identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
317DC3B3F9855176B2C776D848B6B7FF.text	317DC3B3F9855176B2C776D848B6B7FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachythecium wangianum Min Li, X. T. Peng & Ignatov 2026	<div><p>Brachythecium wangianum Min Li, X. T. Peng &amp; Ignatov sp. nov.</p><p>Figs 1, 2</p><p>Diagnosis.</p><p>The species is most closely related to Brachythecium laetum phylogenetically, but can be distinguished morphologically from the latter by its longer and more slender leaf acumen, larger median laminal cells and an autoicous condition.</p><p>Type.</p><p>China • Hebei Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=117.55058&amp;materialsCitation.latitude=40.58806" title="Search Plazi for locations around (long 117.55058/lat 40.58806)">Wuling Mountain National Nature Reserve</a>, 40°35'17.01"N, 117°33'02.09"E, 907 m elev., 18 Sep. 2025, M. Li &amp; X. T. Peng L 20250918-66 (holotype: HBNU!) .</p><p>Description.</p><p>Plants medium-sized, 2–6 cm long, light green to yellowish green, slightly lustrous. Stems prostrate, with a central strand and a cortex of 3 (– 4) layers of incrassate cells, irregularly branched; branches robust, 5–15 (– 20) mm long, erect or slightly curved, terete when foliate. Proximal branch leaves triangular. Stem leaves erect when dry, loosely to densely inserted, (1.5 –) 2.0–3.0 × 0.7–1.0 (– 1.2) mm, ovate-lanceolate to broadly ovate-triangular, widest at 1 / 10–1 / 3 leaf length, gradually narrowed towards the apex, slightly constricted near the apex, acumen comprising 1 / 5–1 / 4 of total leaf length; leaves mostly concave, with one deep longitudinal plica on each side of costa extending to leaf margins; leaf margins plane or slightly recurved, denticulate; costa slender, extending to 1 / 2–2 / 3 leaf length, sometimes ending in a spine at apex; median laminal cells linear, (60 –) 70–100 (– 130) × 6–11 μm; leaf base rounded, not decurrent to slightly decurrent, margins plane or partially recurved, cells near base gradually shorter, slightly wider than median cells, 30–50 (– 65) × 8–15 (– 20) μm; alar cells thin-walled, subquadrate to oblong, 17–30 (– 50) × 15–20 (– 25) μm, arranged in 6–8 rows and together with the basal cells forming an indistinctly delimited, pellucid group. Branch leaves erect-spreading when dry, imbricate, lanceolate to ovate-lanceolate, (1.2 –) 1.5–2.6 × 0.4–0.75 (– 1.0) mm, concave, slightly plicate; margins serrulate; costa extending to 1 / 2–3 / 4 leaf length, usually ending in a spine; median laminal cells linear, smooth, 60–100 (– 130) × (6 –) 7–10 μm; leaf base not decurrent, basal cells rectangular to elliptical; alar cells quadrate to subquadrate, 15–30 (– 40) × 8–20 (– 25) μm.</p><p>Autoicous.</p><p>Perigonial leaves broadly ovate, with a single costa, apex acute or shortly acuminate; margins entire, bordered. Inner perichaetial leaves lanceolate, filiform-acuminate, with a single costa; outer perichaetial leaves shorter, ecostate. Seta reddish-brown, smooth, 0.5–1.5 cm long; capsule suberect to inclined, ovoid, curved and asymmetric; operculum long-conic; annulus differentiated, non-inflated, 1–2 rows of cells, fragmenting and deciduous; exostome teeth reddish brown, transversely striolate below, densely papillose above; endostome subequal to exostome in length, basal membrane 1 / 3–1 / 2 the height of exostome, cilia 1–3, distinctly nodose or protuberant; stomata phaneroporous, restricted to the capsule neck; spores ca. 14–20 μm in diameter, finely papillose.</p><p>Additional specimens examined</p><p>(paratypes). China • Jilin, Baishan, Changbai Mountain, 02 Sep. 2020, J. Kou 2020090233, 03 Sep. 2020, J. Kou 20200903049 (HBNU!) . China • Hebei, Chengde, Wuling Mountain National Nature Reserve, 18 Sep. 2025, M. Li &amp; X. T. Peng L 20250918-66 (HBNU!) ; • Shijiazhuang, Hebei Normal University, 05 Nov. 2024, M. Li &amp; Y. T. Wang 2024110502 (HBNU!) ; • Zhangjiakou, Tianlu Yanpian Mountain, 08 Aug. 2015, M. Li 2015080805 (HBNU!) . China • Inner Mongolia, Ulanqab, Erlongshitai National Forest Park, 03 Aug. 2014, M. Li 2014080301, 2014080343 (HBNU!) . China • Xinjiang, Ili, West Tianshan National Nature Reserve, 04 May 2024, M. Li &amp; M. Sulayman 2024050413 (HBNU!) .</p><p>Diagnostic features.</p><p>Leaves ovate-lanceolate to broadly ovate-triangular, distinctly concave; alar cells thin-walled, quadrate to subquadrate, not reaching the costa, together with the basal cells forming an indistinctly delimited, pellucid group; basal cells rectangular to elliptical, arranged in distinctly transverse rows; autoicous; seta smooth; annulus of 1–2 rows of non-inflated cells, fragmenting and deciduous.</p><p>Etymology.</p><p>The species epithet honorus Prof. Youfang Wang, an eminent bryologist who has long been dedicated to the taxonomic study of Brachytheciaceae in China. She compiled the Brachytheciaceae section in “ Bryophytes of China ” (Hu et al. 2008) and made important contributions to the research on this family.</p><p>Habitat and distribution.</p><p>Brachythecium wangianum grows on soil, humus or fallen logs in shaded habitats. Currently, it is known from northern China (Hebei, Inner Mongolia, Jilin, and Xinjiang) and in Baikal Lake area in southern Russia.</p><p>The species is currently known only from China and Russia. In China, it occurs disjunct in mountainous regions ranging from northeast China and north China to northwest China. In Russia, it grows on calcareous rocks within the mixed conifer and deciduous trees forests at low elevations. Brachythecium wangianum is absent from the core distribution ranges of B. laetum (e. g. eastern North America and western Europe), as well as from subtropical and tropical regions. Instead, its distribution is restricted to hemi-boreal to warm-temperate mountains of East and North Asia, producing a distinct pattern of geographical isolation and complementary distribution relative to B. laetum, which in Eurasia exhibits a strict western distribution.</p></div>	https://treatment.plazi.org/id/317DC3B3F9855176B2C776D848B6B7FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Peng, Xiao-Tong;Ignatov, Michael S.;Gu, Qian-Nan;Liang, Hong-Zhu;Chen, Cai-Xia;Wang, Yu-Ting;Li, Min	Peng, Xiao-Tong, Ignatov, Michael S., Gu, Qian-Nan, Liang, Hong-Zhu, Chen, Cai-Xia, Wang, Yu-Ting, Li, Min (2026): Brachythecium wangianum (Brachytheciaceae, Bryophyta), a new species from Northern China and Southern Russia. PhytoKeys 278: 191-204, DOI: 10.3897/phytokeys.278.205832
