identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E80787D7FFC31675FF723A99FDF1FF26.text	E80787D7FFC31675FF723A99FDF1FF26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara axillaris (Paykull 1799)	<div><p>Mycetochara (s. str.) axillaris (Paykull, 1799)</p><p>(Fig. 1)</p><p>Material. About 40 specimens were examined from different regions of European Russia: Kaliningrad, Yaroslavl, Moscow, Tula, Orenburg and Rostov regions (ZIN, ZMMU, ZMSFU, PCVA) .</p><p>Distribution. Europe from Scandinavia to Mediterranean region (Novák 2014, 2020b). Widely distributed in the European part of Russia north to Yaroslavl Region (ZIN, ZMMU; Dubrovina et al. 1979), south to central areas of Rostov Region (Volgodonsk, ZMSFU), west to Kaliningrad Region (Alekseev &amp; Bukejs 2014), east to Tumen Region in West Siberia (Sergeeva &amp; Stolbov 2020). The species was also recorded for Mordovia (Egorov et al. 2018), Chuvashia (Egorov 2016), Moscow (Nikitsky 2016, 2019), Tambov (Volodchenko 2020), Tula (Dorofeev 2009), Kaluga (S. Alekseev et al. 2020), Saratov (Volodchenko &amp; Sazhnev 2023), Lipetsk (Tsurikov 2009) and Voronezh (Negrobov et al. 2005) regions.</p></div>	https://treatment.plazi.org/id/E80787D7FFC31675FF723A99FDF1FF26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC31676FF7239D5FB43F98E.text	E80787D7FFC31676FF7239D5FB43F98E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara Guerin-Meneville 1827	<div><p>Subgenus Mycetochara Guérin-Méneville, 1827</p><p>Type species: type species Cistela scapularis Illiger, 1805 (= Cistela humeralis Fabricius, 1787).</p></div>	https://treatment.plazi.org/id/E80787D7FFC31676FF7239D5FB43F98E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC61673FF723EEAFC71FCEB.text	E80787D7FFC61673FF723EEAFC71FCEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Ernocharis) C. G. Thomson 1859	<div><p>Subgenus Ernocharis C.G. Thomson, 1859</p><p>Type species: Cistela brevis Illiger, 1794 (= Cistela maura Fabricius, 1792)</p></div>	https://treatment.plazi.org/id/E80787D7FFC61673FF723EEAFC71FCEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC61673FF723FBEFA16FA21.text	E80787D7FFC61673FF723FBEFA16FA21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) angustifrons Reitter 1899	<div><p>Mycetochara (Ernocharis) angustifrons Reitter, 1899</p><p>(Fig. 3)</p><p>= Mycetochara (Ernocharis) seidlitzi Reitter, 1899, synonymized by Iablokoff-Khnzorian (1983: 129)</p><p>Material. 1♀ (ZIN), Russia, Krasnodar Region, Kamyshanova glade, 1200 m, Fagus and Abies, 21.vi.1982 (B.A. Korotyaev); 1♀ (ZMSFU), Russia, Krasnodar Region, Mzymta valley, Engelmanova glade, 28.vii.1997 (M.V. Nabozhenko); 1♀ (ZMSFU), Russia, Karachay-Cherkessia, Arkasara mountain range, 3.viii.1996 (D.G. Kasatkin); 2♂♂, 5♀♀ (ZIN, PCMN), Russia, Karachay-Cherkessiya, Zakan, 15– 19.06.2025 (M.V. &amp; S.V. Nabozhenko). 1♂, 4♀♀ (PCMN), Georgia, Abkhazia, Amzhular natural boundary, 18–19.vi.2019, 1600–2000 m (A. Zubov).</p><p>Distribution. The species is distributed in the Western Caucasus. It is known from Russia (Krasnodar Region, Karachay-Cherkess Republic, Adygea), Georgia: Svanetia (Iablokoff-Khnzorian 1983, Nikitsky et al. 2008, Nabozhenko et al. 2010; Novák 2020a; ZIN, ZMMU, PCMN) and Abkhazia (new record).</p><p>Notes. The name Mycetochara seidlitzi Reitter, 1899 was correctly synonymized with M. angustifrons Reitter, 1899 (Iablokoff-Khnzorian 1983), but this synonymy was missed in the catalogue (Novák 2020b). Mycetochara seidlitzi was described based on two females from Uch-Dere (now in Krasnodar Region, Russia) (Reitter 1899), the same type locality as for M. angustifrons (was described by one male) (Novák 2020a). The female syntype from HNHM (photograph was kindly provided by Vladimír Novák) is conspecific to those of M. angustifrons females.</p></div>	https://treatment.plazi.org/id/E80787D7FFC61673FF723FBEFA16FA21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC61673FF723CACFC7BFD5F.text	E80787D7FFC61673FF723CACFC7BFD5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara flavipes (Fabricius 1792)	<div><p>Mycetochara (s. str.) flavipes (Fabricius, 1792)</p><p>(Fig. 2)</p><p>Near 70 specimens were examined from many regions of European Russia (ZIN, ZMMU, ZMSFU, PCVA) .</p><p>Distribution. Europe from Scandinavia to Mediterranean region, European Russia, Siberia, Russian Far East, Japan, Mongolia (Novák 2020b), Kazakhstan (Nakládal et al. 2017). Widely distributed in the European part of Russia north to Southern Karelia (Polevoi et al. 2018), south to the Northern Cis-Azov region (Rostov Region: Rostov-on-Don, Novocherassk, ZMSFU), west to Kaliningrad Region (V. Alekseev et al. 2015), east to Sakhalin and Japan (Novák 2020b). Most sources for M. axillaris also list M. flavipes and a very long list of references for different regions of Russia, which are not given here.</p><p>Iablokoff-Khnzorian (1983) recorded this species also for the Caucasus, later Novák (2020b) listed M. flavipes for Georgia, but the presence of this species here is not confirmed. Nabozhenko et al. (2010) recorded M. adygea sp. nov. from Nikel village (Adygea Republic) under the name M. flavipes .</p></div>	https://treatment.plazi.org/id/E80787D7FFC61673FF723CACFC7BFD5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC61670FF723A6BFE9DFE56.text	E80787D7FFC61670FF723A6BFE9DFE56.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) excelsa Reitter 1884	<div><p>Mycetochara (Ernocharis) excelsa Reitter, 1884</p><p>(Fig. 4)</p><p>Material. 2♂, 6♀♀ (PCMN, ZIN, ZMSFU), Russia, Dagestan, Magaramkent Distr., lower reaches of Samur River, “Samur forest” National park, 12–14.v.2008 (M. V. and S.V. Nabozhenko, G.V. Chesnokov) .</p><p>Distribution. This species inhabits relict Hyrcanian forests fragmentarily represented in the Eastern Caspian region (Russia, Southern Dagestan; Azervaijan: Ismailli State Reserve) and Southern Caspian region (Azerbaijan, Iran: Talysh and North Alborz). Novák (2020a) also recorded this species for Georgia, which requires verification, because Hyrcanian forests are absent in Georgia. The record for the Northwestern Caucasus (Adygea: Azishtau) in Nabozhenko et al. (2010) is erroneous and belongs to the immature specimen of M. zolotareffi .</p><p>Note. Novák (2022) excluded M. excelsa from the Russian faunistic list (assuming that it was confused with the species M. strejceki). However, the south of Russia is an extremely diverse territory and on the Russian Caucasus both subtropical Colchian forests in the western part and fragments of relict Hyrcanian forests in the eastern part are present. Mycetochara strejceki occurs in western Colchian forests, while M. excelsa is one of the numerous and most common species in the Hyrcanian forests of the south of Dagestan (Russia).</p><p>Bionomics. Adults were collected at night on trunks of Quercus pedunculiflora K. Koch in Samur forest, Dagestan .</p></div>	https://treatment.plazi.org/id/E80787D7FFC61670FF723A6BFE9DFE56	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC51670FF723DE1FB22FB86.text	E80787D7FFC51670FF723DE1FB22FB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) gracilis (Faldermann 1837)	<div><p>Mycetochara (Ernocharis) gracilis (Faldermann, 1837)</p><p>(Fig. 5)</p><p>Material. 1♂ (ZMMU), Russia, Krasnodar Region, Ubinskaya, 18.v.1975 (V. Belov) ; 1♀ (ZMMU), Russia, Stavropol Region, Pyatigorsk, 24.v.1911 (B. Il’in) ; 1♀ (ZMMU), Russia, Stavropol Region, Zheleznovodsk, 1913 (collection of Artobolevsky); 1♂ (ZMSFU), same locality, 5.vi.1949 (A.P. Runich); 1♀ (ZMSFU), same locality, forest, 26.vi.1949 (A.P. Runich); 1♀ (ZMSFU), Russia, Rostov Region, Rostov-on-Don, 7.vi.1991 (E.A. Khachikov) .</p><p>Notes. All specimens from ZMMU are additionally labeled as “ Holotypus [Paratypus] Mycetochara (Ernocharis) ciscaucasica Dubr. ”, but this “taxon” was not described. We compared Armenian specimens of M. gracilis, collected and examined by Iablokoff-Khnzorian (1983) (PCMK) and specimens from the North Caucasus and didn’t find significant differences.</p><p>Distribution. The species was described from “Transcaucasia” and occurs in Armenia, Georgia and the Caucasian Black Sea coast (Iablokoff-Khnzorian 1983). In Russia, M. gracilis is a very rare species, which is known only by several specimens from the Caucasian Mineral Waters area and Krasnodar Region. One female was collected in Cis-Azov Region (Rostov-on-Don). This specimen can be a result of unintentional invasion of this species from the resorts “Caucasian Mineral Waters” with planting material for landscaping Rostov-on-Don.</p><p>Bionomics. The species was collected on maples in Armenia (Iablokoff-Khnzorian 1983).</p></div>	https://treatment.plazi.org/id/E80787D7FFC51670FF723DE1FB22FB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC51670FF723891FC44F9F2.text	E80787D7FFC51670FF723891FC44F9F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) humeralis (Fabricius 1787)	<div><p>Mycetochara (Ernocharis) humeralis (Fabricius, 1787)</p><p>(Fig. 6)</p><p>Material. We examined 1♂ and 1♀ specimens, listed by Alekseev &amp; Bukejs (2017) from Kaliningrad Region and 2♂♂ and 1♀ from Lvov Region, Yavor District in Western Ukraine (ZMMU), listed by Dubrovina et al. (1979).</p><p>1♀ (PCVA), Russia, Kaliningrad Region, Slavsky Distr., Vysokoe, 20.vi.2025, dry side of roadside linden tree (V.I. Alekseev) ; 1♀ (ZMMU), Ukraine, Khmelnitsy Region, Kamenets-Podolsky, 21.v.1908 ( V. and I. Yakubovsky) .</p><p>Distribution. This nemoral species is widely distributed in Europe (Novák 2020b). In Russia it was collected only from Kaliningrad (Alekseev &amp; Bukejs 2017) and Voronezh (https://www.zin.ru/animalia/coleoptera/rus/ mychumkm.htm) regions.</p><p>Bionomics. Oak forests, on damaged trees (Alekseev &amp; Bukejs 2017).</p></div>	https://treatment.plazi.org/id/E80787D7FFC51670FF723891FC44F9F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC5167CFF723A45FE5CFD5A.text	E80787D7FFC5167CFF723A45FE5CFD5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) ingushetica Nabozhenko & Gadaborsheva 2024	<div><p>Mycetochara (Ernocharis) ingushetica Nabozhenko &amp; Gadaborsheva, 2024</p><p>(Fig. 7)</p><p>Type material. See Nabozhenko &amp; Gadaborsheva (2024).</p><p>Material. 9♂♂, 1♀ (ZMMU), Russia, North Ossetia, Buron, iv.1976 (M.I. Dubrovina) ; 1♀ (PCMN), Russia, Karachay-Cherkessia, upper reaches of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=5.5&amp;materialsCitation.latitude=30.0" title="Search Plazi for locations around (long 5.5/lat 30.0)">Rechepsta River</a>, 43˚34ʹ 30.0ʺN, 41˚05ʹ 05.5ʺE, 2034 m, on Acer trautvetteri Medw. 3– 4.07.2020 (M.V. Nabozhenko, D.G. Kasatkin) ; 2♀♀ (PCMN), Russia, Karachay-Cherkessia, turn to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.9&amp;materialsCitation.latitude=19.7" title="Search Plazi for locations around (long 3.9/lat 19.7)">Nizhniy Arkhyz</a>, 43˚40ʹ 19.7ʺN, 41˚26ʹ 03.9ʺE, 1222 m, 6.07.2020 (M.V. Nabozhenko) ; 2♀♀ (ZIN), Karachay-Cherkessia, Zakan, 43˚41ʹ56ʺN / 40˚47ʹ35ʺE, 1250–1300 m, under the bark of a rotten Acer platanoides L., 1753, 15–19.vi.2025 (M.V. Nabozhenko) .</p><p>Notes. This species was listed as M. hirsuta Pic, 1925 by Dubrovina et al. (1979) from North Ossetia; ten specimens were hatched from larvae, collected in rotten maple. Later, M. hirsuta was interpreted as a junior synonym of M. maura (Novák 2020b) . We believe that both opinions are erroneous. Mycetochara hirsuta was described from “Borjom” (Transcaucasia, Georgia: Borjomi) as a large species with suberected body setation (Pic 1925a). The species M. maura does not inhabit the Caucasus, where it is replaced by a similar species M. zolotareffi . Thus, the Transcaucasian M. hirsuta is a valid species that should be reestablished from the junior synonyms of M. maura: M. hirsuta Pic, 1925, sp. resurr. Thus, two large black unicolorous species remain in the Caucasus: M. ingushetica and M. hirsuta . The first one is known only from the Northern Caucasus, and the second one from Transcaucasia. The second species, collected from Abkhazia, was tentatively identified as M. cf. abschasica and well illustrated by Nabozhenko (2022) and Nabozhenko &amp; Gadaborsheva (2024). This is a large specimen belonging to M. hirsuta .</p><p>Distribution. Russia, the North Caucasus: Karachay-Cherkessia, North Ossetia, Ingushetia.</p><p>Bionomics. The species is distributed from 600 to ~ 2000 m. Most specimens (larvae, pupae and imagines) were found on different Acer spp. In Ingushetia all specimens were collected on window trap with exposure on damaged Quercus robur L.</p></div>	https://treatment.plazi.org/id/E80787D7FFC5167CFF723A45FE5CFD5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC9167CFF723EEDFBB5FB7E.text	E80787D7FFC9167CFF723EEDFBB5FB7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) maura (Fabricius 1792)	<div><p>Mycetochara (Ernocharis) maura (Fabricius, 1792)</p><p>(Fig. 8)</p><p>Material. We examined 4♂♂ and 7♀♀ from Kaliningrad Region of Russia, listed by V. Alekseev et al. (2015) and Alekseev (2022).</p><p>1♀ (PCVA), Russia, Kaliningrad Region, Mechnikovo ( Baltiysk), 31.05.2025, dried oak (V.I. Alekseev) ; 1♀ (ZMMU), Ukraine, “Lesienice” (now, Lvov Region, Lisinichi); 1♂ (ZMMU), Germany, Berlin (ex coll. of Muche) .</p><p>Distribution. The species is widely distributed in Europe (Novák 2020b). In Russia, it is known only from Kaliningrad Region (V. Alekseev et al. 2015, V. Alekseev 2022; in the second reference as M. linearis). The record for Krasnodar Region (Ubinskaya) (Nikitsky et al. 2008) belongs possibly to M. zolotareffi .</p><p>Bionomics. This species was collected in Kaliningrad Region in different broadleaf damaged trees: Acer platanoides, Tilia cordata, Populus tremula (V. Alekseev et al. 2015), Quercus sp.</p></div>	https://treatment.plazi.org/id/E80787D7FFC9167CFF723EEDFBB5FB7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC9167CFF7238C9FCD7F962.text	E80787D7FFC9167CFF7238C9FCD7F962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) obscura (Zetterstedt 1840)	<div><p>Mycetochara (Ernocharis) obscura (Zetterstedt, 1840)</p><p>(Fig. 9)</p><p>Material. We examined all specimens (only females), listed by Nikitsky (2016, 2019) for Moscow Region and one female, recorded by Egorov (2019) and Egorov &amp; Semionenkov (2023) for Chuvash Republic.</p><p>1♂, 1♀ (PCMN), Russia, Tver Region, Kimry Distr., Shelkovnikovo village environs, 25–28.vi.2022 (collector unknown) .</p><p>Distribution. North and Central Europe (Novák 2020b). Russia: Komi Republic (Yurkina 2011), Udmurtia (Dedukhin &amp; Nikitsky 2009), Chuvashia (Egorov &amp; Semionenkov 2023), Moscow (Nikitsky 2016, 2019) and Tver regions; it was listed for Kaliningrad Region (Bercio &amp; Folwaczny 1979); Western Siberia: Tyumen Region (Sergeeva &amp; Stolbov 2020). The record for the steppe Ciz-Ural region (Kozminykh 2015) needs verification.</p><p>Bionomics. Rare boreal species, usually on rotten old Picea trees at forest edges, sometimes in Pinus -Betula forests (Čížek &amp; Hauck 2006, Egorov &amp; Semionenkov 2023).</p></div>	https://treatment.plazi.org/id/E80787D7FFC9167CFF7238C9FCD7F962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFC91678FF723B35FC11FE2A.text	E80787D7FFC91678FF723B35FC11FE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) ruthenica Nabozhenko & Martynov & Bulysheva 2025	<div><p>Mycetochara (Ernocharis) ruthenica Nabozhenko, sp. nov.</p><p>(Fig. 10)</p><p>Material. Holotype, ♂ (ZMSFU): Russia, Rostov Region, Myasnikovsky District, environs of educational and experimental farm “Nedvigovka” of Southern Federal University, 47.272952˚N, 39.318149˚E, 29.iv.2024 (E.A. Khachikov); paratype, ♂ (ZMSFU), the same locality and collector, but 12.v.2025.</p><p>Description. Holotype (Fig. 10A). Body slender, narrow, unicolour, dark-brown, legs ochre-reddish, antennae with brown 5–8 antennomeres and ochre-reddish rest ones. Body covered with dense suberected brown setae, yellowish on head and pronotum.</p><p>Head.Anterior margin of epistoma straight, lateral margin of genae slightly evenly rounded, strongly converging from eyes to anterior margin of epistome. Dorsal side with sparse smooth (edges are not distinct) puncturation (interpuncture distance near twice as long as puncture diameter), each puncture bears yellowish long suberected setae. Head ventrally and laterally with long smooth wrinkles along posterior edge of eyes, with fine and sparse puncturation. OI = 41.6. Antennae long, reaching basal third of elytral length when directed backward.</p><p>Prothorax. Pronotum (Fig. 10B) transverse (1.34 times as wide as long), widest at anterior third, 1.25 times as wide as head; PI = 74.38. Lateral margins slightly rounded, converging from widest portion to base. Anterior margin straight, base slightly bisinuate. Disc slightly convex only laterally, slightly depressed in basal half of middle and stronger depressed at basal portions on sides from middle. Puncturation of disc coarse and sparse, punctures round, interpuncture distance 2–3 times as long as puncture diameter; each puncture bears long subrecumbent yellowish seta. Anterior angles not expressed, pronotum widely rounded antero-laterally; posterior angles slightly obtuse. Edges of pronotum not margined. Prothoracic hypomera and prosternum smooth and shiny, without setation.</p><p>Pterothorax. Elytra elongate, subparallel when completely closed, 1.28 times wider at base (immediately behind humeral angles) than base of pronotum. Humeral angles widely rounded. Striae consist of round slightly raduliform dense punctures, interstriae 1 and 2 slightly convex, rest ones flattened, each interstria contains one irregular row of raduliform punctures; each puncture bears long subrecumbent light-brown setae; elytral setation dense. Mesoventrite and metepimera with dense and coarse puncturation; mesepisterna and mesepimera with sparse irregular puncturation; metaventrite convex, sparsely punctured at anterior portion and smooth at basal half.</p><p>Legs slender, long, reddish-ochre; profemora shortest, metafemora longest; tibiae straight; each protarsal claw with 7 teeth on inner side.</p><p>Abdomen. Abdominal ventrites punctured by sparse and fine raduliform punctures and covered with recumbent setae, shorter than on dorsal surface. Aedeagus (Fig. 10D, E) with basal piece 2.85 times longer than parameres. Parameres dorsally with slightly rounded lateral margins converging to apex and narrowly rounded tip, slightly Cshaped laterally, with directed backward strong spines on sides. Penis with cone-like sculpture apically.</p><p>Variability. The second male (paratype) differs from the holotype in the darker body with brown antennomeres 4–10 (both 4 and 10 antennomeres are bicolour, with reddish base and brown rest surface), shape of the pronotum with lateral margins straight in the basal half and right posterior angles (Fig. 10C).</p><p>Etymology. The name translated from Latin as pertaining to Rus. Feminine adjective.</p><p>Comparative diagnosis. The species is externally and in small size very similar to the coastal and foothill specimens of M. zolotareffi, but differs from the latter by larger eyes (OI = 40.81–41.6 vs 49.4–51.7 in M. zolotareffi). From both similar taxa, M. zolotareffi and M. maura, the new species differs in the structure of the aedeagus (compare Figs 8C, D, E, 10D, E, 13C, D and 15F, G) and penis with cone-like sculpture. The species differs from the similar M. ingushetica in the shape of the pronotum (lateral margins of the pronotum are not emarginated vs emarginated and pronotal shape trapezoidal in M. ingushetica); aedeagus is also different in M. ingushetica (compare Figs 7C, D and 10D, E)</p><p>Bionomics. Both specimens flew to specially moistened dry firewood (information from the collector, E.A. Khachikov). The type locality contains several old oak trees among steppe landscape and a small planted oak grove of middle-aged trees. The species probably lives on these oak trees.</p></div>	https://treatment.plazi.org/id/E80787D7FFC91678FF723B35FC11FE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFCD1678FF723E7DFEBCF86C.text	E80787D7FFCD1678FF723E7DFEBCF86C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) abschasica Pic 1925	<div><p>Mycetochara (Ernocharis) abschasica Pic, 1925</p><p>(Figs 11, 12)</p><p>? = Mycetochara (Ernocharis) strejceki Novák, 2022</p><p>Type material. Holotype of M. abschasica, 1♀ (MNHN) (Fig. 12), examined by photo images courtesy Antoine Mantilleri, credit for images is “MNHN/Maxime Boutin”: “ Abchasien Kavkas.” (print), “type” (handwritten by Pic), “ abschasica n. sp. ” (handwritten by Pic), “TYPE” (red, print), “ Museum Paris Coll. M. Pic ”, “ HOLOTYPE ” (red, print), “ Holotype Mycetochara abschasica Pic, 1925 ”, “MNHM EC 50607 [+ QR-code]”.</p><p>Material. 2♂♂ (ZMSFU), Krasnodar Region, Khosta, “Yew-Boxwood grove”, window traps in Taxus - Fagus forest, exposition on Fagus, 5.v.–6.vi.2006 (A.R. Bibin); 1♀ (ZMSFU), same locality, soil traps in Taxus forest, v.2006 (A.R. Bibin, Yu.A. Chumachenko).</p><p>Notes. The female of this species has red-brown or bicolour body (brown head and pronotum and red-brown elytra). The body shape is similar to those in M. zolotareffi, but the pronotum is narrower (pronotum 1.24–1.25 times as wide as long in M. abschasica vs 1.27–1.34 in M. zolotareffi). Very rarely, immature females of M. zolotareffi also have bicolour body with ochre elytra and brown pronotum and head (at least we have one such female from Goryachy Klyuch, Russia in large series of unicolour beetles, one male and one female from Gudauta (Abkhazia) within series of unicolour specimens). In this case the most distinct distinguish character for females of both species is the proportion of the pronotum, and for males is the structure of the aedeagus.</p><p>Pic (1925b) described Mycetochara abschasica based on one female from Abkhasia.He compared M. abschasica with M. zolotareffi (misspelled as “ M. zolotarewi ”) and mentioned as diagnostic characters different body colouration and more elongate elytra. The holotype female of M. abschasica is conspecific to the female from Yew-Boxwood grove, and two males from the same locality correspond to M. strejceki . Males from Khosta externally slightly differ from those, imaged in Novák (2022) by the shape of the pronotum: widest at base (Novák 2022, fig. 28) or lateral margins parallel at basal half (Novák 2022, fig. 29). We examined one male from Khosta with subparallel lateral margins of the pronotum, and the second one (Fig. 11E) has margins converging from the widest portion to the base (but also straight). The shape of the pronotum in males of Mycetochara spp. can be very variable between different micropopulations and even within one micropopulation (including among beetles collected in different years in one locality). Both, the holotype of M. strejceki and specimens (males and females) of M. abschasica were collected in different parts of Sochi city: Matsesta and Khosta. The distance between Matsesta (the type locality of M. strejceki) and Khosta (our specimens) is 7 km. Shapes of the aedeagus figured by Novák (2022) and those shown on our Fig. 11C, D are identical. Thus, we assume that both names are synonyms, but to be absolutely sure, it is necessary to collect females and males together from Matsesta. We do not include M. strejceki in the key below, because we believe that its distinguishing characters of the pronotum are only variability (this variability was also shown by Novák (2022) on his figures 28 and 28).</p><p>Distribution. The species is known only in southern environs of Sochi (Russia, Krasnodar Region: Khosta, Macesta) (Novák 2022; present material) and Abkhazia (Georgia).</p><p>Bionomics. The species was collected in window traps, with exposition on mouldering Fagus and soil trap in Taxus forest.</p></div>	https://treatment.plazi.org/id/E80787D7FFCD1678FF723E7DFEBCF86C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD31662FF72389BFBB2F962.text	E80787D7FFD31662FF72389BFBB2F962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) zolotareffi Reitter 1896	<div><p>Mycetochara (Ernocharis) zolotareffi Reitter, 1896</p><p>(Figs 13, 14, 15)</p><p>= Mycetochara (Ernocharis) gracilicornis Roubal, 1935, syn. nov.</p><p>Type material. Syntype of M. zolotareffi, 1♀ (HNHM), examined by photo images courtesy Vladimír Novák .</p><p>Syntypes of M. gracilicornis, 1♂, 1♀ (ZMMU) pinned on one pin: “Ca. b. Teberda VI. 912 Roubal”, 2 red labels “ Syntypus ”, 2 pink labels “№ ZMMU Col 02840 Zool. Mus. Mosq. Univ. (Mosquae, ROSSIA)”, “№ ZMMU Col 02841 Zool. Mus. Mosq. Univ. ( Mosquae, ROSSIA)” . Syntypes were received by N.N. Dubrovin in an exchange with his Czech colleagues.</p><p>Material. Russia. 1♀ (ZMSFU), Krasnodar Region, Mezmay, vi.1990 (P. Bagin) ; 1♀ (ZMSFU), same place, 15– 20.vi.1992 (E.A. Khachikov); 1♀ (ZMSFU), same place, 20.voo.1996 (Yu. G. Arzanov); 1♀ (ZMSFU), Krasnodar Region, Verkhnee Makopse, 10.viii.1996 (M.V. Nabozhenko) ; 1♂ (ZMSFU), Krasnodar Region, Temnolesskaya, 17–22.vii.1999 (V. Grebennikov) ; 1♂ (ZMSFU), Krasnodar Region, Novoprokhladnyy, 27.v.2007 (M. Saprykin) ; 7♂♂, 13♀♀ (ZIN, PCMN), Krasnodar Region, Goryachy Klyuch env., 6–7.v.2019 (M. V. and S.V. Nabozhenko) ; 2♂♂, 2♀♀ (PCMN), Krasnodar Region, Sochi, Adler District, Imereti lowland, 22.iv.–3.v.2021 (E.A. Khachikov) ; 1♂, 6♀♀ (PCMN), Krasnodar Region, Kabardinks, 9–10.v.2022 (M. V. and S.V. Nabozhenko) ; 5♀♀ (PCMN), Krasnodar Region, Bakinskaya, 44.7411ʹN / 39.2887˚E, 18–19.v.2023 (I.A. Zabaluev) ; 1♂ (ZMSFU), Krasnodar Region, Sukko environs, 26–30.iv.2018 (E.A. Khachikov) ; 1♀ (PCMN), Krasnodar Region, 4 km E Sukko village, 44.7892˚N / 37.4794˚E, 15.v.2023 (I.A. Zabaluev) ; 1♀ (PCMN), Krasnodar Region, Gelendzhik, Yashamba River, 44.5916˚N / 37.9866˚E (I.A. Zabaluev) ; 1♂ (PCMN), Krasnodar Region, Novokrymsky, 44.9246˚N / 37.7946˚E, 22.v.2023 (I.A. Zabaluev) ; 1♂ (PCMN), Krasnodar Region, Ubinskaya, 44.7225˚N / 38.5330˚E, 20.v.2023 (I.A. Zabaluev) ; 1♀ (ZMSFU), Adygea, Maykop env., N coast of Belaya River, 21.vi.1997 (collector unknown) ; 1♂ (ZMSFU), Adygea, environs of Guzeripl, window trap on pine, 1.vi–22.vi.2001 (A.R. Bibin) ; 1♀ (ZMSFU), Adygea, Maykop Distr., sanatorium “Lesnaya skazka” ( Forest tale), environs of ravin “Polkovtitskaya”, 44˚20.724ʹN, 40˚11.368ʹ, soil traps, 1–25.v.2014 (A.R. Bibin) ; 1♂ (PCMN), Stavropol Region, Zheleznovodsk, road to Zheleznaya Mt., 44.1437˚N / 43.02230˚E, litter, 6.v.2023 (I.A. Zabaluev); Karachay-Cherkessia, Teberda, 20.vi.1993 (Yu. G. Arzanov) ; 4♂♂, 3♀♀ (ZIN, PCMN), Karachay-Cherkessia, Zakan, 43˚41ʹ56ʺN / 40˚47ʹ35ʺE, 1250–1300 m, 15–19.vi.2025 (M. V. and S.V. Nabozhenko) . Georgia. 1♂, 8♀♀ (PCMN), Abkhazia, Gudauta, Dzindzaria Street, 6.v.2022 (O.S. Guskova, S.V. Nabozhenko) ; 1♀ (ZMSFU), Adjara, “Batum”; 2♀♀ (PCMN), Oni District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.547806&amp;materialsCitation.latitude=42.6455" title="Search Plazi for locations around (long 43.547806/lat 42.6455)">Utsera</a>, 42°38ʹ43.8ʺN / 43°32ʹ52.1ʺE, 1050 m, 14– 15.06.2013 (M. V. and S.V. Nabozhenko) .</p><p>Variability. Specimens from foothills and the Caucasian coastal zone of Black Sea have dorsal pubescence of light yellowish setae (correspond to syntypes of M. zolotareffi from Uch-Dere) (Fig. 15). Specimens from mountain areas of the Caucasus and mountain Crimea (to 1400 m both in the Caucasus and Crimea) are different entirely black pubescence (correspond to syntypes of M. gracilicornis from Teberda) (Figs 13, 14). Crimean specimens often have elytra pubescent by black or dark-brown setae and lighter pubescence of pronotum and head (although specimens with entirely light pubescence are also presented). The body colouration is also variable: from ochre, light brown or brown body and yellowish legs and distal and proximal antennomeres in foothills and coastal micropopulations to black body and reddish legs and proximal and distal antennomeres in mountain micropopulations. Middle antennomeres are also darker in mountain individuals. The male pronotum is also very variable: from almost square in foothill and coastal specimens to trapezoidal in several mountain micropopulations. It is worth noting that a wide range of the pronotal shape can be represented even in a single series of beetles collected together or in different years. Males from Karachay-Cherkessia have antennomeres slightly thinner, than males from other territories (compare Figs. 13A and 15C), but this character variable in different micropopulations with thickest ones in some Crimean specimens. The male genitalia are identical throughout all the range.</p><p>Synonymy. Roubal (1935) described darker mountain specimens of M. zolotareffi from Teberda (Karachay-Cherkessia, Russia) as a separate species. Two examined syntypes have dark-brown body with brown dorsal pubescence. Thus, the following synonymy is established: Mycetochara (Ernocharis) zolotareffi Reitter, 1896 = Mycetochara (Ernocharis) gracilicornis Roubal, 1935, syn. nov.</p><p>Notes. Novák (2020a) recorded M. obtusicollis Reitter, 1899 for Armenia based on the female holotype and Russia (Karachay-Cherkessia: Zakan) based on five males. We conducted a comprehensive collection of beetles in Zakan village and environs in 2025 and found three species: M. zolotareffi (dark form), M. angustifrons and M. ingushetica . This is the largest number of sympatric species we have found in a very small area. We collected seven specimens of the M. zolotareffi, including females, which are distinctly different from the female holotype of M. obtusicollis by the shape of the pronotum. The contour of parameres drawn by Novák (2020a) corresponds to those in M. zolotareffi, but without spines on lateral sides. In our experience, spines can easily fall off during dissection, so we are confident that the five males from Zakan listed as M. obtusicollis belong to M. zolotareffi .</p><p>Iablokoff-Khnzorian (1983) incorrectly synonymized M. obtusicollis (type locality “Alagoes” = Aragats Mt.) with M. linearis Illiger, 1794 (now the junior synonym of M. maura (Fabricius, 1792)) . He mentioned that in Armenia this species occurs in open habitats, mountain steppes, under stones and in gerbil burrows. Only one Transcaucasian species, M. armeniaca Novák, 2022, was collected in such habitats (Nabozhenko &amp; Gadaborsheva 2023), which sharply distinguishes it from the forest xylobiont M. maura . We checked specimens in the collection of Iablokoff-Khnzorian (PCMK) from the type locality Aragats as well as specimens from other localities. All of them belong to M. obtusicollis / M. armeniaca . We have not found any significant differences between M. obtusicollis and M. armeniaca and believe that these names should be synonymized. In any case, M. obtusicollis should be excluded from the fauna of Russia.</p><p>Distribution. Russia: Krasnodar (foothills and low mountains, coast of Black Sea) (Nikitsky et al. 2008; as M. gracilicornis) and Stavropol (Caucasian Mineral Waters area) regions (Nabozhenko 2022, as M. gracilicornis; PCMN), Karachay-Cherkessia (Teberda, Zakan), Crimean peninsula: southern part, Yalta, Alma valley, Baydar valley (ZIN); Georgia, Abkhazia (Nabozhenko 2022, as M. gracilicornis), Kazbek Mt. (Reitter 1896), Utsera, Batumi (present material). The species was erroneously listed also for Turkey (Klein-Asien) by Seidlitz (1896) based on specimens from Reitter’s collection and repeatedly in several catalogues, including the last one (Novák 2020b). Reitter (1908) already wrote about this error of G. Seidlitz and once again reminded that the species originates from the Caucasus. Thus, the species must be excluded from Turkish fauna. Since the species was collected in Batumi 20 km from the Turkish border, it may be found in northeastern Turkey in the future.</p></div>	https://treatment.plazi.org/id/E80787D7FFD31662FF72389BFBB2F962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD71662FF723B35FD50F8AE.text	E80787D7FFD71662FF723B35FD50F8AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Oculochara) Novak 2020	<div><p>Subgenus Oculochara Novák, 2020</p><p>Type species: Mycetochara ocularis Reitter, 1884</p></div>	https://treatment.plazi.org/id/E80787D7FFD71662FF723B35FD50F8AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD41660FF723CACFEA8FB7E.text	E80787D7FFD41660FF723CACFEA8FB7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Oculochara) adygea Nabozhenko & Martynov & Bulysheva 2025	<div><p>Mycetochara (Oculochara) adygea Nabozhenko, sp. nov.</p><p>(Fig. 16)</p><p>Material. Holotype, ♂ (ZMSFU): Russia, Adygea Republic, Nikel, 26.vii.1996 (D.G. Kasatkin) ; Paratypes, 2♂♂ (ZMSFU): Russia, Adygea Republic, Guzeripl’ environs, window trap on Pinus, 22.vi.2001 (A.R. Bibin). All type specimens with incomplete antennae, maximum with eight antennomeres .</p><p>Description. Holotype. Body slender, narrow, shiny, head and pronotum light-brown, elytra reddish-brown, with reddish spot in basal 3/4 (spots large, but not reaching elytral suture), legs, mouthparts and prosternum reddish, antennae reddish, with antennomeres 4–8 slightly darker. Body covered with suberected light-reddish setae.</p><p>Head. Anterior margin of epistoma slightly rounded, lateral margin of genae straight, strongly converging from eyes to anterior margin of epistome. Dorsal side with sparse smooth (edges not distinct) puncturation (interpuncture distance near twice as long as puncture diameter), each puncture bears long suberected setae. Head ventrally and laterally with long smooth wrinkles along posterior edge of eyes, without visible puncturation. Eyes large, transverse section of one eye 1.12 times as long as interocular space. OI = 31.</p><p>Prothorax. Pronotum transverse (1.4 times as wide as long), widest at base, 1.27 times as wide as head; PI = 72. Lateral margins rounded at anterior two thirds and slightly emarginated at basal third, converging from base to anterior margin. Anterior margin slightly rounded, base bisinuate, middle portion protruded. Disc slightly convex only laterally in anterior half, slightly depressed at middle basally and with stronger transverse impressions at basal portions on sides from middle. Puncturation of disc moderately coarse and sparse, punctures round, interpuncture distance 2–3 times as long as puncture diameter, punctures larger along base and in posterior angles; each puncture bears long subrecumbent light-reddish seta. Anterior angles not expressed, pronotum widely rounded antero-laterally; posterior angles acute. Edges of pronotum not margined. Prothoracic hypomera and prosternum shiny, with fine and sparse puncturation and short setation.</p><p>Pterothorax. Elytra elongate, subparallel when completely closed, 1.29–1.3 times wider at base (immediately behind humeral angles) than base of pronotum. Humeral angles widely rounded. Striae consist of round slightly raduliform dense punctures, interstriae 1 and 2 slightly convex, rest ones flattened, each interstria contains one irregular row of punctures; strial punctures 2 times larger than interstrial ones; each puncture bears subrecumbent light-reddish setae. Mesoventrite and metepimera with dense and coarse puncturation; mesepisterna and mesepimera with sparse irregular puncturation; metaventrite convex, moderately coarsely punctured at anterior portion and sparser punctured at basal half.</p><p>Legs slender, long, reddish; profemora shortest, metafemora longest; pro- and metatibiae straight, mesotibiae slightly curved in holotype and straight in paratypes; each protarsal claw with 10 teeth on inner side.</p><p>Abdomen. Abdominal ventrites punctured by sparse and fine raduliform punctures and covered with recumbent setae. Aedeagus with basal piece 3.2 times longer than parameres. Parameres flattened dorso-ventrally, dorsally with slightly emarginated in middle lateral margins, widely rounded at apex, slightly S-shaped laterally, with two rows of spines dorsally. Penis smooth, without sculpture.</p><p>Variability. In males (paratypes) from Guzeripl PI = 75 and 75.7. One male has reddish elytral spots reaching sutural angles and forms band in the basal quarter of the elytra. OI in both specimens = 31.8.</p><p>Etymology. The name derives from Adygea, the republic in Russia, from which this species is described. Indeclinable non-Latinized name (Article 31.2.3 of IZCN).</p><p>Comparative diagnosis. The new species is most similar to three species from Talysh Mts: M. ocularis Reitter, 1884, M. orszuliki Novák, 2020 and M. masalliica Novák, 2020 by the presence of reddish spots in humeral area of elytra, but differs from them by the ocular index (OI = 31–31.8 in M. adygea sp. nov. vs 18.35–25.93 in compared species) and the shape of the aedeagus with distinct spines (compare Fig. 16E, F in the present paper with Figs 60–61, 67–68 and 71–72 in Novák (2020a)).</p><p>Distribution. Mountain part of Adygea Republic (North Caucasus, Russia). This is the first record of the subgenus Oculochara in the Greater Caucasus. Other species are known from the south of Anatolia (Mardin and Muğla provinces) and the South Caspian region (Talysh in Iran and Azerbaijan and Zagros Mts in Iran) (Novák 2020a, 2022).</p></div>	https://treatment.plazi.org/id/E80787D7FFD41660FF723CACFEA8FB7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD51660FF7238C9FD5EFA8A.text	E80787D7FFD51660FF7238C9FD5EFA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Pterna) Reitter 1884	<div><p>Subgenus Pterna Reitter, 1884</p><p>Type species: Ernocharis auricoma Reitter, 1884</p></div>	https://treatment.plazi.org/id/E80787D7FFD51660FF7238C9FD5EFA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD51660FF72399DFCCCF866.text	E80787D7FFD51660FF72399DFCCCF866.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Pterna) auricoma (Reitter 1884)	<div><p>Mycetochara (Pterna) auricoma (Reitter, 1884)</p><p>(Fig. 17)</p><p>Material. We examined more than 20 specimens in the collection of ZMMU (Crimea, Lower Volga region, W Kazakhstan), listed by Dubrovina (1976).</p><p>10 specimens (ZIN), Russia, Stavropol Region, Divnoe, 19.v.1953 (O.L. Kryzhanovskij) ; 1♂, 1♀ (ZIN), Russia, Krasnodar Region, Taman Peninsula, Volna, marine coast, 23. v. and 21.vi.1979 (B.A. Korotyaev) ; 1♀ (ZIN), Russia, Krasnodar Region, Taman Peninsula, Taman, 27.v.1979 (B.A. Korotyaev) ; 2♂♂, 3♀♀ (ZMSFU), Russia, Kalmykia, Lysyy Liman Lake, 8.vi.1977 (Kuznetsova) ; 1♀ (ZMSFU), Russia, Rostov Region, Rostov State Reserve, Manych-Gudilo Lake, Vodny island, 2.vi.2017 (E.A. Khachikov) ; 1♀ (PCEK), Russia, Volgograd Region, Volgograd, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=44.218647&amp;materialsCitation.latitude=48.614594" title="Search Plazi for locations around (long 44.218647/lat 48.614594)">Sovetsky District</a>, 3.5 km NE of Vodny village, 48°36’52.54”N, 44°13’7.13”E, five-year fallow, soil trap, 13.vi.2024 (E.V. Komarov) .</p><p>Distribution. Widely distributed in steppes (usually saline habitats) from mouth of the Danube in Odessa Region of Ukraine through Crimean and Taman peninsulas, Rostov, Stavropol and Volgograd regions, Kalmykia to Western Kazakhstan (Zhanybek) in the east (Dubrovina 1976).</p></div>	https://treatment.plazi.org/id/E80787D7FFD51660FF72399DFCCCF866	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
E80787D7FFD8166DFF723E88FC2CFC5D.text	E80787D7FFD8166DFF723E88FC2CFC5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mycetochara (Ernocharis) recondita Novak 2022	<div><p>Mycetochara (Ernocharis) recondita Novák, 2022</p><p>Material. 1♀ (PCMK), Armenia, Lori Prov., Teghut mine, 41.0743˚N, 448399˚E, 1105 m, soil traps, 20.vi.–1.vii.2013 (M.Yu. Kalashian) ; 1♂ (PCMK), Armenia, Lori Prov., Teghut mine, near refuse heap, 41.0705˚N, 448378˚E, 1210 m, 19.vi.–23.vii.2015 (G.H. Karagyan, M.Yu. Kalashian) .</p><p>Distribution. Georgia, Armenia (new record for the country), Iran.</p></div>	https://treatment.plazi.org/id/E80787D7FFD8166DFF723E88FC2CFC5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Nabozhenko, Maxim V.;Martynov, Vladimir V.;Bulysheva, Natalia I.	Nabozhenko, Maxim V., Martynov, Vladimir V., Bulysheva, Natalia I. (2025): Two new species of Mycetochara Guérin-Méneville, 1827 (Coleoptera: Tenebrionidae: Alleculinae) from the Northern Cis-Azov region and the Northwestern Caucasus (Russia) with a new synonymy and a key to species of the European part of Russia. Zootaxa 5706 (3): 397-425, DOI: 10.11646/zootaxa.5706.3.5, URL: https://doi.org/10.11646/zootaxa.5706.3.5
