taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529599/files/figure.png	https://doi.org/10.5281/zenodo.20529599	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529633/files/figure.png	https://doi.org/10.5281/zenodo.20529633	Fig. 14. Photographs of live Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0018. G–H. Habitat at the type locality.	Fig. 14. Photographs of live Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0018. G–H. Habitat at the type locality.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529651/files/figure.png	https://doi.org/10.5281/zenodo.20529651	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529653/files/figure.png	https://doi.org/10.5281/zenodo.20529653	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529639/files/figure.png	https://doi.org/10.5281/zenodo.20529639	Fig. 16. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0018 — SEM. A–H, K–Q. Male. I–J, R. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–M. Left leg 10, posterior views. N. Left coxal process on leg 10, posterior view. O. Femur of left leg 10, posterior view. P. Right first pleurite, ventral view. Q. Right midbody pleurite, ventral view. R. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go =gonopore; Mp=membranous process; O =operculum; po =postfemur; pre= prefemur; rdg=femoral ridge; sb =sensilla basiconica; Sp= small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	Fig. 16. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0018 — SEM. A–H, K–Q. Male. I–J, R. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–M. Left leg 10, posterior views. N. Left coxal process on leg 10, posterior view. O. Femur of left leg 10, posterior view. P. Right first pleurite, ventral view. Q. Right midbody pleurite, ventral view. R. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go =gonopore; Mp=membranous process; O =operculum; po =postfemur; pre= prefemur; rdg=femoral ridge; sb =sensilla basiconica; Sp= small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529641/files/figure.png	https://doi.org/10.5281/zenodo.20529641	Fig. 17. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left and right posterior telopod, posterior views, respectively. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih = inner horns; iL= inner lobes of telopod; ml = membranous lobe; scl-s= sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	Fig. 17. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left and right posterior telopod, posterior views, respectively. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih = inner horns; iL= inner lobes of telopod; ml = membranous lobe; scl-s= sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529605/files/figure.png	https://doi.org/10.5281/zenodo.20529605	Fig. 6. Photographs of live Zephronia macula Srisonchai & Wesener, 2024 and habitat. A–B. Specimens from Chanthaburi Province described by Srisonchai et al. (2024), “spotted” morph (paratypes CUMZzeph0016). C–F. Newly collected “green” morph specimens from Wat Khao Hin Sorn, Chachoengsao Province and Wat Tham Khao Chan, Sa Kaeo Province, respectively. G–H. Habitat at Wat Khao Hin Sorn.	Fig. 6. Photographs of live Zephronia macula Srisonchai & Wesener, 2024 and habitat. A–B. Specimens from Chanthaburi Province described by Srisonchai et al. (2024), “spotted” morph (paratypes CUMZzeph0016). C–F. Newly collected “green” morph specimens from Wat Khao Hin Sorn, Chachoengsao Province and Wat Tham Khao Chan, Sa Kaeo Province, respectively. G–H. Habitat at Wat Khao Hin Sorn.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529613/files/figure.png	https://doi.org/10.5281/zenodo.20529613	Fig. 8. Zephronia macula Srisonchai & Wesener, 2024, specimens from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior view. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac =apical cone; ad= antennal disc; bu =bursa; Cl=claw; cx= coxa; fe= femur; go = gonopore; O= operculum; po = postfemur; pre= prefemur; rdg= femoral ridge; sb =sensilla basiconica; Sp =small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	Fig. 8. Zephronia macula Srisonchai & Wesener, 2024, specimens from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior view. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac =apical cone; ad= antennal disc; bu =bursa; Cl=claw; cx= coxa; fe= femur; go = gonopore; O= operculum; po = postfemur; pre= prefemur; rdg= femoral ridge; sb =sensilla basiconica; Sp =small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529630/files/figure.png	https://doi.org/10.5281/zenodo.20529630	Fig. 13. Zephronia ovalis Gray, 1832, male specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM.A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; syn-cx =syncoxite. Scale bars in micrometres (μm).	Fig. 13. Zephronia ovalis Gray, 1832, male specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM.A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; syn-cx =syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529603/files/figure.png	https://doi.org/10.5281/zenodo.20529603	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE17FFA4FDFCFD98FEEB4EC5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529649/files/figure.png	https://doi.org/10.5281/zenodo.20529649	Fig. 21. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; sp =sclerotized spine; syn-cx =syncoxite. Scale bars in micrometres (μm).	Fig. 21. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; sp =sclerotized spine; syn-cx =syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE5BFE89FB104B43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE5BFE89FB104B43.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE77FD1CFB784D16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE77FD1CFB784D16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE77FD1CFB784D16.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529599/files/figure.png	https://doi.org/10.5281/zenodo.20529599	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE35FBF0FB384F29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FE35FBF0FB384F29.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529603/files/figure.png	https://doi.org/10.5281/zenodo.20529603	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FD96F9C7FBBC4EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FD96F9C7FBBC4EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE16FFA5FD96F9C7FBBC4EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529599/files/figure.png	https://doi.org/10.5281/zenodo.20529599	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE09FE89FEB04BEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE09FE89FEB04BEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE09FE89FEB04BEB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529603/files/figure.png	https://doi.org/10.5281/zenodo.20529603	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE44FC84FEAF4DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE44FC84FEAF4DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB9FE44FC84FEAF4DE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529603/files/figure.png	https://doi.org/10.5281/zenodo.20529603	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529605/files/figure.png	https://doi.org/10.5281/zenodo.20529605	Fig. 6. Photographs of live Zephronia macula Srisonchai & Wesener, 2024 and habitat. A–B. Specimens from Chanthaburi Province described by Srisonchai et al. (2024), “spotted” morph (paratypes CUMZzeph0016). C–F. Newly collected “green” morph specimens from Wat Khao Hin Sorn, Chachoengsao Province and Wat Tham Khao Chan, Sa Kaeo Province, respectively. G–H. Habitat at Wat Khao Hin Sorn.	Fig. 6. Photographs of live Zephronia macula Srisonchai & Wesener, 2024 and habitat. A–B. Specimens from Chanthaburi Province described by Srisonchai et al. (2024), “spotted” morph (paratypes CUMZzeph0016). C–F. Newly collected “green” morph specimens from Wat Khao Hin Sorn, Chachoengsao Province and Wat Tham Khao Chan, Sa Kaeo Province, respectively. G–H. Habitat at Wat Khao Hin Sorn.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529607/files/figure.png	https://doi.org/10.5281/zenodo.20529607	Fig. 7. Zephronia macula Srisonchai & Wesener, 2024, a male specimen from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A. Head, anterior view. B. Left ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal view. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it = 3-combed inner tooth; ant = antenna; cl= clypeus; co= condylus; cpd = central pad; cs = cuticular scales; ct= central tooth; et = external tooth of epipharynx; Et =external tooth of mandible; Ia= inner area of mandible; il = incisura lateralis; ip = inner palpi; ll = lamellae lingulales; me = mentum; mp = molar plate; o = ommatidia; pl = pectinate lamellae; rsp= row of spines; sc = sensory cone; st = stipites; to= organ of Tömösváry. Scale bars in micrometres (μm).	Fig. 7. Zephronia macula Srisonchai & Wesener, 2024, a male specimen from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A. Head, anterior view. B. Left ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal view. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it = 3-combed inner tooth; ant = antenna; cl= clypeus; co= condylus; cpd = central pad; cs = cuticular scales; ct= central tooth; et = external tooth of epipharynx; Et =external tooth of mandible; Ia= inner area of mandible; il = incisura lateralis; ip = inner palpi; ll = lamellae lingulales; me = mentum; mp = molar plate; o = ommatidia; pl = pectinate lamellae; rsp= row of spines; sc = sensory cone; st = stipites; to= organ of Tömösváry. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529613/files/figure.png	https://doi.org/10.5281/zenodo.20529613	Fig. 8. Zephronia macula Srisonchai & Wesener, 2024, specimens from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior view. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac =apical cone; ad= antennal disc; bu =bursa; Cl=claw; cx= coxa; fe= femur; go = gonopore; O= operculum; po = postfemur; pre= prefemur; rdg= femoral ridge; sb =sensilla basiconica; Sp =small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	Fig. 8. Zephronia macula Srisonchai & Wesener, 2024, specimens from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior view. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac =apical cone; ad= antennal disc; bu =bursa; Cl=claw; cx= coxa; fe= femur; go = gonopore; O= operculum; po = postfemur; pre= prefemur; rdg= femoral ridge; sb =sensilla basiconica; Sp =small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529615/files/figure.png	https://doi.org/10.5281/zenodo.20529615	Fig. 9. Zephronia macula Srisonchai & Wesener, 2024, a male specimen from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih =inner horns; iL= inner lobes of telopod; ml=membranous lobe; scl-s = sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	Fig. 9. Zephronia macula Srisonchai & Wesener, 2024, a male specimen from Wat Khao Hin Sorn, Chachoengsao Province, Thailand — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih =inner horns; iL= inner lobes of telopod; ml=membranous lobe; scl-s = sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529651/files/figure.png	https://doi.org/10.5281/zenodo.20529651	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE0AFFB7FE6FFA80FC614EFA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529653/files/figure.png	https://doi.org/10.5281/zenodo.20529653	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529619/files/figure.png	https://doi.org/10.5281/zenodo.20529619	Fig. 10. Photographs of live Zephronia ovalis Gray, 1832 and habitat. A–F. Specimens of a population from Sisaket Province, Thailand. G–H. Sandstone forest habitat.	Fig. 10. Photographs of live Zephronia ovalis Gray, 1832 and habitat. A–F. Specimens of a population from Sisaket Province, Thailand. G–H. Sandstone forest habitat.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529623/files/figure.png	https://doi.org/10.5281/zenodo.20529623	Fig. 11. Zephronia ovalis Gray, 1832, a male specimen from Huai Chan Waterfall, Sisaket Province, Thailand — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal views. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it= 3-combed inner tooth; ant= antenna; cl = clypeus; co = condylus; cpd = central pad; cs = cuticular scales; ct = central tooth; et =external tooth of epipharynx; Et =external tooth of mandible; hyp =hypopharynx; Ia =inner area of mandible; il =incisura lateralis; ip= inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o =ommatidia; pl =pectinate lamellae; rsp =row of spines; sc=sensory cone; st =stipites; to= organ of Tömösváry. Scale bars in micrometres (μm).	Fig. 11. Zephronia ovalis Gray, 1832, a male specimen from Huai Chan Waterfall, Sisaket Province, Thailand — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal views. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it= 3-combed inner tooth; ant= antenna; cl = clypeus; co = condylus; cpd = central pad; cs = cuticular scales; ct = central tooth; et =external tooth of epipharynx; Et =external tooth of mandible; hyp =hypopharynx; Ia =inner area of mandible; il =incisura lateralis; ip= inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o =ommatidia; pl =pectinate lamellae; rsp =row of spines; sc=sensory cone; st =stipites; to= organ of Tömösváry. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529627/files/figure.png	https://doi.org/10.5281/zenodo.20529627	Fig. 12. Zephronia ovalis Gray, 1832, specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go = gonopore; O = operculum; po = postfemur; pre = prefemur; rdg = femoral ridge; sb = sensilla basiconica; Sp = small triangular spines; st-pl = stigmatic plate; ta= tarsus; tib = tibia. Scale bars in micrometres (μm).	Fig. 12. Zephronia ovalis Gray, 1832, specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go = gonopore; O = operculum; po = postfemur; pre = prefemur; rdg = femoral ridge; sb = sensilla basiconica; Sp = small triangular spines; st-pl = stigmatic plate; ta= tarsus; tib = tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529630/files/figure.png	https://doi.org/10.5281/zenodo.20529630	Fig. 13. Zephronia ovalis Gray, 1832, male specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM.A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; syn-cx =syncoxite. Scale bars in micrometres (μm).	Fig. 13. Zephronia ovalis Gray, 1832, male specimens from Huai Chan Waterfall, Sisaket Province, Thailand — SEM.A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; syn-cx =syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529651/files/figure.png	https://doi.org/10.5281/zenodo.20529651	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE07FF8FFDFFFE89FCB74C0B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529653/files/figure.png	https://doi.org/10.5281/zenodo.20529653	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3CFF8FFE3DFAE4FB5B4E25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3CFF8FFE3DFAE4FB5B4E25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3CFF8FFE3DFAE4FB5B4E25.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529599/files/figure.png	https://doi.org/10.5281/zenodo.20529599	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3CFF8CFE7CF8C2FAD64B04.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529589/files/figure.png	https://doi.org/10.5281/zenodo.20529589	Fig. 1. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 1), inferred using maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. The blue letters represent species of the genus Zephronia, while the black letters indicate other genera.	Fig. 1. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 1), inferred using maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. The blue letters represent species of the genus Zephronia, while the black letters indicate other genera.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3CFF8CFE7CF8C2FAD64B04.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF8CFDC6FDE5FACD4D4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF8CFDC6FDE5FACD4D4D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529599/files/figure.png	https://doi.org/10.5281/zenodo.20529599	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	Fig. 4. Line drawings of male telopods and average body length of Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. (paratype), compared with the three closely related species. A–F. Zephronia chrysomallos Bhansali & Wesener, 2022. G–L. Zephronia erawani Bhansali & Wesener, 2022. M–R. Zephronia panhai Srisonchai, Sutcharit & Likhitrakarn, 2021. S–X. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior views. B, E, H, K, N, Q, T, W. Posterior views. C, I, O, U. Lateral views. Abbreviations: cr-t = crenulated teeth; ml= membranous lobe; sp= sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W =0.5 mm; F, L, R, X= 20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529633/files/figure.png	https://doi.org/10.5281/zenodo.20529633	Fig. 14. Photographs of live Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0018. G–H. Habitat at the type locality.	Fig. 14. Photographs of live Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0018. G–H. Habitat at the type locality.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529637/files/figure.png	https://doi.org/10.5281/zenodo.20529637	Fig. 15. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0018 — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it=3-combed inner tooth; ant= antenna; cl= clypeus; co= condylus; cpd = central pad; cs= cuticular scales; ct = central tooth; et= external tooth of epipharynx; Et = external tooth of mandible; Ia = inner area of mandible; il = incisura lateralis; ip = inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o=ommatidia; pl =pectinate lamellae; rsp =row of spines; sc =sensory cone; st=stipites; to=organ of Tömösváry. Scale bars in micrometres (μm).	Fig. 15. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0018 — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it=3-combed inner tooth; ant= antenna; cl= clypeus; co= condylus; cpd = central pad; cs= cuticular scales; ct = central tooth; et= external tooth of epipharynx; Et = external tooth of mandible; Ia = inner area of mandible; il = incisura lateralis; ip = inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o=ommatidia; pl =pectinate lamellae; rsp =row of spines; sc =sensory cone; st=stipites; to=organ of Tömösváry. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529639/files/figure.png	https://doi.org/10.5281/zenodo.20529639	Fig. 16. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0018 — SEM. A–H, K–Q. Male. I–J, R. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–M. Left leg 10, posterior views. N. Left coxal process on leg 10, posterior view. O. Femur of left leg 10, posterior view. P. Right first pleurite, ventral view. Q. Right midbody pleurite, ventral view. R. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go =gonopore; Mp=membranous process; O =operculum; po =postfemur; pre= prefemur; rdg=femoral ridge; sb =sensilla basiconica; Sp= small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	Fig. 16. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0018 — SEM. A–H, K–Q. Male. I–J, R. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–M. Left leg 10, posterior views. N. Left coxal process on leg 10, posterior view. O. Femur of left leg 10, posterior view. P. Right first pleurite, ventral view. Q. Right midbody pleurite, ventral view. R. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx =coxa; fe = femur; go =gonopore; Mp=membranous process; O =operculum; po =postfemur; pre= prefemur; rdg=femoral ridge; sb =sensilla basiconica; Sp= small triangular spines; st-pl =stigmatic plate; ta =tarsus; tib =tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529641/files/figure.png	https://doi.org/10.5281/zenodo.20529641	Fig. 17. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left and right posterior telopod, posterior views, respectively. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih = inner horns; iL= inner lobes of telopod; ml = membranous lobe; scl-s= sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	Fig. 17. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left and right posterior telopod, posterior views, respectively. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t= crenulated teeth; ih = inner horns; iL= inner lobes of telopod; ml = membranous lobe; scl-s= sclerotized spots; sp = sclerotized spine; syn-cx = syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529651/files/figure.png	https://doi.org/10.5281/zenodo.20529651	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE3FFF84FE22FB2AFDE348FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529653/files/figure.png	https://doi.org/10.5281/zenodo.20529653	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529591/files/figure.png	https://doi.org/10.5281/zenodo.20529591	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	Fig. 2. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 2), inferred from a maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. Background colours indicate species: purple =Z. macula Srisonchai & Wesener, 2024 “spotted” morph; green =Z. macula “green” morph; sky blue = Z. ovalis Gray, 1832; pink= Z. tratensis Srimongkol & Srisonchai sp. nov.; yellow= Z. sukhothaiensis Srimongkol & Srisonchai sp. nov. Scale bar represents branch length.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529603/files/figure.png	https://doi.org/10.5281/zenodo.20529603	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	Fig. 5. Line drawings of male telopods and average body length of Zephronia tratensis Srimongkol & Srisonchai sp. nov. (paratype) compared with the three closely related species. A–F. Zephronia enghoffi Srisonchai, Sutcharit & Likhitrakarn, 2021. G–L. Zephronia golovatchi Srisonchai, Sutcharit & Likhitrakarn, 2021. M–R. Zephronia lannaensis Likhitrakarn & Golovatch, 2021. S–X. Zephronia tratensis Srimongkol & Srisonchai sp. nov. A–C, G–I, M–O, S–U. Left anterior telopods. D–E, J–K, P–Q, V–W. Left posterior telopods. F, L, R, X. Comparison of average body length of each species. A, D, G, J, M, P, S, V. Anterior view. B, E, H, K, N, Q, T, W. Posterior view. C, I, O, U. Lateral view. Abbreviations: cr-t= crenulated teeth; ml = membranous lobe; sp =sclerotized spine. Scale bars: A–E, G–K, M–Q, S–W=0.5 mm; F, L, R, X =20 mm.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529643/files/figure.png	https://doi.org/10.5281/zenodo.20529643	Fig. 18. Photographs of live Zephronia tratensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0020. G–H. Habitat at the type locality.	Fig. 18. Photographs of live Zephronia tratensis Srimongkol & Srisonchai sp. nov. and habitat. A–F. Male paratypes MZKKU-MYR0020. G–H. Habitat at the type locality.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529645/files/figure.png	https://doi.org/10.5281/zenodo.20529645	Fig. 19. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it=3-combed inner tooth; ant= antenna; cl= clypeus; co= condylus; cpd = central pad; cs= cuticular scales; ct = central tooth; et= external tooth of epipharynx; Et = external tooth of mandible; Ia = inner area of mandible; il = incisura lateralis; ip = inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o=ommatidia; pl =pectinate lamellae; rsp =row of spines; sc =sensory cone; st=stipites; to=organ of Tömösváry. Scale bars in micrometres (μm).	Fig. 19. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A. Head, anterior view. B. Right ommatidia, lateral view. C. Left Tömösváry’s organ, lateral view. D. Epipharynx, ventral view. E–F. Gnathochilarium, ventral and frontal views, respectively. G. Left central pad, frontal view. H–I. Right lateral palpi, frontal views. J–L. Right mandible, dorsal, mesal and anterior views, respectively. Abbreviations: 3it=3-combed inner tooth; ant= antenna; cl= clypeus; co= condylus; cpd = central pad; cs= cuticular scales; ct = central tooth; et= external tooth of epipharynx; Et = external tooth of mandible; Ia = inner area of mandible; il = incisura lateralis; ip = inner palpi; ll =lamellae lingulales; me=mentum; mp =molar plate; o=ommatidia; pl =pectinate lamellae; rsp =row of spines; sc =sensory cone; st=stipites; to=organ of Tömösváry. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529647/files/figure.png	https://doi.org/10.5281/zenodo.20529647	Fig. 20. Zephronia tratensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0020 — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx = coxa; fe = femur; go = gonopore; O = operculum; po = postfemur; pre = prefemur; rdg = femoral ridge; sb = sensilla basiconica; Sp = small triangular spines; st-pl = stigmatic plate; ta= tarsus; tib = tibia. Scale bars in micrometres (μm).	Fig. 20. Zephronia tratensis Srimongkol & Srisonchai sp. nov., paratypes MZKKU-MYR0020 — SEM. A–H, K–P. Male. I–J, Q. Female. A–B. Antenna, anterior and mesal views. C–D. Antennal disc and apical cone. E–F. First coxae with stigmatic plates, posterior views. G–H. Coxae of legs 2 with gonopores, posterior views. I–J. Coxae of legs 2 with vulvae, posterior views. K–L. Left leg 10, posterior views. M. Left coxal process on leg 10, posterior view. N. Femur of left leg 10, posterior view. O. Right first pleurite, ventral view. P. Right midbody pleurite, ventral view. Q. Subanal plate, ventral view. Abbreviations: ac= apical cone; ad =antennal disc; bu = bursa; Cl = claw; cx = coxa; fe = femur; go = gonopore; O = operculum; po = postfemur; pre = prefemur; rdg = femoral ridge; sb = sensilla basiconica; Sp = small triangular spines; st-pl = stigmatic plate; ta= tarsus; tib = tibia. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529649/files/figure.png	https://doi.org/10.5281/zenodo.20529649	Fig. 21. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; sp =sclerotized spine; syn-cx =syncoxite. Scale bars in micrometres (μm).	Fig. 21. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU-MYR0020 — SEM. A–C. Telopods, anterior, posterior and ventral views, respectively. D, F. Right anterior telopod, ventral views. E. Left anterior telopod, anterior view. G–H. Sclerotized spots on right anterior telopod, ventral views. I–J. Crenulated teeth on right anterior telopod, ventral views. K, M. Crenulated teeth on left posterior telopod, posterior views. L. Sclerotized spots on right posterior telopod, ventral view. N. Membranous lobe on right posterior telopod, anterior view. Abbreviations: cr-t=crenulated teeth; ih= inner horns; iL= inner lobes of telopod; ml= membranous lobe; scl-s = sclerotized spots; sp =sclerotized spine; syn-cx =syncoxite. Scale bars in micrometres (μm).	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529651/files/figure.png	https://doi.org/10.5281/zenodo.20529651	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	Fig. 22. Endoterga of thoracic shield and tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype MZKKU- MYR0020. Abbreviations: cp =cuticular impression; ia = inner area; ma =middle area; oa= outer area; pm = posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE37FF9DFE49FD9AFEB84DDC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529653/files/figure.png	https://doi.org/10.5281/zenodo.20529653	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	Fig. 23. Endoterga of tergite 7, all in ventral view — SEM. A–B. Zephronia macula Srisonchai & Wesener, 2024 (Wat Khao Hin Sorn). C–D. Zephronia ovalis Gray, 1832 (Huai Chan Waterfall, Thailand). E–F. Zephronia sukhothaiensis Srimongkol & Srisonchai sp. nov., male paratype CUMZzeph0018. G–H. Zephronia tratensis Srimongkol & Srisonchai sp. nov., male paratype CUMZ-zeph0020. Abbreviations: cp = cuticular impression; ia= inner area; ma= middle area; oa = outer area; pm= posterior margin.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE2EFF9DFE75FABCFB594FFD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529589/files/figure.png	https://doi.org/10.5281/zenodo.20529589	Fig. 1. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 1), inferred using maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. The blue letters represent species of the genus Zephronia, while the black letters indicate other genera.	Fig. 1. COI-based barcoding tree of the genus Zephronia Gray, 1832 (part 1), inferred using maximum likelihood (ML) analysis. Numbers at the nodes indicate bootstrap support (BS) from ML and posterior probabilities (PP) from Bayesian inference (BI), shown as ML/BI. A hyphen “-” at nodes indicates a different topology between the ML and BI analyses. The scale bar represents branch length. The blue letters represent species of the genus Zephronia, while the black letters indicate other genera.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
F24EBD4FCE2EFF9DFE75FABCFB594FFD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20529595/files/figure.png	https://doi.org/10.5281/zenodo.20529595	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	Fig. 3. Distribution map of Zephronia Gray, 1832 in Thailand and nearby areas. Identical shapes of both black and white symbols represent same species.	2026-05-13	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon		Zenodo	biologists	Srimongkol, Nirun;Ruangchai, Sukhum;Tungpairojwong, Nisarat;Srisonchai, Ruttapon			
