally. Antero-dorsal margin slightly concave.</p><p>Remarks</p><p>It is difficult to identify this internal mould of Lucinidae J. Fleming, 1828 precisely, but its flat, moderaltely subquadrangular shape is slightly reminiscent of Recticardo Cossmann, 1908, a genus found in the Late Cretaceous of Europe (Cox et al. 1969) and in the Danian of the Paris and Mons basins (Cossmann 1908; Pacaud 2018a). However, this mould has a rather central almost rectigyrate, blunt umbo, which does not fit Recticardo very well.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E17FFFF1355FD55FD7F96D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E17FFFE1365F9D4FC27906E.text	F12B8A1C7E17FFFE1365F9D4FC27906E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euheterodonta Giribet & Distel 2003	<div><p>Euheterodonta indet. (?lucinoid shape)</p><p>Fig. 20G–J</p><p>Material examined (6 specs)</p><p>FRANCE – Marne • 1 spec., LV, EF; Vertus ( Grand Mont), sandy limestone; MMM coll.; Fig. 20G; MNHN.F.C02996 • 1 spec., LV, EF; same data as for preceding; MMM coll.; Fig. 20H; MNHN.F.C02997 • 1 spec., RV, EF, Bergères-lès-Vertus ( Mont Aimé), sandy limestone with shell concentrations; MMM coll.; Fig. 20I; MNHN.F.C02998 • 1 spec., LV, IF; Vertus ( Grand Mont), sandy limestone; MMM coll.; Fig. 20J; MNHN.F.C02999 • 1 spec., RV, IF; same data as for preceding; MMM coll.; MNHN.F.C03000 • 1 spec., LV, EF; same data as for preceding; MMM coll.; MNHN.F.C03001 .</p><p>Measurements</p><p>MNHN.F.C02996, DVL: 6.8 mm, APL: 7.5 mm. MNHN.F.C02997, DVL: 5 mm, APL: 5.1 mm. MNHN.F.C02998, DVL: 6.2 mm, APL: 6 mm. MNHN.F.C02999, DVL: 7.5 mm, APL: 8.2 mm. MNHN.F.C03000, DVL: 7.2 mm, APL: 7 mm. MNHN.F.C03001, DVL: 7 mm, APL: 9 mm.</p><p>Description</p><p>Small and subcircular shell with lucinid shape. Postero-dorsal margin curved, posterior, ventral and dorsal margins rounded. Antero-dorsal margin slightly concave near the beak. Irregular commarginal growth ridges. No radial ribs. Hinge missing but apparently weak.</p><p>Remarks</p><p>These small bivalves display a lucinoid shape with more or less inflated valves, but it is not certain that they belong to the same species. The specimen MNHN.F.C02996 (Fig. 20G) has an anteriorly shifted, blunt umbo. In the specimen MNHN.F.C02997 (Fig. 20H), the umbo is central and rather pointed. The specimen MNHN.F.C02999 (Fig. 20J) is a more inflated shell than the others. Additional shells are therefore necessary to understand this complex of small bivalves.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E17FFFE1365F9D4FC27906E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E16FFFE1358FC78FBB097F4.text	F12B8A1C7E16FFFE1358FC78FBB097F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fimbria Megerle von Muhlfeld 1811	<div><p>Genus Fimbria Megerle von Mühlfeld, 1811</p><p>Type species</p><p>Fimbria magna Megerle von Mühlfeld, 1811 accepted as Fimbria fimbriata (Linnaeus, 1758) by monotypy. Present day, Indo-Pacific.</p><p>Stratigraphic range of the genus</p><p>Middle Jurassic (Bathonian) to present day, tropical seas (Cox et al. 1969).</p></div>	https://treatment.plazi.org/id/F12B8A1C7E16FFFE1358FC78FBB097F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E16FFE11356FB3BFD49949F.text	F12B8A1C7E16FFE11356FB3BFD49949F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fimbria multilamellosa (d'Orbigny 1850)	<div><p>Fimbria multilamellosa (d’Orbigny, 1850)</p><p>Fig. 20K</p><p>Corbis multilamellosa d’Orbigny, 1850: 131, n° 34.</p><p>Lucina contorta – C. d’Orbigny 1836: 284 (non Defrance, 1823).</p><p>? Corbis (Fimbria) multilamellosa – Combes 1908: 14.</p><p>Corbis multilamellosa – Chavan 1948: 566–567, 569. — Pacaud &amp; Merle 2002: 595, fig. 1.1a–b.</p><p>Fimbria multilamellosa – Pacaud 2018a: 252, fig. 198d–f; 2018b: 51. — Merle &amp; Pacaud 2018a: 176, fig. 159f. — Merle 2023: 25.</p><p>For a detailed synonymic list, see Pacaud (2018b: 51).</p><p>Type locality</p><p>Meudon, Hauts-de-Seine, France.</p><p>Type horizon</p><p>Pisolitic limestone of Meudon, Danian.</p><p>Type material</p><p>Lectotype FRANCE – Hauts-de-Seine • RV, EF; Meudon; Danian; A. d’Orbigny coll.; designated by Pacaud (2018a: 253, 256, fig. 198f); MNHN.F.R63237.</p><p>Paralectotypes (2 specs) FRANCE – Hauts-de-Seine • 1 spec., LV, EF; same data as for lectotype; A. d’Orbigny coll.; figured in Pacaud (2018a: 156, fig. 198d; MNHN.F.R63246. – Yvelines • 1 spec., LV, EF; Le Port-Marly; Danian; A. d’Orbigny coll.; MNHN.F.R63253.</p><p>Other material examined</p><p>FRANCE – Marne • 1 spec., RV, EF; Vertus; probably sandy limestone of Vertus; Ch. d’Orbigny coll., ex n°GG.gg2005/5961; figured in Merle &amp; Pacaud (2018a: 176, fig. 159e) and herein Fig. 20K; MNHN.F.C03002 .</p><p>Measurements</p><p>MNHN.F.C03002, DVL: 13 mm; APL: 18.5 mm.</p><p>Description</p><p>Original description (A. d’Orbigny 1850: 131): “Grande espèce ovale, voisine du C. pectunculus, mais beaucoup moins bombée, à côtes concentriques très rapprochées, à côtes rayonnantes à peine visibles”. [English translation: “Large oval species, similar to C. pectunculus, but much less rounded, with very closely spaced concentric ribs and barely visible radial ribs”.]</p><p>Description of the Maastrichtian material: small shell (juvenile), transversely elliptical, subequilateral. Posterior margin shorter than anterior margin. Around 40 low, lamellose commarginal ribs. Radial ribs slightly pronounced. Intersections between marginal and radial ribs forming small scales, more distinct on anterior margin than posterior margin.</p><p>Remarks</p><p>d’Orbigny (1850) described this species based on two syntypes from the Danian of the Paris Basin (Meudon and Le Port Marly) and one syntype from Vertus (MNHN.F.R63253). Pacaud (2018a: 253, 256, fig. 198f) designated the syntype from Meudon (MNHN.F.R63237) as the lectotype. According to Merle &amp; Pacaud (2018a), the Vertus paralectotype has more widely spaced commarginal ribs. It does not correspond to Fimbria multilamellosa, but to Fimbria sp. 1, a species discussed below. However, another specimen from Vertus (MNHN.F.C03002), found in the geology collection of the Muséum, bears close similarities to the Danian specimens. In particular, the tightly packed commarginal lamellae at the beginning of growth are completely identical. Because of these strong similarities, this specimen is thus identified herein as Fimbria multilamellosa . Combes (1908: 14) cited two species of Fimbria from the Bois d’Esmans, Fimbria multilamellosa and F. sublamellosa (d’Orbigny, 1850), but in the absence of material, these identifications cannot be confirmed. Squires &amp; Godaert (1994) described Linearia (Linearia) louellasaulae Squires &amp; Godaert, 1994, a Tellinidae from the Lower Eocene of the Crescent Black Hills Formation, Southwestern Washington (USA), whose sculpture is somewhat reminiscent of that of Fimbria multilamellosa . However, the radial ribs are much more pronounced and the American species is taller in shape.</p><p>Stratigraphic range</p><p>Maastrichtian to Danian, Paris Basin, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E16FFE11356FB3BFD49949F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E08FFE01383FEC4FBE29555.text	F12B8A1C7E08FFE01383FEC4FBE29555.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fimbria undefined-1	<div><p>Fimbria sp. 1</p><p>Fig. 20L–M</p><p>Fimbria sp. 1 – Merle &amp; Pacaud 2018a: 177, fig. 159f–g. — Merle 2023: 25.</p><p>Material examined (3 specs)</p><p>FRANCE – Marne • 1 spec., RV, EF, fragment; Bergères-lès-Vertus ( Mont Aimé); sandy limestone with shell concentrations; MMM coll.; figured in Merle &amp; Pacaud (2018a: 177, fig. 159f) and herein Fig. 20M; MNHN.F.A66779 • 1 spec.,?RV, EF; same data as for preceding; MMM coll.; figured in Merle &amp; Pacaud (2018a: 177, fig. 159g) and herein Fig. 20L; MNHN.F.A66780 • 1 spec., ex syntype of Corbis sublamellosa d’Orbigny, 1850; Vertus; A. d’Orbigny coll.; MNHN.F.R63253 .</p><p>Measurements</p><p>MNHN.F. A66780, DVL: 23.3 mm, APL (posterior area missing): 23 mm. Other material incomplete.</p><p>Description</p><p>Middle sized shell. Anterior and ventral margins rounded. Dorso-posterior margin almost straight. Around 34–35 commarginal lamellose ribs, slightly wrinkled in anterior region when crossing radial riblets. Commarginal ribs fairly spaced near the umbo, becoming closer together at end of growth. Commarginal sculpture crossed by numerous radial riblets.</p><p>Remarks</p><p>Fimbria sublamellosa is cited from Vertus (d’Orbigny 1850), Mont Aimé (Chavan 1948: 566) and Bois d’Esmans (Combes 1908: 14). d’Orbigny (1850) clearly stated that it differs from F. multilamellosa in having fewer commarginal ridges. The type material of F. sublamellosa included a Danian specimens from Le Port Marly and two specimens from Vertus. To stabilise the nomenclature of F. sublamellosa, Pacaud (2018a: 253, fig. 198g) designated the Danian specimen (MNHN.F.A46124) from Le Port Marly as the lectotype. The lamellose commarginal ribs of the Maastrichtian specimens are much more widely spaced throughout their growth than those of the typical Danian F. sublamellosa . These specimens from Vertus are named herein as Fimbria sp. 2 (see Fig. 20N). Like the lectotype of F. sublamellosa, Fimbria sp. 1 has spaced commarginal lamellae, but in the lectotype of F. sublamellosa they remain spaced at the end of growth, whereas in Fimbria sp. 1 they become closer together. The lectotype of F. sublamellosa has 22 ribs across the entire shell, while the two specimens of Fimbria sp. 1 have between 30 and 34 ribs. This difference in ontogeny suggests that Fimbria sp. 1 is a different species from F. sublamellosa . However, more specimens would be needed to better understand the sculptural variability of F. sublamellosa, Fimbria sp. 1 and Fimbria sp. 2 and clearly define these three species, which are distinguished by subtle differences. The citations of F. sublamellosa by Combes (1908) at Bois d’Esmans and by Chavan (1948) at Mont Aimé undoubtedly correspond to a species different from F. multilamellosa, but without figure or description, it is impossible to specify whether it is Fimbria sp. 1 or Fimbria sp. 2 .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E08FFE01383FEC4FBE29555	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E08FFE31383F954FEDF91C6.text	F12B8A1C7E08FFE31383F954FEDF91C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fimbria undefined-2	<div><p>Fimbria sp. 2</p><p>Fig. 20N</p><p>Fimbria sp. 2 – Merle &amp; Pacaud 2018a: 176, fig. 159j.</p><p>Material examined (2 specs)</p><p>FRANCE – Marne • 2 specs, ex syntypes of Corbis sublamellosa d’Orbigny, 1850, historical cast including 1 spec.?RV, EF; Vertus; A. d’Orbigny coll.; figured in Merle &amp; Pacaud (2018a: 176, fig. 159j) and herein Fig. 20N; MNHN.F.R63680 .</p><p>Measurements</p><p>MNHN.R63680 (figured specimen), DVL: 15 mm, APL: 24 mm. Other specimen incomplete.</p><p>Description</p><p>Middle sized shell, flattened. Anterior and ventral margins rounded. Around ten commarginal lamellose ribs. Commarginal ribs regularly spaced from umbo to ventral margin. Commarginal sculpture crossed by numerous radial riblets.</p><p>Remarks</p><p>As seen above Fimbria sp. 2 differs from Fimbria sp. 1 in that it has fewer marginal ribs (20 instead of 30–35 in Fimbria sp. 1) which are significantly more widely spaced. The poor state of preservation of the specimens calls for caution in generic attribution, but the numerous small radial riblets are typical of Fimbria .</p></div>	https://treatment.plazi.org/id/F12B8A1C7E08FFE31383F954FEDF91C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E0BFFE31320FC5CFF6E97F5.text	F12B8A1C7E0BFFE31320FC5CFF6E97F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chama Linnaeus 1758	<div><p>Genus Chama Linnaeus,1758</p><p>Type species</p><p>Chama lazarus Linnaeus, 1758 by subsequent designation Children (1823) and Opinion 484 (ICZN 1957). Present day, Indo-Pacific.</p><p>Stratigraphic range of the genus</p><p>Late Cretaceous (Coniacian) from Austria (Kennedy et al. 1970) to present day, subtropical and tropical seas.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E0BFFE31320FC5CFF6E97F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E0BFFE213C4FB3BFDBA97F4.text	F12B8A1C7E0BFFE213C4FB3BFDBA97F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chama loubryi Merle 2026	<div><p>Chama loubryi sp. nov.</p><p>urn:lsid:zoobank.org:act: 04AA310B-337C-4B12-90B9-399B562FD406</p><p>Fig. 20O–R</p><p>Chama sp. – Merle &amp; Pacaud 2018a: 176, fig. 159l–m. — Merle 2023: 25.</p><p>Etymology</p><p>Dedicated to Philippe Loubry for his important photographical work on the Maastrichtian shells of the Paris Basin.</p><p>Type material</p><p>Holotype FRANCE – Marne • LV, EF; Bergères-lès-Vertus; sandy limestone with shell concentrations; MMM coll.; MNHN.F.C03003 (Fig. 20O).</p><p>Paratypes (5 specs) FRANCE – Marne • 1 spec., LV, EF; same data as for holotype; MMM coll.; figured in Merle &amp; Pacaud (2018a: 176, fig. 159m) and herein Fig. 20R; MNHN.F.A66785 • 1 spec., LV, EF incomplete, historical cast; Vertus ( Faloise); Hébert coll.; figured in Merle &amp; Pacaud (2018a: 176, fig. 159l) and herein Fig. 20Q; UPMC.PAL.2017.0.19 • 1 spec., RV, EF; Vertus ( Grand Mont); sandy limestone; MMM coll.; Fig. 20P; MNHN.F.C03004 • 1 spec., LV, EF; same data as for preceding; MMM coll.; MNHN.F.C03006 • 1 spec. (LV, IF), Vertus ( Grand Mont); basal biocalcarenite with pectinids; MMM coll.; MNHN.F.C03007 .</p><p>Measurements</p><p>Holotype MNHN.F.C03003, DVL: 23 mm, APL: 18.8 mm. Paratype MNHN.F. A66785, DVL: 17.8 mm, APL: 18.2 mm. Paratype MNHN.F.C03004, DVL: 13.2 mm, APL: 12.2 mm. Paratype MNHN.F.C03006, DVL: 10.9 mm, APL: 9 mm.</p><p>Description</p><p>Middle sized shell, solid. Outline rounded to slightly elongated dorso-ventrally. Valves moderately to strongly convex and inequilateral. Umbo prosogyrate. Postero-dorsal, posterior and ventral margins rounded; antero-dorsal margin slightly concave or expanded by accretion of commarginal lamellae (see Fig. 20Q–R). Between 15–20 commarginal lamellae. Lamellae low, not spiny, tending to be tighter at end of growth.</p><p>Comparisons</p><p>Chama loubryi sp. nov. resembles Chama ciplyensis Vincent, 1930, from the Danian of the Paris Basin and Belgium, in its fairly smooth lamellar commarginal ribs, which are more widely spaced at the beginning than at the end of the growth. Chama loubryi differs from C. ciplyensis by lacking developed spiny lamellae at the end of its growth (see Vincent 1930: pl. 5, fig. 17a and Merle &amp; Pacaud 2018b: fig. 233c–d). It has less flat valves and is smaller in size (15 mm in C. loubryi versus 18–23 mm in larger specimens of C. cypliensis). Chama loubryi is smaller (DVL: 11 mm) Chama ancestralis Cossmann, 1908 from the Danian of the Mons Basin and doesn’t display a high hollow valve shaped like a cornucopia, as seen in C. ancestralis .</p><p>Stratigraphic range</p><p>Maastrichtian: Marne, France.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E0BFFE213C4FB3BFDBA97F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E0AFFE51388FAF4FC1492FA.text	F12B8A1C7E0AFFE51388FAF4FC1492FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bivalvia Linnaeus 1758	<div><p>Bivalvia indet.</p><p>Fig. 20S</p><p>Inceramidae indet. – Merle 2023: 25.</p><p>Material examined</p><p>FRANCE – Marne • 1 spec., LV, internal mould; Vertus ( Grand Mont); basal biocalcarenite with pectinids; MMM coll.; MNHN.F.C03005 .</p><p>Measurements</p><p>DVL: 13 mm; APL: 20 mm</p><p>Description</p><p>Shell small and subcircular. Antero-dorsal margin slightly concave. Anterior margin curved, slightly convex passing to rounded and narrower ventral margin. Posterior margin rounded and dorso-posterior margin almost straight. Six strong and thick commarginal folds.</p><p>Remarks</p><p>It is difficult to identify this internal mould, but its shape and sculpture with pronounced commarginal folds are reminiscent of inoceramid shells. Since inoceramids generally have folds rather than ribs, an internal mould should preserve this sculpture. The early stage of growth of the internal mould of Inoceramus aff. bererensis Sornay, 1975 from the Campanian of Austria illustrated by Tröger et al. (1999: pl. 1 fig. 1) displays similar strong commarginal folds.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E0AFFE51388FAF4FC1492FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
F12B8A1C7E0DFFE713F0FD41FA9C9182.text	F12B8A1C7E0DFFE713F0FD41FA9C9182.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nautilida Agassiz 1847	<div><p>Nautilida indet.</p><p>Fig. 21A–F</p><p>Nautilus hebertinus d’Orbigny, 1850: 127, no 34 pars, only the syntype of Montereau.</p><p>Material examined (2 specs)</p><p>FRANCE – Marne • 1 spec.; Bergères-lès-Vertus ( Mont Aimé); base of the section (1 m above the top of the Campanian chalk; Gilles Fronteau leg; Fig. 21A–D; MNHN.F.C03008. – Seine-et-Marne • 1 spec., fragments of an internal mould, ex syntype of Nautilus hebertinus d’Orbigny, 1850; Montereau ( probably Bois d’Esmans); A. d’Orbigny coll.; Fig. 21E–F; MNHN.F.B46110.</p><p>Measurements</p><p>MNHN.F.C03008, D: 83 mm, W: 67 mm, WH: 47 mm, DWH: 25 mm, UD: 4 mm.</p><p>Description</p><p>Small internal mould slightly deformed, probably juvenile specimen. Shape slightly depressed (W/WE = 1.42), involute (umbilicus generally about 5% of shell diameter). Umbilicus very small. Flanks slightly convex, converging towards rounded venter; venter rather wide. Peristome rounded, ventrally enlarged. Sinuous suture towards umbilicus.</p><p>Remarks</p><p>The nautilid from Bois d’Esmans is known only from a very fragmentary specimen (Fig. 21E–F) that was part of the type material for Nautilus hebertinus by Orbigny A., 1850, a species from the Danian of the Paris Basin (Montainville and Vigny). It was the occurrence of this species that prompted Chavan (1948) to recognise the Danian at Bois d’Esmans. The specimen collected at Bergères-lès-Vertus is better preserved, but does not allow for precise identification as it is a young specimen with poorly visible sutures (Stijn Goolaerts, written communication).</p><p>Composition of the assemblage</p><p>The list obtained from the detailed study of the mollusc assemblage at the Vertus (Grand Mont and Faloise), Bergères-lès-Vertus (Mont Aimé) and Esmans (Bois d’Esmans) sites includes 57 taxa (Table 1). It provides a much clearer picture of the assemblage than those presented in Merle &amp; Pacaud (2018a) and Merle (2023), which included only 24 and 41 taxa, respectively. For gastropods, 30 taxa were recognised. Sixteen species were identified, 14 of which are new (? Calliostoma monswavinarensis sp. nov., Sparnollonia bicarinata gen et sp. nov.,? Tectus acmantensis sp. nov.,? T. fraterculus sp. nov.,? T. senarmonti sp. nov.,? Clypeostoma vertusensis sp. nov., Calliomphalus (Planolateralus) montenati sp. nov., Rhinoclavis parvisulcatus sp. nov.,? Vicinocerithium heberti sp. nov., Granulolabium cretaceum sp. nov., Haustator bergierensis sp. nov., Sveltella binkhorsti sp. nov., Fusinus bouchardae sp. nov. and? Pseudopisania cretacea sp. nov.), eight are left in open nomenclature due to a lack of material, and five taxa are identified at the higher ranks. For bivalves, 26 taxa were recognised. Fifteen species were identified, including five new ones ( Nucula bauti sp. nov., Arca (s. str.) submontensis sp. nov., Plicatula pacaudi sp. nov., Bathytormus sparnacensis sp. nov. and Chama loubryi sp. nov.), five were left in open nomenclature and six taxa were identified at the family level or higher ranks. For cephalopods there is one unidentified Nautilida . The fauna thus appears to be quite diversified. The analysis of size distribution (Fig. 22A) shows that taxa measuring between 1 and 10 mm (the average size of the taxa is taken as a reference) represent only 27% of the assemblage for gastropods and 4% for bivalves. For the present day, small forms can represent more than 50% of the extant species (Bouchet et al. 2002) and they likely represent a high part of the post-Palaeozoic mollusc diversity (Cherns &amp; Wright 2011). These low values for the number of small species from the Maastrichtian of the Paris Basin, together with the sedimentary context (sandy limestone), clearly indicate a bias in the record of the palaeobiodiversity due to the lithification of the studied beds. The analysis the lithification bias in the Lutetian beds of Grignon (Sanders et al. 2015) reveals that nearly 80% of the species richness is not recorded in the lithified facies, and that loss can reach 100% for species smaller than 2 mm. Consequently, the fauna studied could originally have been much richer and this lithification bias cannot be compensated by simply adjusting the sample size. Given the limited amount of field research conducted so far, it is highly likely that future field trips will be able to supplement this study.</p></div>	https://treatment.plazi.org/id/F12B8A1C7E0DFFE713F0FD41FA9C9182	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Merle, Didier	Merle, Didier (2026): Biodiversity and palaeoecology of the Maastrichtian marine molluscs from the Paris Basin (France). European Journal of Taxonomy 1083: 1-108, DOI: 10.5852/ejt.2026.1083.3313, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3313/14563
