identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
EE4BE285C0D15529BB4BC86382DC91A5.text	EE4BE285C0D15529BB4BC86382DC91A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla australis (Fabricius 1787)	<div><p>Deleproctophylla australis (Fabricius, 1787)</p><p>Figs 2 A, 2 B, 4 A – D, 5 A, 5 B, 6–8</p><p>Ascalaphus australis Fabricius, 1787: 250 (type locality: “ Europa australiore ” = southern Europe; holotype in NHMD). Gmelin 1788: 2645 (Myrmeleon). Villers 1789: 62 (Myrmeleon). Olivier 1790: 245 (Ascalaphus). Fabricius 1793: 96 (Ascalaphus). Turton 1800: 405 (Myrmeleon). Angelini 1827: 468 (Ascalaphus). Lefèbvre 1842: 6 (Deleproctophylla). Rambur 1842: 351 (Theleproctophylla). Schneider 1845: 341 (Theleproctophylla). Walker 1853: 416 (Ascalaphus). Hagen 1860: 53 (Theleproctophylla). Stein 1863: 420 (Ascalaphus). Hagen 1866: 402 (Deleproctophylla). Costa 1866: 14 (Theleproctophylla). McLachlan 1873: 260 (Theleproctophylla). Navás 1905: 547 (Theleproctophylla). Navás 1911: 92 (Theleproctophylla). Navás 1912 a: 124 (Theleproctophylla). Navás 1912 b: 34 (Theleproctophylla). Lacroix 1913: 100 (Theleproctophylla). Kimmins 1930: 188 (Theleproctophylla). Mosely 1932: 182 (Theleproctophylla). Táborský 1936: 164 (Deleproctophylla). Capra 1936: 213 (Theleproctophylla). Capra 1937: 58 (Theleproctophylla). Zimsen 1964: 612 (Ascalaphus). Zelený 1964: 326 (Theleproctophylla). Popov 1977: 276 (Deleproctophylla). H. Aspöck et al. 1980: 315 (Deleproctophylla). Devetak 1992: 111 (Deleproctophylla). Zakharenko and Krivokhatsky 1993: 73 (Deleproctophylla). Bernardi et al. 1995: 11 (Deleproctophylla). Devetak 1995: 194 (Deleproctophylla). Letardi and Pantaleoni 1996: 299 (Deleproctophylla). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). Pantaleoni and Letardi 1998: 40 (Deleproctophylla). H. Aspöck et al. 2001: 301 (Deleproctophylla). Whittington 2002: 377 (Deleproctophylla). Popov 2004: 231 (Deleproctophylla). Canbulat 2007: 36 (Deleproctophylla). Dobosz and Ábrahám 2007: 24 (Deleproctophylla). Devetak and Janžekovič 2012: 183 (Deleproctophylla). Colombo et al. 2012: 125 (Deleproctophylla). Badano and Pantaleoni 2014: 301 (larva, Deleproctophylla). Letardi 2016: 194 (Deleproctophylla). Aistleitner 2019: 123 (Deleproctophylla). Jones 2019: 3 (Deleproctophylla). Poggi 2024: 9 (Deleproctophylla).</p><p>Theleproctophylla australis var. impar Navás, 1923: 154 (type locality: Greece: Thessaloniki; holotype in MNHN but lost at present).</p><p>Examined material.</p><p>• Croatia: Dalmatien, Hvar, 17. VII. 1962, R. Löberbauer, 1 ♀ (HUAC) ; • Croatia: Dalmatien-Split, Küste, 22. VI. 1953, H. Hamann, 1 ♂ 1 ♀ (HUAC) ; • Croatia: Ribnica, Otavice 10 km N v. Titov Velea, 28. VII. 1965, 1 ♂ 1 ♀ (RMNH) ; Croatia: • Rovinj, 15. V. 1953, 1 ♂ (HUAC) ; • Croatia: Ninski Stanovi, 40 m, 14. 07. 2004, L. Ábrahám, 31 ♂♂ 34 ♀♀ (SCM) ; • Croatia: Zadar, Benkovac, 11. 07. 2002, D. Szalóki, 3 ♀♀ (SCM) ; • Croatia: Dalmacija, Gor. Poličnik, 12 km E of Zadar, 20. 06. 2007, leg. Zs. Józan, 1 ♀ (SCM) ; • Croatia: Dalmacija, Primošten-Bilo, 09. 06. 2003, Nikola Rahmé 1 ♂ (SCM) ; • Croatia: Vinodol Obc., Crikvenica, Selce, 180 m, 01. 07. 2001, K. Székely 1 ♂ (SCM) ; • Croatia: Ninski Stanovi, 40 m, 14. 07. 2004, L. Ábrahám, 3 ♂♂ 3 ♀♀ (USMB) ; • France: Corse, Calvi, 13–27. VII. 1971, Fam. V. Oorschot, 1 ♀ (RMNH) ; • France: Corse, Belgodère, 350 m, 14. VII. 1971, Fam. V. Oorschot, 1 ♀ (RMNH) ; • France: Corse, Belvedere, VII. 1973, H. M. Steiner, 3 ♂ 1 ♀ (HUAC) ; • France: Corsica, Valle de Restino, 02. 06. 2000, leg. L. Nádai, 1 ♀ (SCM) ; • France: Haute-Corse, Castifao, 25. 06. 2002, leg. J. Bard, 1 ♀ (SCM) ; • Greece: Ahaia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.883333&amp;materialsCitation.latitude=38.009445" title="Search Plazi for locations around (long 21.883333/lat 38.009445)">Michas</a>, 1300 m, 38°00'34"N, 21°53'00"E, 7. VII. 2008, Dils J. and Faes J., 1 ♀ (RMNH) ; • Greece: Chalcide, Tristinika, 20. VI. 2015, 1 ♂ (MDC) ; • Greece: Cos, Tigaki, 3–10. VII. 1978, M. C. and G. Kruseman, 2 ♀ (RMNH) ; • Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.266666&amp;materialsCitation.latitude=35.35" title="Search Plazi for locations around (long 24.266666/lat 35.35)">Georgioupolis</a>, 35°21'N, 24°16'E, H. u. L. Hölzel, 24 ♂ 19 ♀ (NHMW) ; • Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.266666&amp;materialsCitation.latitude=35.35" title="Search Plazi for locations around (long 24.266666/lat 35.35)">Georgioupolis</a>, 35°21'N, 24°16'E, H. u. L. Hölzel, 1 ♂ 1 ♀ (HUAC) ; • Greece: Elis, Andritsaina, 1000 m, 29. VII. 1978, L. Willemse and J. Tilmans, 1 ♂ (RMNH) ; • Greece: Ioannina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=20.636778&amp;materialsCitation.latitude=39.938557" title="Search Plazi for locations around (long 20.636778/lat 39.938557)">Mesovouni</a>, 650 m, 39°56'18.8"N, 20°38'12.4"E, 28. VII. 2006, T., W. and J. Garrevoet, 2 ♀ (RMNH) ; • Greece: Lesbos, Sigri, 6–8. VI. 1988, J. P. Duffels, 1 ♂ 1 ♀ (RMNH) ; • Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.0&amp;materialsCitation.latitude=38.0" title="Search Plazi for locations around (long 23.0/lat 38.0)">Loutraki-Perachora</a>, 38°N, 23°E, 30. V. 1969, Rausch, 1 ♀ (HUAC) ; • Greece: Pierias, Litochoro, 5. VII. 1976, F. and L. Willemse and J. Tilmans, 1 ♀ (RMNH) ; • Greece, Creta Nord, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.398306&amp;materialsCitation.latitude=35.31008" title="Search Plazi for locations around (long 25.398306/lat 35.31008)">Hersonissos</a>, 35°18'36.3"N, 25°23'53.9"E, 15. V. 2024, J. Vetrovec, 1 ♂ (DB) ; • Greece: Creta, Lasithi, Xerokampos, 2. VI. 1972, M. C. and G. Kruseman, 1 ♂ 2 ♀ (RMNH) ; • Greece: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.3095&amp;materialsCitation.latitude=37.656364" title="Search Plazi for locations around (long 22.3095/lat 37.656364)">Kardaras</a>, 800 m, 37°39'22.9"N, 22°18'34.2"E, 7. VII. 2008, Dils J. and Faes J., 1 ♂ (RMNH) ; • Greece: Tripolis-Davia 3–8 km NW of Tripolis, 800 m, 13. VII. 1975, J. P. Duffels, 1 ♀ (RMNH) ; • Greece: Leptokaria, 28. 06. 1992, Balásházy, 1 ♀ (SCM) ; • Greece: Sithonia, Sarti, 0–50 m, 02–09.07. 1997, D. Szalóki, 1 ♂ 4 ♀♀ (SCM) ; • Greece: Sithonia, Sarti, 0–50 m, 02–09.07. 1997, D. Szalóki, 1 ♀ (USMB) ; • Greece: Argolída, SE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.15&amp;materialsCitation.latitude=37.666668" title="Search Plazi for locations around (long 23.15/lat 37.666668)">Néa Epídavros</a>, 37°40'N, 23°09'E, 120 m, 09. 06. 2007, R. Królik, 1 ♂ (USMB) ; • Greece: Corfu, Acharavi [CK 90], xerothermic meadow, 06. 07. 2011, W. Żyła, 3 ♂♂ (USMB) ; • Greece: Corfu, Acharavi [CK 90], xerothermic meadow, 07. 07. 2011, W. Żyła, 1 ♂ (USMB) ; • Greece: Chalkidiki, Paliouri env., 39°56'31"N, 23°39'29"E, SE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.658054&amp;materialsCitation.latitude=39.941944" title="Search Plazi for locations around (long 23.658054/lat 39.941944)">Kassandria</a>, 06–11.06. 2013, M. Šnižek, 2 ♂♂ (USMB) ; • Greece: Chalkidiki, Paliouri env., 39°56'31"N, 23°39'29"E, SE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.658054&amp;materialsCitation.latitude=39.941944" title="Search Plazi for locations around (long 23.658054/lat 39.941944)">Kassandria</a>, 12–14.06. 2013, M. Šnižek, 3 ♂♂ (USMB) ; • Greece: Thessaly, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.783333&amp;materialsCitation.latitude=39.816666" title="Search Plazi for locations around (long 22.783333/lat 39.816666)">Kokkino Nero</a>, 39°49'N, 22°47'E, environs, 17–29.05. 2015, A. Skiba, 1 ♂ (USMB) ; • Greece: Chalkidiki, E of Paliouri, 39°56'N, 23°41'E, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.683332&amp;materialsCitation.latitude=39.933334" title="Search Plazi for locations around (long 23.683332/lat 39.933334)">Kassandria</a>, 11. 06. 2022, M. Šnižek, 1 ♂ (USMB) ; • Greece: Peloponnese, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.11028&amp;materialsCitation.latitude=37.853333" title="Search Plazi for locations around (long 21.11028/lat 37.853333)">Loutra Killinis</a>, 37°51'12"N, 21°6'37"E, dunes and grass, 19. 06. 2023, W. Żyła, 1 ♀ (USMB) ; • Greece: Peloponnese, Messenia, SE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.119083&amp;materialsCitation.latitude=37.870415" title="Search Plazi for locations around (long 21.119083/lat 37.870415)">Kalamia</a>, 37°52'13.5"N, 21°07'08.7"E, olive garden, 13. 06. 2024, W. Żyła, 2 ♀♀ (USMB) ; • Greece: Peloponnese, Messenia, SE of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=21.119083&amp;materialsCitation.latitude=37.870415" title="Search Plazi for locations around (long 21.119083/lat 37.870415)">Kalamia</a>, 37°52'13.5"N, 21°07'08.7"E, olive garden, 14. 06. 2024, W. Żyła, 1 ♂ 1 ♀ (in absolute alcohol) (USMB) ; • Italy: Basilicata, 10 km uoru, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.75&amp;materialsCitation.latitude=40.583332" title="Search Plazi for locations around (long 15.75/lat 40.583332)">Potenza</a>, 700 m, 40°35'N, 15°45'E, 1 ♂ (HUAC) ; • Italy: Calabria, Crotone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.916666&amp;materialsCitation.latitude=38.916668" title="Search Plazi for locations around (long 16.916666/lat 38.916668)">Catanzaro</a>, 30–100 m, 38°55'N, 16°55'E, 4. VIII, 1 ♂ (HUAC) ; • Italy: Prov. Messina, Lipari, Isole, Stromboli, San Vincenzo, 17. VII. 1995, Alain Olivier, 1 ♀ (RMNH) ; • Italy: Tuscany, Grosseto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.1472225&amp;materialsCitation.latitude=42.83972" title="Search Plazi for locations around (long 11.1472225/lat 42.83972)">Nomadelfia</a> (GR), 42°50'23"N, 11°08'50"E, 29. VI. 2024, Davide Badano and Yuchen Zheng, 2 ♂ 2 ♀ (ZCAU) ; • same locality • 1 ♂ 3 ♀ (DB); • Italy: Tuscany, Grosseto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.168056&amp;materialsCitation.latitude=42.82611" title="Search Plazi for locations around (long 11.168056/lat 42.82611)">Roselle</a> (GR), 42°49'34"N, 11°10'05"E, 29. VI. 2024, Davide Badano and Yuchen Zheng, 1 ♂ 2 ♀ (ZCAU) ; • same locality data • 1 ♀ (DB); • Italy: Tuscany, Grosseto, Parco Nazionale della Maremma, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.040278&amp;materialsCitation.latitude=42.66417" title="Search Plazi for locations around (long 11.040278/lat 42.66417)">Marina di Alberese (GR)</a>, 42°39'51"N, 11°02'25"E 20. VI. 2025, Davide Badano leg., 1 ♂ 2 ♀ (DB) ; • Italy: Lazio, Roma (RM), Via Appia Antica, 11. VII. 1937, L. Barbera, 1 ♀ (MSNG) ; • Italy: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=11.147611&amp;materialsCitation.latitude=42.839775" title="Search Plazi for locations around (long 11.147611/lat 42.839775)">Nomadelfia (GR)</a>, 42°50'23.2"N, 11°08'51.4"E, 30. VI. 2024, D. Badano and Y. Zheng, (DB) ; • Italy: Puglia, Bari, Masseria Pellicciari, 450 m, 9. VII. 2000, Fontana P., Kleukers R. and Cogo A., 1 ♀ (RMNH) ; • Italy: Puglia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.271194&amp;materialsCitation.latitude=41.089138" title="Search Plazi for locations around (long 16.271194/lat 41.089138)">Castel del Monte</a>, 450 m, 41°05'20.9"N, 16°16'16.3"E, 2. VII. 2001, T. and W. Garrevoet, 1 ♀ (RMNH) ; • Italy: Prov. Salerno, Pioppi, Cost Cilentana, 14. VII. 1971, Holzinger, 1 ♀ (HUAC) ; • Italy: Sardegna, Marrubiu (OR), Sant’Anna, stazione e dintorni, 1. VI. 2003, C. Meloni, 1 same locality, ♂ (MSNG) ; • same locality, 2. VI. 2006, 1 ♀ (MSNG); • same locality, 26. VI. 2009, C. Meloni, 3 ♂ 12 ♀ (MSNG); • Italy: Sardegna, Alghero (SS), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.324917&amp;materialsCitation.latitude=40.54036" title="Search Plazi for locations around (long 8.324917/lat 40.54036)">Lu Cantar</a>, 40°32'25.3"N, 8°19'29.7"E, 12. VII. 2012, D. Badano, 3 ♂ (DB) ; • Italy: Sardegna, Alghero (SS), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.251667&amp;materialsCitation.latitude=40.605583" title="Search Plazi for locations around (long 8.251667/lat 40.605583)">Monte Doglia</a>, 40°36'20.1"N, 8°15'06.0"E, 7. VII. 2012, D. Badano, 3 ♀ (DB) ; • Italy: Sardegna, Monti (OT), 2. VII. 2011, D. Badano, 2 ♀ (DB) ; • Italy: Sicilia, Sud di Castrofilippo (AG), m 450, 4. VI. 2015, E. Gallo, 1 ♀ (MSNG) ; • Italy: Sicilia, Caltanissetta (CL), strada Enna, bivio Gela, 8. VII. 1954, Adalberto Parvis, 2 ♂ 3 ♀ (MSNG) ; • Italy: Sicilia, Peloritani, 10 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=15.07&amp;materialsCitation.latitude=38.06" title="Search Plazi for locations around (long 15.07/lat 38.06)">Barcellona</a>, 100 m, 38.06°N, 15.07°E, 25–26.1974, 2 ♂ 2 ♀ (HUAC) ; • Italy: Sardinia, Siniscola, Su Tilio, 27. 06. 2004, V. Feik, 1 ♀ (NMPC) ; • Italy: Sardinia, “ Sardinien, Mti Gennargentu Mte, Spada ”, 1250 m, 17–21.06. 1968, Ströhle, 2 ♂♂ (SCM) ; • North Macedonia: Demir Kapija, Barovo, 27. VII. 1965, 1 ♀ (RMNH) ; • North Macedonia: Štip, 23. VII. 1965, 1 ♂ (RMNH); • Turkey: Aydin, Kusadasi, 11. VII. 1980, H. v. Oorschot, 1 ♀ (RMNH) ; • Turkey: Balıkesir, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=27.47&amp;materialsCitation.latitude=40.15" title="Search Plazi for locations around (long 27.47/lat 40.15)">nahe Bezirci (zw. Edincik u. Bugdayli)</a>, 150 m, 40.15°N, 27.47°E, 22. VII. 1978, H. and U. Aspöck, H and R Rausch, P. Ressl, 3 ♂ 4 ♀ (HUAC) ; • Turkey: Edirne, 08–13.06. [19] 47, 1 ♂, Exp. N. Mus. ČRS (NMPC) .</p><p>Diagnostic characters.</p><p>Metafemur with brown marking; forewing with brown marking below pterostigma (often absent in eastern populations); hindwing elongate, posterior margin curved; not angled; hindwing with large marking below pterostigma; male ectoproct with branch in apical half of branch, after mid-length; setae on branch only at tip; setae on branch and tip of ectoproct stout and curved.</p><p>Redescription.</p><p>Size (mm, based on 5 specimens). Head + thorax length 7.7–8.6; antenna length 15.1–2; fore wing length 21.7–22.9; fore wing width 6.4–7.4; hindwing length 17.1–18.7; hindwing width 5.0–5.7.</p><p>Head. Vertex yellowish brown; setae dense, long, pale yellow (Fig. 6 C). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 6 B). Mandible yellowish brown, transitioning to dark brown apically. Labium and palp yellowish brown, concolorous with mandible. Antenna with pedicel with dense pale and dark setae, flagellomeres brown, nodes slightly darker; club brown, anterior face paler (Fig. 6 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum brown with yellowish brown markings: an anterior rounded pair, a median triangular pair, and a posterior rounded pair; mesoscutellum anteriorly dark brown, posteriorly yellowish brown. Mesonotum with long pale yellow setae. Metanotum largely yellowish brown, medially dark brown; setae pale yellow (Fig. 6 C). Pleurae yellowish brown; setae pale yellow (Fig. 6 D).</p><p>Legs. Pro- and mesothoracic legs yellowish brown, tarsi slightly darker. Metathoracic leg yellowish brown, metafemur with dorsal dark brown marking. Coxae with thin pale yellow setae; femora and tibiae with thin pale yellow setae and ventral thicker blackish setae; tibial spurs narrow, blackish; tarsi with robust blackish setae (Fig. 6 D).</p><p>Wings. Forewing long, apex rounded. Pterostigma with four to five forked and unforked brown veinlets; membrane brown. Presectoral area with ~ 6 crossveins. RP with five branches. MA gradually curving toward hind margin in distal portion. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly pronounced. Forewing venation largely dark brown, paler toward wing base; Sc and R light brown; anal veins pale brown. Forewing membrane hyaline (in old specimens with faint brownish), distal section usually with a brown marking in radial area, just below pterostigma (absent in some populations) (Figs 2 A, 2 B, 4 A – D, 6 A). Hindwing elongated, broadest at RP fork; hind margin smoothly curved gradually narrowing toward rounded apex. Presectoral area with three crossveins. RP with four branches. MA largely straight, slightly curved toward hind margin distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation dark brown, paler near wing base. Hindwing membrane hyaline, distal section with a rather large brown marking extending from pterostigma across radial area until first branch of RP (Figs 2 A, 2 B, 4 A – D, 6 A).</p><p>Abdomen. Tergites pale brown, each with a caudal pair of yellowish markings. Sternites pale brown, paler than tergites. Pleural membrane brown with large paler markings.</p><p>Male genitalia. Ectoproct narrow, with branch in apical half, after mid-length. Setae of ectoproct long, dark brown; setae on branch limited to tip, dense, black, stout, curved, and spine-like; setae only on tip of ectoproct, black, stout, curved, and spine-like (Fig. 7 A – C). Tergite 9 sub-rectangular in lateral view, with ventral projection; setae long and black, longer and denser on projection. Sternite 9 with three lobes, of which median largest; setae thin, spine-like on median lobe (Fig. 7 B, C). Gx 11 arch-shaped, slightly curved downward in lateral view; gx 9 strongly sclerotized, with upward fold; gp 9 lightly sclerotized forming a shallow median furrow, with short setae (Fig. 7 F – J).</p><p>Female genitalia. Ectoproct elongated; setae long, black (Fig. 7 D, E). Tergite 9 sub-rectangular. Gx 9 wider than long ventrally; setae long. Tergite 8 subrectangular. Gx 8 wider than long, with long setae. Gp 8 prominent; setae thin (Fig. 7 D, E).</p><p>Larva. See Badano and Pantaleoni (2014) (Fig. 5 A, B).</p><p>Distribution.</p><p>Asia: Azerbaijan (see remarks for D. variegata), Georgia, Turkey (see remarks). Europe: Albania, Bulgaria, Croatia, France (Corsica), Greece, Italy (including Sardinia and Sicily), North Macedonia, Turkey (Fig. 25) (H. Aspöck et al. 1980; Devetak 1995; H. Aspöck et al. 2001; Popov 2004; Devetak and Janžekovič 2012; Kerimova et al. 2022; Japaridze et al. 2024; Oswald 2025).</p><p>Life history notes.</p><p>Deleproctophylla australis is typically associated with arid grasslands, meadows, and clearings (H. Aspöck et al. 1980; Devetak 1995; Popov 2004) (Fig. 8). Deleproctophylla australis is usually active at daylight, despite flights at dusk with hot weather. The female seems to prefer ovipositing on short grass stems (Fig. 8 A, B), the eggs are placed in rows, as typical for the tribe (Fig. 8 D).</p><p>Remarks.</p><p>Deleproctophylla australis is the type and first described species of Deleproctophylla . Among all congeners, D. australis is also the species with the widest distribution, ranging from the Tyrrhenian islands to western Asia (Fig. 25). However, this owlfly is usually localized and not particularly common (H. Aspöck et al. 1980, 2001).</p><p>The reliability of the forewing marks as a diagnostic character of D. australis is challenged by its absence in some specimens both due to individual variation and geographic differences (Fig. 4 A – D). Deleproctophylla australis is distinguished from congeners, particularly the highly similar D. dusmeti, by a combination of wing and genital characters; D. australis is characterized by the elongated hindwing without a prominent posterior angle (present in D. dusmeti) and male ectoproct with branch arising after mid-length (at mid-length in D. dusmeti). Western populations of D. australis are typically characterized by the presence of a forewing spot below pterostigma, a trait often considered diagnostic in distinguishing this species from its European congeners (e. g., H. Aspöck et al. 1980) (Figs 2 A, 2 B, 4 A – D, 6 A). However, the frequency of this forewing marking diminishes eastward, with the populations of the Balkan Peninsula including spotted and unspotted specimens and those of Anatolia lacking the spot (Fig. 4 D). These Asian unmarked specimens were assigned to a “ dusmeti - like morph ”, due to the shared absence of forewing marking and presence of spot on metafemur (H. Aspöck et al. 1980). The re-examination of specimens from these populations suggests that these enigmatic individuals from Anatolia belong to D. australis, as supported by wing shape and characters of male genitalia. Nevertheless, specimens of D. australis without spotted fore wings can be present in all populations, including Italy (Fig. 4 B).</p></div>	https://treatment.plazi.org/id/EE4BE285C0D15529BB4BC86382DC91A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
81ED67A9402A5FD4A0B8B49545A71696.text	81ED67A9402A5FD4A0B8B49545A71696.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla bleusei Kimmins 1949	<div><p>Deleproctophylla bleusei Kimmins, 1949</p><p>Figs 2 C, 2 D, 4 E, 4 F, 9, 10, 24 A, 24 B</p><p>Deleproctophylla bleusei Kimmins, 1949: 28 (type locality: Algeria: Naama, Mécheria; holotype in NHMUK). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). H. Aspöck et al. 2001: 302 (Deleproctophylla). Güsten 2003: 133 (Deleproctophylla). Monserrat et al. 2014: 157 (Deleproctophylla). Monserrat 2015: 478 (Deleproctophylla). Michel and François 2019: 183 (Deleproctophylla). Monserrat et al. 2022: 81 (Deleproctophylla).</p><p>Material examined.</p><p>Type specimen: Holotype. Images available at https://data.nhm.ac.uk/object/73295e3e-26cd-4bf4-8c10-0879722377ff.</p><p>Additional material.</p><p>• Morocco: Fes-el-Bali (60 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.16&amp;materialsCitation.latitude=34.38" title="Search Plazi for locations around (long -5.16/lat 34.38)">Ouezzane</a>), 34.38°N, 5.16°W, 300 m, 8. VII. 1977, 4 ♂, 6 ♀ (HUAC) ; • Spain: Alicante, Cox, VIII. 1911, Andréu, 1 ♂ (MCNM) ; • Morocco: Middle Atlas, Parc National de Tazekka, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-4.3079724&amp;materialsCitation.latitude=34.00164" title="Search Plazi for locations around (long -4.3079724/lat 34.00164)">env. of Barrage Bab Louta Lake</a>, 34°0'5.90"N, 4°18'28.70"W, 590 m a. s. l., dry, ruderal meadow, 15. 06. 2022, 2 ♂, W. Szczepański (USMB) ; • some locality, date, 1 ♂, W. Szczepański (SCM); • Spain: Almería, Cabo de Gata, 30 SWF 67, 27. V. 2010, F. Rodríguez and J. R. Correas, 1 ♂ (VM) ; • same locality 16. VI. 2013, 2 ♂, 1 ♀ (VM); • same locality 6. VII. 2013, 1 ♂, 2 ♀ (VM), 1 ♀ (DB); • Almería, Cabo de Gata, Salinas, Las Amoladeras, 30 SWF 67, 14. VI. 2012, 1 ♀ (VM) ; • same locality, 6. VII. 2013, 2 ♀ (VM); • same locality, 30. VII. 2013, 1 ♀ 1 ♂ (VM); • Spain: Almería, San José, Playa de los Genoveses, 14. VI. 2012, 3 ♂ 2 ♀ (VM) ; • Spain: Almería, San José, Playa de Mónsul, 10 m, 14. VI. 2012, 1 ♀ (VM) ; • Spain: Murcia, Lorca, Purias, 30 SXG 25, 550 m, 12. VI. 2010, José Carrillo, 1 ♀ (VM) ; • same locality, 19. VI. 2010, José Carrillo, 1 ♂ (VM) .</p><p>Diagnostic characters.</p><p>Metafemur unmarked; forewing unmarked; hindwing posterior margin markedly curved, convex; hindwing with marking below pterostigma; male ectoproct with branch long, in apical half of branch, just after mid-length; setae on branch only at tip; setae on branch and tip of ectoproct stout and curved.</p><p>Redescription.</p><p>Size (mm, based on 4 specimens) Head + thorax length 6.7–7.1; antenna length 11.6–14.5; fore wing length 15.7–22.4; fore wing width 6.5–6.7; hindwing length 19.2–22.7; hindwing width 4.2–5.5.</p><p>Head. Occiput yellowish brown; setae dense, long, pale yellow (Fig. 9 C). Vertex yellowish brown; setae dense, long, pale yellow (Fig. 9 B). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale. Mandible yellowish brown, darker apically. Labium and palp yellowish brown. Antenna with pedicel with dense pale and dark setae, flagellomeres brown, nodes slightly darker; club brown, paler anteriorly (Fig. 9 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum yellowish brown; internal margins of sclerites with darker brown shades; mesoscutellum yellowish brown with faint median brown stripe; setae long, pale yellow. Metanotum yellowish brown, with faint median marking; setae pale yellow (Fig. 9 C). Pleurae yellowish brown; setae pale yellow (Fig. 9 D).</p><p>Legs. Pro- and mesothoracic legs entirely yellowish brown. Metathoracic leg yellowish brown, unmarked. Setae of coxae pale yellow; setae of femora and tibiae thin, pale yellow, thicker blackish setae ventrally; tibial spurs narrow; setae of tarsi blackish (Fig. 9 D).</p><p>Wings. Forewing long, apex rounded. Pterostigma with ~ 4 forked and unforked brown veinlets; membrane very pale brown, almost whitish in young (but not freshly emerged) individuals. Presectoral area with ~ 5 crossveins. RP with five branches. MA progressively curving toward hind margin in distal portion. MP - CuA area with ~ 3 irregular rows of cells. Axillary lobe slightly prominent. Forewing venation largely brown, paler toward wing base; Sc and R yellowish, very pale in young individuals; anal veins yellowish. Forewing membrane hyaline (in old specimens faint brownish), unmarked (Figs 2 C, 2 D, 4 E, 4 F, 9 A). Hindwing broad, wider at RP fork; hind margin markedly convex, with a pronounced curve. Presectoral area with three crossveins. RP with four branches. MA largely straight, slightly curved toward hind margin distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation brown, paler near wing base; Sc yellowish. Hindwing membrane hyaline, distal section with a long narrow brown marking, sometimes stripe-like, extending from pterostigma to first branch of RP (Figs 2 C, 2 D, 4 E, 4 F, 9 A).</p><p>Abdomen. Tergites pale brown, each with a caudal pair of yellowish markings. Sternites pale brown, paler than tergites. Pleural membrane pale brown.</p><p>Male genitalia. Ectoproct narrow, with long and thin branch bifurcating beyond mid-length, in apical half. Setae of ectoproct long, dark brown; setae on branch only at apex, dense, black, stout, curved, and spine-like; setae on tip of ectoproct black, stout, curved, and spine-like (Fig. 10 A – C). Tergite 9 sub-rectangular in lateral view, with prominent ventral projection; setae long and black, longer and denser on projection. Sternite 9 with three lobes, of which median largest; setae thin, spine-like on median lobe (Fig. 10 B, C). Gx 11 arch-shaped, projecting caudally and curved downward laterally; gx 9 sclerotized, dorsally folded; gp 9 lightly sclerotized forming a shallow median furrow (Fig. 10 F – J).</p><p>Female genitalia. Ectoproct elongated; setae long, black (Fig. 10 D, E). Tergite 9 sub-rectangular. Gx 9 wider than long; setae long. Tergite 8 subrectangular. Gx 8 wider than long, with long setae. Gp 8 projecting ventrally; setae thin (Fig. 10 D, E).</p><p>Comparative notes.</p><p>Deleproctophylla bleusei has a long history of taxonomic confusion with D. dusmeti, to which it is sympatric in the Iberian Peninsula and closely resembles in wing pattern (Monserrat et al. 2014). However, D. bleusei can be distinguished by the unmarked metafemur (Fig. 9 D) and characters of male ectoproct, such as long and thin branch arising after mid-length (Fig. 10 A). The shape of the hindwing provides a reliable diagnostic trait: D. bleusei has smoothly curved posterior margin in D. bleusei, whereas D. dusmeti is characterized by a distinctly triangular shape. Additionally, young (but not newly hatched) specimens of D. bleusei are characterized by a pale C, Sc and R veins, as well as an almost whitish pterostigma. This distinctive venation coloration is also evident in the holotype.</p><p>Distribution.</p><p>Africa: Algeria, Morocco, Tunisia. Europe: Spain (H. Aspöck et al. 2001; Monserrat et al. 2014; Oswald 2025) (Fig. 25).</p></div>	https://treatment.plazi.org/id/81ED67A9402A5FD4A0B8B49545A71696	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
40445BE1F3D756FD871858C579950214.text	40445BE1F3D756FD871858C579950214.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla dandizenor Badano, Zheng, U. Aspock & Dobosz 2026	<div><p>Deleproctophylla dandizenor Badano, Zheng, U. Aspöck &amp; Dobosz sp. nov.</p><p>Figs 2 E, 2 F, 11, 12</p><p>Type material.</p><p>Holotype • Afghanistan: Pr. Kunar, Nuristan ob. LindaiSin-Tal vic. Barg e Matal, Dandizenor Mts., 2600 m, 13. VII. 1970, Naumann 1 ♂ (NHMW) ; paratype, same information as holotype, 1 ♀ (NHMW) . Paratypes • Afghanistan: Province Bamiyan, Yakawlang District, SW Kotak village, ca 2800 m, 03. 07. 2016, I. Pljushtch, 1 ♂ (SCM) ; • Afghanistan: Province Bamiyan, Bande-Amir National Park, 2950 m, 26. 06. 2016, I. Pljushtch, 1 ♂ (USMB) ; • Pakistan: Hindukush Mts., Booni, 2000 m, 11. 07. 1994, B. Herczig, Gy. M. László and G. Ronkay, 1 ♂ (SCM) .</p><p>Diagnosis.</p><p>Metafemur with large brown marking; pterostigma of both wings dark brown; wing venation rather dense, with many crossveins; forewing hyaline; hindwing broad; hindwing with large band-shaped marking from pterostigma to posterior margin of wing; male ectoproct robust, with branch in apical two-thirds of branch; ectoproct, including branch and apex densely covered with setae; setae on branch and apex spine-like. Male tergite 9 with caudal margin with long hair-like setae, ventral projection stout, hairy. Female ectoproct with short ventral projection. Female gx 9 and gp 8 relatively prominent.</p><p>Description.</p><p>Size (mm, based on 2 specimens) Head + thorax length 8.1–8.2; antenna length 15.4–15.5; fore wing length 20.3–21.9; fore wing width 6.5–6.8; hindwing length 15.5–17.7; hindwing width 5.4–5.5.</p><p>Head. Vertex yellowish brown; setae dense, long, pale yellow (Fig. 11 C). Para-ocular band yellowish brown, darker anteriorly. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 11 B). Mandible yellowish brown, transitioning to brown apically. Labium and palp yellowish brown, concolorous with mandible. Antenna with pedicel with pale and dark setae, flagellomeres brown; club brown (Fig. 11 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum brown with yellowish brown markings: an anterior small, rounded pair, a median subtriangular pair, and a posterior rounded pair; mesoscutellum dark brown, caudal margin yellowish brown. Mesonotum with long pale-yellow setae. Metanotum brown, medially dark brown; setae pale yellow (Fig. 11 C). Pleurae yellowish brown; setae pale yellow (Fig. 11 D).</p><p>Legs. Pro- and mesothoracic legs yellowish brown, tarsi dark brown. Metathoracic leg yellowish brown, metafemur with large dark brown marking. Coxae with thin pale yellow setae; femora and tibiae with thin pale yellow setae and ventral thicker blackish setae; tibial spurs narrow, blackish; tarsi with robust blackish setae (Fig. 11 D).</p><p>Wings. Forewing long, broad, apex rounded. Venation dense, with many crossveins. Pterostigma with four or five forked and unforked brown veinlets; membrane dark brown. Presectoral area with ~ 4 crossveins. RP with five branches. MA gradually curving towards hind margin in distal portion. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe inconspicuous. Forewing venation dark brown, Sc and R paler; anal veins pale brown. Forewing membrane hyaline. Hindwing broad, widest after RP fork; hind margin smoothly curved toward rounded apex. Presectoral area with three crossveins. RP with four branches. MA straight, only slightly curved. MP - CuA area with ~ 2 irregular rows of cells (Figs 2 E, 2 F, 11 A). Hindwing venation dark brown, paler near wing base. Hindwing membrane hyaline, distal section with a large brown marking extending from pterostigma to cubital area, shaped as curved band gradually narrowing posteriorly, pigmentation bordering crossveins, central part of cells paler; marking fainter in males than in females (Figs 2 E, 2 F, 11 A).</p><p>Abdomen. Tergites brown. Sternites brown, paler than tergites. Pleural membrane brown with large paler markings.</p><p>Male genitalia. Ectoproct robust, with branch at two-thirds of length and apex stout. Setae of ectoproct dense, long, dark brown, gradually longer caudally; setae on whole branch, denser, and spine-like at apex; setae on apex of ectoproct spine-like (Fig. 12 A – C). Tergite 9 sub-rectangular in lateral view, with robust ventral projection; setae long and black, longer and denser on caudal margin of the sclerite and on projection. Sternite 9 with lateral lobes inconspicuous, rather small, median lobe prominent, apically rounded (Fig. 12 B, C). Gx 11 arch-shaped, slightly curved; gx 9 strongly sclerotized, with prominent dorso-apical fold, giving to apex a hooked appearance, gp 9 relatively small, lightly sclerotized forming a shallow median furrow, with short setae (Fig. 12 F – J).</p><p>Female genitalia. Ectoproct elongated, relatively short and stout ventral projection; setae long, black (Fig. 12 D, E). Tergite 9 sub-rectangular. Gx 9 wider than long, prominent; setae long. Tergite 8 subrectangular. Gx 8 prominent, as long as wide, with long setae. Gp 8 projecting ventrally, with prominent caudal margin; setae thin (Fig. 12 D, E).</p><p>Distribution.</p><p>Asia: Afghanistan, Pakistan (Fig. 25).</p><p>Etymology.</p><p>The species epitheton is derived from the name of a mountain range in the east of Afghanistan. It is a substantive and an apposition to the genus.</p><p>Comparative notes.</p><p>Deleproctophylla dandizenor is a broad-winged species, easily distinguishable from all its congeners by the narrow, curved, hindwing marking that extends from pterostigma almost to the posterior wing margin. The hindwing appears more pigmented in female in comparison to male, in which the marking is reduced to an infuscation (Fig. 2 E, F). A similar difference in wing pattern is also observed in D. tengri sp. nov., despite the shape of markings is very different in these two species. However, due to the limited number of known specimens, it remains unclear whether these differences are attributable to sexual dimorphism or individual variation. The male of D. dandizenor is characterized by the robust hairy ectoproct with branch that bifurcates after mid-length, and stout apex. Moreover, the spine-like setae on the branch and on the apex of the ectoproct are rather long and slightly curved and they extend for the whole length of both structures (Fig. 12 A – C). The hair-like setae on the caudal margin of tergite 9 are also diagnostic of this species. The female of D. dandizenor differs from the other species of the genus by the relatively stout ectoproct, with very short ventral projection and the prominent gx 8 and gp 8 (Fig. 12 D, E).</p></div>	https://treatment.plazi.org/id/40445BE1F3D756FD871858C579950214	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
B7B52E33475C580D9865317C008BD42E.text	B7B52E33475C580D9865317C008BD42E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla dusmeti (Navas 1914)	<div><p>Deleproctophylla dusmeti (Navás, 1914)</p><p>Figs 2 G, 2 H, 4 G, 4 H, 5 C, 5 D, 13, 14</p><p>Theleproctophylla dusmeti Navás, 1914: 57 (type locality: Spain: Madrid, Montarco; lectotype in MNCN). Navás 1915 a: 68 (Theleproctophylla). Navás 1915 b: 461 (Theleproctophylla). Bohigas and Sánchez 1917: 311 (Theleproctophylla). Navás 1920: 7 (Deleproctophylla). Escribano 1921: 365 (Theleproctophylla). Monserrat 1978: 182 (Deleproctophylla). H. Aspöck et al. 1980: 316 (with some questionable distribution, Deleproctophylla). Monserrat 1982: 70 (Deleproctophylla). H. Aspöck and Hölzel 1996: 78 (with some questionable distribution, Deleproctophylla). Sziráki 1998: 61 (with some questionable distribution, Deleproctophylla). H. Aspöck et al. 2001: 302 (Deleproctophylla); Whittington 2002: 377 (with some questionable distribution, Deleproctophylla). Canbulat and Kiyak 2004: 114 (probably misidentification, Deleproctophylla). Canbulat 2007: 36 (probably misidentification, Deleproctophylla). Dobosz and Ábrahám 2007: 24 (probably misidentification, Deleproctophylla). Koçak and Kemal 2012: 17 (probably misidentification, Deleproctophylla). Monserrat et al. 2012: 38 (Deleproctophylla). Monserrat 2013: 298 (Deleproctophylla). Monserrat et al. 2014: 153 (Deleproctophylla). Badano and Pantaleoni 2014: 301 (Deleproctophylla). Aistleitner 2019: 123 (with some questionable distribution, Deleproctophylla). Hassan et al. 2019: 513 (questionable distribution, Deleproctophylla). Aistleitner and Lencina 2020: 102 (Deleproctophylla). Monnerat and Ábrahám 2020: 144 (questionable distribution, Deleproctophylla). Hassan and Liu 2021: 416 (questionable distribution, Deleproctophylla). Poggi 2024: 9 (Deleproctophylla).</p><p>Examined material.</p><p>Type specimen. Paralectotype: • Granada, 8. VII. 01, Typus, Theleproctophylla dusmeti Navás SJ det., Theleproctophylla dusmeti Navás, 1914, Paralectotype, J. Legrand det, 1 ♀ (MNHN) (Fig. 24 A, B).</p><p>Other specimens.</p><p>• France: Alpes Maritimes, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.6767056&amp;materialsCitation.latitude=43.662247" title="Search Plazi for locations around (long 3.6767056/lat 43.662247)">Villeneuve Loubet</a>, 43°39'44.08"N, 3°40'36.14"E, 16. VII. 2011, D. Badano, 4 ♂ 5 ♀ (DB) ; • France: Bouches-du-Rhône, Eygalières, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=4.921583&amp;materialsCitation.latitude=43.74686" title="Search Plazi for locations around (long 4.921583/lat 43.74686)">Le Mas de Montfort</a>, m 170, 43°44'48.7"N, 04°55'17.7"E, 26. VI. 2019, 2 ♂ (DB) ; • France: Besse / Var, 250 m, 18. VII. 1964, Bilek, 1 ♂ 1 ♀ (HUAC) ; • France: Hérault, Gigean, 26–29. VI. 2014, A. Prost, 2 ♂ 1 ♀ (MDC) ; • France: Hérault, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.8759167&amp;materialsCitation.latitude=43.682995" title="Search Plazi for locations around (long 3.8759167/lat 43.682995)">Prades Le Lez</a>, 43°40'58.78"N, 3°52'33.30"E, 11. VII. 2011, D. Badano, 3 ♀ (DB) ; • France: Hérault, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=3.6767056&amp;materialsCitation.latitude=43.61004" title="Search Plazi for locations around (long 3.6767056/lat 43.61004)">St. Paul et Valmalle</a>, 43°36'36.14"N, 3°40'36.14"E, 26. VI. 2011, D. Badano, 2 ♂ 1 ♀ (DB) ; • same locality, 29. VI. 2011, D. Badano, 2 ♀ (DB); • same locality, 5. VII. 2011, D. Badano, 4 ♂ 1 ♀ (DB); • same locality, 10. VII. 2011, D. Badano, 3 ♂ 3 ♀ (DB); • France: Rochefort-du-Gard, 18. 07. 2002, J. Bard, 1 ♂ (SCM) ; • Portugal: Algarve, Odelouca, 14. VI. 1970, C. and A. Jeekel, 1 ♀ (RMNH) ; • Spain: Aragona, Sena, 17. VII. 1923, L. Navás, 1 ♂ 1 ♀ (MSNG) ; • Spain: Granada, 8. VII. 1901, 1 ♀ (MNHN) ; • Spain: Madrid, Coll. Lacroix, 1 ♂ 2 ♀ (MNHN) ; • Spain: Zaragoza, Pina de Ebro, 05. 07. 1991, V. Redondo, 1 ♂ (SCM) .</p><p>Diagnostic characters.</p><p>Metafemur with brown marking; forewing unmarked; hindwing subtriangular, posterior margin angled; hindwing with marking below pterostigma; male ectoproct with branch at mid-length; setae on branch only at tip; setae on branch and tip of ectoproct stout and curved.</p><p>Redescription.</p><p>Size (mm, based on 5 specimens). Head + thorax length 7.8–8.3; antenna length 14.1–16.5; fore wing length 19.7–23.8; fore wing width 5.6–6.4; hindwing length 15.3–18.4; hindwing width 4.7–5.8.</p><p>Head. Occiput yellowish brown; setae dense, long, pale yellow. Vertex yellowish brown; setae dense, long, pale yellow (Fig. 13 C). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 13 B). Mandible yellowish brown, dark brown apically. Labium and palp yellowish brown. Antenna with pedicel with dense pale and dark setae, flagellomeres brown, nodes slightly darker; club brown, paler anteriorly (Fig. 13 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum brown with yellowish brown markings: an anterior stripe-like pair, a median triangular pair, and a posterior rounded pair; mesoscutellum dark brown with a pair of yellowish rounded markings and posterior margin; setae pale yellow. Metanotum largely yellowish brown, medially dark brown; setae pale yellow (Fig. 13 C). Pleurae yellowish brown; setae pale yellow (Fig. 13 D).</p><p>Legs. Pro- and mesothoracic legs yellowish brown, tarsi slightly darker. Metathoracic leg yellowish brown, dorsal side of metafemur with dark brown marking. Setae of coxae pale yellow setae; setae of femora and tibiae thin and pale yellow dorsally, thicker and blackish ventrally; tibial spurs narrow; setae of tarsi blackish (Fig. 13 D).</p><p>Wings. Forewing long, apex rounded. Pterostigma with four to five forked and unforked brown veinlets; membrane brown. Presectoral area with five crossveins. RP with five branches. MA gently toward hind margin. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly pronounced. Forewing venation largely dark brown, paler toward wing base; Sc and R pale brown; anal veins pale brown. Forewing membrane hyaline (Figs 2 G, 2 H, 4 G, 4 H, 13 A). Hindwing broad, subtriangular, widest after RP fork; hind margin angled, gradually narrowing toward rounded apex. Presectoral area with three crossveins. RP with four branches. MA largely straight, slightly curved distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation dark brown, paler near wing base. Hindwing membrane hyaline, distal section with a rather large brown marking extending from pterostigma across to second branch of RP (Figs 2 G, 2 H, 4 G, 4 H, 13 A).</p><p>Abdomen. Tergites pale brown, with caudal pairs of yellowish markings. Sternites pale brown, paler than tergites. Pleural membrane brown with large paler markings.</p><p>Male genitalia. Ectoproct narrow, with branch at mid-length. Setae of ectoproct long, dark brown; setae on branch limited to tip, dense, black, stout, curved, and spine-like; setae only on tip of ectoproct, black, stout, curved, and spine-like (Fig. 14 A – C). Tergite 9 sub-rectangular in lateral view, with ventral projection; setae long and black, longer and denser on projection. Sternite 9 with paired lateral and a single median lobe; setae thin, spine-like on median lobe (Fig. 14 B, C). Gx 11 arch-shaped, projecting caudally and slightly curved downward in lateral view; gp 11 lightly sclerotized, forming a median furrow, with minute setae; gx 9 widely set apart, strongly sclerotized, dorsally folded (Fig. 14 F – J).</p><p>Female genitalia. Ectoproct elongated, projecting ventrally; setae long, black. Tergite 9 sub-rectangular. Gx 9 wider than long; setae long (Fig. 14 D, E). Tergite 8 subrectangular. Gx 8 wider than long, with long setae. Gp 8 projecting ventrally; setae thin (Fig. 14 D, E).</p><p>Larva. See Badano and Pantaleoni (2014) (Fig. 5 C, D).</p><p>Distribution.</p><p>Europe: France, Portugal, Spain (H. Aspöck et al. 1980; Rehfeldt 1989; H. Aspöck et al. 2001; Monserrat et al. 2014; Oswald 2025) (Fig. 25).</p><p>Comparative notes.</p><p>Deleproctophylla dusmeti may be difficult to distinguish from its two congeners in western Europe, D. australis and D. bleusei . The wing patterns traditionally used to differentiate these species are unreliable and exhibit high individual variation (Fig. 4). The diagnostic characters of D. dusmeti include a triangular-shaped hindwing (in contrast to the elongated hindwing of D. australis) and a male ectoproct branch that protrudes at mid-length (whereas in D. australis and D. bleusei the branch arises after mid-length) (Figs 13 A, 14 A).</p><p>Remarks.</p><p>D. dusmeti has been reported from several Asian countries, including Azerbaijan, Turkey, and Pakistan (H. Aspöck et al. 2001; Canbulat and Kiyak 2004; Canbulat 2007; Dobosz and Ábrahám 2007; Koçak and Kemal 2012; Hassan et al. 2019; Monnerat and Ábrahám 2020; Hassan and Liu 2021; Oswald 2025). However, the examination of the specimens assigned to the “ dusmeti - like form ” (see H. Aspöck et al. 1980) suggest that these instead belong to D. australis, representing individuals without forewing marking, a common trait in eastern populations of this species. The vague report from Pakistan (Hassan et al. 2019; Hassan and Liu 2021; Oswald 2025), which lacks any locality information cannot be verified at present. It may actually refer to D. dandizenor sp. nov., another species with unmarked forewing. The current observations suggest that D. dusmeti is exclusively a western European species. Deleproctophylla dusmeti is typically distinguished from D. australis by the absence of forewing spots. However, some populations along the southeastern coast of France include individuals with marked forewings mixed with unmarked ones (Fig. 4 H). The study of genital morphology and species delimitation analyses confirm that these specimens belong to D. dusmeti (Fig. 1). These spotted individuals are likely the source of records of D. australis in southern France, despite this species is absent from the French mainland (though it is present in Corsica). The distributions of D. dusmeti and D. australis do not overlap, being separated by a relatively small geographic gap along the northern Tyrrhenian coast of Italy (Fig. 25).</p></div>	https://treatment.plazi.org/id/B7B52E33475C580D9865317C008BD42E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
F401A582E6BF578888B5DC172056FE51.text	F401A582E6BF578888B5DC172056FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla gelini (Navas 1919)	<div><p>Deleproctophylla gelini (Navás, 1919)</p><p>Figs 3 A, 3 B, 15, 16</p><p>Theleproctophylla gelini Navás, 1919: 21 (type locality: Morocco: Safi; holotype in MNHN but lost at present). Navás 1930: 121 (Theleproctophylla). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). H. Aspöck et al. 2001: 302 (Deleproctophylla). Whittington 2002: 377 (Deleproctophylla). Ábrahám and Mészáros 2004: 333 (Deleproctophylla). Faucheux 2006: 204 (Deleproctophylla). Faucheux et al. 2012: 171 (Deleproctophylla). Ábrahám 2017: 126 (Deleproctophylla).</p><p>Examined material.</p><p>• Morocco: Agadir, 27. V – 6. VI. 1973, C. and A. Jeekel, 2 ♀ (RMNH) ; • Morocco: Bou Tazzert, Pres Mogadoa, 1928, R. Benoist. 2 ♂, 1 ♀, 2 ex. (MNHN) ; • Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.85725&amp;materialsCitation.latitude=30.708862" title="Search Plazi for locations around (long -9.85725/lat 30.708862)">Tamri</a>, 30°42'31.9"N, 09°51'26.1"W, 15 m, 30. 06. 2008, L. Ábrahám, L. Bognár and L. Nagy, 5 ♂♂ 1 ♀ (SCM) ; • Morocco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.85725&amp;materialsCitation.latitude=30.708862" title="Search Plazi for locations around (long -9.85725/lat 30.708862)">Tamri</a>, 30°42'31.9"N, 09°51'26.1"W, 15 m, 30. 06. 2008, L. Ábrahám, L. Bognár and L. Nagy, 1 ♂ 1 ♀ (USMB) ; • Morocco: Atlantic Plain, 4 km W <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-9.857278&amp;materialsCitation.latitude=30.708694" title="Search Plazi for locations around (long -9.857278/lat 30.708694)">Agoudi n’Aït Amar</a> 30°42'31.3"N, 9°51'26.2"W, 10 m, 27. 06. 2024, W. Szczepański, 4 ♂♂ 2 ♀♀ (USMB) ;</p><p>Diagnostic characters.</p><p>Vertex as long as wide; metafemur without brown marking; forewing extensively marked; hindwing broad, posterior margin curved; forewing with complex pattern: brown marking at base, subcostal area brown, brown marking at RP bifurcation, and large brown marking from pterostigma to first branch of RP; hindwing with a similar pattern: brown marking at base, subcostal area brown, brown marking at RP bifurcation, and large brown marking from pterostigma to posterior margin; male ectoproct bent inward, with branch at mid-length and protruding dorsally; setae on branch only at tip; setae on tip of ectoproct robust and relatively long; male with gx 11 straight, broad, incapsulating gx 9; gx 9 without apical fold; female ectoproct long; gx 8 and gp 8 not prominent.</p><p>Redescription.</p><p>Size (mm, based on 3 specimens). Head + thorax 7.4–9; antenna length 14.9–16.9; fore wing length 21–24.9; fore wing 6.2–7.3; hindwing 16.1–20.5; hindwing width 5.2–6.1.</p><p>Head. Vertex nearly as long as wide, yellowish brown; setae dense, long, pale yellow (Fig. 15 C). Para-ocular band pale yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale (Fig. 15 B). Mandible yellowish brown, dark brown apically. Labium and palp yellowish brown. Antenna with pedicel with dense pale and dark setae, flagellomeres pale brown, nodes dark brown; club dark brown, paler anteriorly (Fig. 15 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum yellowish brown gradually darker medially and along sclerites margins. Metanotum yellowish brown, medially darker; setae pale yellow (Fig. 15 C). Pleurae yellowish brown; setae pale yellow (Fig. 15 D).</p><p>Legs. Pro- and mesothoracic legs yellowish brown, including tarsi. Metathoracic leg yellowish brown, unmarked. Setae of coxae pale yellow setae; setae of femora and tibiae thin and pale yellow dorsally, thicker and blackish ventrally; tibial spurs narrow; setae of tarsi blackish (Fig. 15 D).</p><p>Wings. Forewing broad, apex rounded. Pterostigma with ~ 5 forked and unforked brown veinlets; membrane brown. Presectoral area with five crossveins. RP with ~ 5 branches. MA gently toward hind margin. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly pronounced. Forewing venation largely pale brown. Forewing membrane largely hyaline with a complex brown pattern: base of wing brown, apical costal crossveins with brown shade at base, subcostal area brown, brown marking at RP branching, apical portion of wing with a large brown marking extending from subcostal area and pterostigma to first branch of RP (Figs 3 A, 3 B, 15 A). Hindwing broad, widest at RP fork; hind margin rounded, smoothly curved toward rounded apex. Presectoral area with three crossveins. RP with five branches. MA largely straight, slightly curved distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation pale brown, Hindwing membrane hyaline, with a complex brown pattern: base of wing brown, apical costal crossveins with brown shade at base, subcostal area brown, brown marking at RP branching, apical portion of wing with a large brown marking extending from subcostal area and pterostigma to posterior margin of wing (Figs 3 A, 3 B, 15 A).</p><p>Abdomen. Tergites pale brown. Sternites pale brown, paler than tergites. Pleural membrane brown.</p><p>Male genitalia. Ectoproct robust, strongly bent inward, with branch slightly beyond mid-length. Branch projecting dorsally, broad, and relatively thick, progressively widening at apex. Apex of ectoproct club-shaped expanding internally. Setae of ectoproct short, dark brown; setae on branch sparse, those on apical section, black, stout, and curved; setae on tip of ectoproct progressively denser caudally, black, stout, curved, and spine-like on the internal side (Fig. 16 A – C). Tergite 9 trapezoid in lateral view, with broad ventral projection; setae long and black, longer and denser on projection (Fig. 16 B, C). Sternite 9 without paired lateral lobes, being composed only by the median lobe; setae thin, spine-like lobe. Gx 11 arch-shaped, projecting caudally, broad and incapsulating gx 9, straight in lateral view; gp 11 lightly sclerotized, elongated, with a shallow median furrow, with minute setae; gx 9 widely set apart, strongly sclerotized, flattened, without dorsal fold (Fig. 16 F – J).</p><p>Female genitalia. Ectoproct strongly elongated, projecting ventrally; setae long, black (Fig. 16 D, E). Tergite 9 sub-rectangular. Gx 9 wider than long; setae long. Tergite 8 subrectangular. Gx 8 not prominent, with long setae. Gp 8 not prominent; setae thin. Sternite 7 with sparse stout setae (Fig. 16 D, E).</p><p>Distribution.</p><p>Africa: Morocco (H. Aspöck et al. 2001; Ábrahám and Mészáros 2004; Oswald 2025).</p><p>Comparative notes.</p><p>Among all Deleproctophylla species, D. gelini is the easiest to identify due to its unique fore- and hindwing pattern (Ábrahám and Mészáros 2004) (Fig. 3 A, B). Additionally, this species differs from all congeners in both male and female genitalia, which are characterized by several distinctive characters (Ábrahám and Mészáros 2004) (Fig. 16). The male ectoproct is stout and distinctly bent inward and the branch arises dorso-caudally at mid-length. The branch stacks on the ectoproct, instead of being directed inward, as seen in all other species. Furthermore, the sternite 9 of the male lacks the lateral lobes found in other species (Fig. 16 A – C). The gx 9 + 11 complex is also different being, shaped like a capsule. The Gx 11 is broad, not curved downward, and encases the gx 9, which lacks the apical dorsal fold present in all other species (Fig. 16 F – J). The female of D. gelini is characterized by an elongated ectoproct projecting ventrally and reduced gx 8 (Fig. 16 D, E).</p></div>	https://treatment.plazi.org/id/F401A582E6BF578888B5DC172056FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
BEA9AA02FEDA5955B3495403B133D1E3.text	BEA9AA02FEDA5955B3495403B133D1E3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla Lefebvre 1842	<div><p>Deleproctophylla Lefèbvre, 1842</p><p>Figs 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24</p><p>Deleproctophylla Lefèbvre, 1842: 6 (emended to “ Theleproctophylla ” by Rambur, 1842: 351). Type species: Ascalaphus australis Fabricius, 1893, by subsequent designation by van der Weele (1908).</p><p>Diagnosis.</p><p>Adult. Head breadth at widest point, including compound eyes, slightly larger to width of mesothorax at wing base. Compound eye with deep median furrow, lateral margins slightly indented at furrow. Antenna not reaching pterostigma; club pyriform, wide. Forewing elongate; hindwing broadest at RP fork. Forewing membrane largely hyaline, with brown markings in a few species; hindwing membrane largely hyaline with brown markings or infuscations in apical half. Abdomen without pleuritocavae.</p><p>Male genitalia. Ectoproct with prominent “ pincer-like ” ventral projection, with bifurcating cylindrical branch in distal half, apex bent downward and inward; branch and apex with robust, spiniform setae. Tergite 9 sub-rectangular in lateral view, with prominent ventral thumb-shaped projection. Sternite 9 with caudal paired lateral lobes and a median, prominent sub-rectangular lobe; setae on median lobe spine-like. Complex of gx 9 + 11 proportionally small in comparison to tip of abdomen; gx 11 arch-shaped, projecting caudally and slightly curved downward in lateral view; gp 11 forming a furrow, with short setae; gx 9 widely set apart, with a dorso-apical, upward fold, resembling a smooth hook in lateral view.</p><p>Female genitalia. Ectoproct elongate, with short ventral projection. Tergite 9 subrectangular. Gx 9 wider than long. Tergite 8 subrectangular. Gx 8 distinct, not fused and widely set apart, wider than long, slightly prominent. Gp 8 caudally prominent, appearing subtriangular in lateral view (except for D. gelini).</p><p>Larva. Head capsule not dilated posteriorly; ocular tubercle cylindrical with small apical protuberance; mandible with three teeth, median tooth largest and positioned closer to apical than to basal tooth; meso- and metathorax each with two pairs of sub-equal cylindrical scolus-like processus; anterior abdominal spiracle lateral; dorsal abdominal series of spiracles composed of eight cylindrical scolus-like processus, ventral series composed of two anterior scolus-like processus and six tubercle-like processus; sternite 8 with short odontoid processus; sternite 9 with short rastra with four digging setae; body dolichasters elongated, stick-shaped.</p><p>Comparative notes.</p><p>Potential synapomorphies of Deleproctophylla include: i) male ectoproct with caudal projection and median bifurcating branch, ii) male gx 9 widely separated from each other and from gp 11, with apical fold; iii) female ectoproct with caudal projection. Deleproctophylla is one of the few genera of Ascalaphini characterized by prominent male ectoproct with median branch, such as Bubopsis McLachlan, 1898, and Phalascusa Kolbe, 1897 (Tjeder 1992). However, in Deleproctophylla species, the ectoproct branch develops inward or dorsally (i. e., D. gelini), while in the sympatric species of Bubopsis, the branch extends basally and ventrally. These traits, combined with the characteristic pigmentation pattern of Deleproctophylla, allow easy identification from other ascalaphid genera with elongated ectoproct. In addition, to the obvious coloration differences, Deleproctophylla is immediately set apart from Libelloides in the shape of the male ectoproct, which is proportionally shorter, simpler in shape and lacking branch in the latter. However, these two genera share a similar conformation of the gx 9 + 11 complex, especially in the shape of gx 9. The larvae of these two genera are also very similar in body shape, tooth arrangement, and the position, development, and shape of thoracic and abdominal series of scolus-like processus. The larvae of Deleproctophylla can be distinguished by those of Libelloides in the head capsule, which is not dilated posteriorly and the dolichasters covering the body, which are mostly stick-shaped and do not expand apically into a wide rosette (Badano and Pantaleoni 2014).</p><p>Remarks.</p><p>One of the most unusual features of Deleproctophylla is the presence of the so-called phylla, large, paired, membranous, leaf-shaped structures protruding from the female abdomen (Figs 2 F, 3 F) (H. Aspöck et al. 1980; Tjeder 1992). The genus name derives from this trait. The phylla consist of the male spermatophore, which remains attached to the female genital opening after mating but is shed when she lays eggs. The evolutionary significance of this prominent structure remains unclear, but it may serve to reduce competition between males by preventing remating with an already fertilized female. In a few other ascalaphid genera, such as Bubopsis McLachlan, females also retain the spermatophore after mating, but in none of them are these structures as large and striking as in Deleproctophylla .</p></div>	https://treatment.plazi.org/id/BEA9AA02FEDA5955B3495403B133D1E3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
67F1EA64EACE5BEF8317C0FA5BB2A1AA.text	67F1EA64EACE5BEF8317C0FA5BB2A1AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla tengri Zheng, Badano, H. Aspock & Liu 2026	<div><p>Deleproctophylla tengri Zheng, Badano, H. Aspöck &amp; Liu sp. nov.</p><p>Figs 3 C, 3 D, 17–21</p><p>Type material.</p><p>Holotype • China: Xinjiang, Shihezi [石河子], Maangou Village [马鞍沟村], Mt. Jiangjunshan [将军山], 650 m, 6. VI. 2020, Haitian Song, 1 ♂ (IZCAS) . Paratypes • Same locality as holotype, 18. VII. 2022, Rui Wang, 1 ♀ (IZCAS); • same locality as holotype, 14. VII. 2025, Yuchen Zheng and Rui Wang, 1 ♂ 1 ♀ (CAU); • Kazakhstan: Alma-Ata (present Almaty), 1 ♀ (NHMW) ; • Kyrgyzstan: Narinskaya Obl. NE Naryn, Aral, 1500 m, 10. VII. 2000, Vladimir Dolin, 1 ♂ (HUAC) ; • Kyrgyzstan: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=74.872635&amp;materialsCitation.latitude=41.426018" title="Search Plazi for locations around (long 74.872635/lat 41.426018)">5 km NE of Ugut</a>, 41°25.561'N, 74°52.358'E, 1616 m, 09. 07. 2013, L. Ábrahám and S. Ilniczky, 1 ♂ 2 ♀♀ (SCM) ; • Kyrgyzstan: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.21455&amp;materialsCitation.latitude=41.444485" title="Search Plazi for locations around (long 72.21455/lat 41.444485)">2 km S of Kala</a>, 41°26.669'N, 72°12.873'E, 789 m, 03. 07. 2013, L. Ábrahám and S. Ilniczky, 6 ♀♀ (SCM) ; • Kyrgyzstan: Jalal-Abad Region, ca 7 km N of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=72.25&amp;materialsCitation.latitude=41.416668" title="Search Plazi for locations around (long 72.25/lat 41.416668)">Tash-Kömür</a>, 41°25'N, 72°15'E, 660 m, 20–21.06. 2003, R. Dobosz, 1 ♀ (Phot. 1) (USMB) ; • Tajikistan: Gorno-Badakhshan, S. side Saghirdasht Pass 10 km N. of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.800446&amp;materialsCitation.latitude=38.547695" title="Search Plazi for locations around (long 70.800446/lat 38.547695)">Kala-i Khumb</a>, 38°32'51.7"N, 70°48'1.6"E, 1850 m, T. and W. Garrevoet, 1 ♀ (RMNH) .</p><p>Larval material.</p><p>China: Xinjiang, Shihezi, 5 km south of the Shihezi City, 686 m, 20. V. 2025, Ziyang Ni (ZCAU).</p><p>Diagnostic characters.</p><p>Metafemur unmarked; forewing with acute apex, hyaline, without marking; hindwing subtriangular, posterior margin angled; hindwing marking generally as shade along crossveins, extending on RP; male ectoproct median branch basally wide, gradually bifurcating, subtriangular shape.</p><p>Description.</p><p>Size (mm, based on 2 specimens). Head + thorax length 6.9; antenna length 11.22–14.15; fore wing length 20.1–21.2; fore wing width 6.1–6.5; hindwing length 15–16; hindwing width 5.7–6.2.</p><p>Head. Vertex dark brown; setae dense, long, pale (Fig. 17 C). Para-ocular band pale yellow. Frons yellow, medially with a dark brown spot; setae dense, long, pale. Clypeus and labrum yellow; setae long, pale (Fig. 17 B). Mandible yellow, transitioning to dark brown apically. Labium and palp yellow. Antenna pedicel with dense pale and dark setae, flagellomeres generally dark brown; club basally dark brown, gradually transitioned to pale yellow to the tip (Fig. 17 A).</p><p>Thorax. Cervical sclerite yellow, with a pair of dark markings. Pronotum black, lateral margin yellow; setae long, pale. Mesonotum black with yellow markings: an anterior transversal pair, a median triangular pair and two lateral pairs; mesoscutellum mostly black, medially with a pair of yellowish markings, posterior margin yellow (Fig. 17 C). Pleurae pale yellow (Fig. 17 D).</p><p>Legs. Pro- and mesothoracic legs pale yellow, tarsi each generally pale yellow and distally dark brown; hind tarsus mostly dark brown. Coxae with thin pale setae; femora and tibiae with thin pale setae and ventral thicker blackish setae; tibial spurs narrow, blackish; tarsi with robust blackish setae (Fig. 17 D).</p><p>Wings. Forewing long, apex acute. Pterostigma with three to four forked and unforked brown veinlets; membrane brown. Presectoral area with five to six crossveins. RP with four branches. MA gradually curving toward hind margin in distal portion. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly pronounced. Forewing venation largely brown, paler toward wing base; Sc pale yellow, paler toward distal part; R pale yellow, darker toward distal part. Forewing membrane hyaline (Figs 3 C, 3 D, 17 A). Hindwing subtriangular, broadest at RP fork; hind margin angular curved narrowing toward rounded apex. Presectoral area with two crossveins. RP with three branches. MA largely straight, slightly curved toward hind margin distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation brown, paler near wing base and distally. Hindwing membrane hyaline, hindwing marking generally as shade along crossveins, extending on RP (Figs 3 C, 3 D, 17 A).</p><p>Abdomen. Tergites black, each with a caudal pair of yellowish markings. Sternites yellow. Pleural membrane brown with large paler markings.</p><p>Male genitalia. Ectoproct narrow, with branch in apical half, basally wide, gradually bifurcating, subtriangular shape. Setae of ectoproct long, dark brown; setae on branch limited to tip, dense, black, stout, curved, and spine-like; setae only on tip of ectoproct, black, stout, curved, and spine-like (Fig. 18 A – C). Tergite 9 sub-rectangular in lateral view, with ventral projection; setae long and black, longer and denser on projection. Sternite 9 wide trapezoid in ventral view; setae thin, spine-like on median part (Fig. 18 B, C). Gx 11 arch-shaped, external tip rounded, slightly curved downward in lateral view; gp 11 lightly sclerotized forming a shallow median furrow, with short setae; gx 9 strongly sclerotized, with upward fold, widely hook-like in lateral view (Fig. 18 F – J).</p><p>Female genitalia. Ectoproct elongated; setae long, black. Tergite 9 sub-rectangular. Gx 9 longer than wide; setae long. Tergite 8 subrectangular. Gx 8 wider than long, with long setae. Gp 8 flatted; setae thin (Fig. 18 D, E).</p><p>Diagnosis of 3 rd instar larva.</p><p>Dorsal head yellowish brown, with many dark brown spots, spots on the middle part relatively sparse, anteromedially with a dark brown marking. Dorsal segments 1–7 medially with a pair of short dark brown stripes, laterally scattered with irregular dense dark brown markings (Fig. 20).</p><p>Description of 3 rd instar larva.</p><p>Size (mm). Body length (excluding mandible): 15.74; head length: 3.83; head width: 4.07; mandible length: 4.06.</p><p>Head. Wider then long, laterally with short setae (Fig. 21 A, B). Clypeo-labrum generally brown, concave internally; anterior margin covered with some dolichasters. Dorsal side of head capsule yellowish brown, with many dark brown spots, spots on the middle part relatively sparse, anteromedially with a dark brown marking; head ventrally yellowish brown, with many short dolichasters, pore developed. Ocular tubercle prominent and large, apical part with a tiny projection. Antennae brown (Fig. 21 A, B). Mandible nearly as long as head width; dorsal mandible basally pale brown, gradually darkening to the distal part; ventral mandible generally dark brown. Mandible basally with three or four pseudo-teeth and three interdental mandibular setae; second tooth largest and close to the third tooth; first tooth longer than third tooth; two pseudo-teeth between first and second teeth (Fig. 21 A, B).</p><p>Thorax. Laterally with short setae. Pronotum generally pale pink, laterally pale yellow, scattered with some dark dots, posterolaterally with a pair of black markings, covered with tiny dark dolichaster. Mesothorax generally pale yellow, with spiracles present on short stout black sclerotized tubercle, lateral margin dark brown. Meso- and metathoracic setiferous processus scolus-like, distally constrict. Anterior mesothoracic setiferous processus longer than the posterior pair (Fig. 21 C).</p><p>Legs. Pale yellowish brown, densely covered by short setae.</p><p>Abdomen. Pale yellow. Dorsal segments 1–7 medially with a pair of short dark brown stripes, laterally scattered with irregular dense dark brown markings. Each ventral segments 1–7 pale, laterally scattered with many dark spots, ventral segments 2–7 medially with a dark spot. Each segment 1–7 with a pair of dorsal scolus-like processus and a pair of ventral short scolus-like processus. Abdominal segment 9 pale yellow, long trapezoid, twice as long as wide, developed rastra with four pairs of digging setae (Fig. 21 D).</p><p>Distribution.</p><p>Asia: China (Xinjiang); Kazakhstan (Almaty Region); Kyrgyzstan (Jalalabad and Narinsky Region); Tajikistan (Gorno-Badakhshan Autonomous Region) (Fig. 25).</p><p>Etymology.</p><p>The species name tengri is derived from the Turkic word “ tengri, ” meaning sky or heaven. The species was found in the surrounding area of the Tianshan Mountains (also known as the Tengri Tagh Mountains), a mountain range whose name, across various language families, signifies “ Mountains of sky or heaven ”. Thus, the name tengri reflects both the geographical origin of the species and the celestial meaning of the term in Turkic languages. The species epitheton is a substantive and an apposition to the genus.</p><p>Comparative notes.</p><p>Deleproctophylla tengri can be readily distinguished from the congeners by the median branch of the male ectoproct is subtriangular and distinctly broad at the base (Fig. 18 A – C). The larva of D. tengri sp. nov. exhibits all the diagnostic larval features of the genus and differs only slightly from the known congeners D. australis and D. dusmeti (Figs 5, 20, 21). The main distinguishing feature of D. tengri compared to its western counterparts is the paler head capsule, which is distinctly mottled dark brown dorsally and paler ventrally (Fig. 21 A, B). In contrast, the larvae of D. australis and D. dusmeti, which are virtually indistinguishable from each other, are characterized by a uniformly dark head capsule (Fig. 5).</p><p>Biology.</p><p>Deleproctophylla tengri inhabits arid hills (Fig. 19 A, B). The larva of this new species was observed preying on a Parnassius apollonius Eversmann, 1847 ( Lepidoptera: Papilionidae) on a hillside meadow (Fig. 19 C).</p><p>Remarks.</p><p>This new species is endemic to the surrounding area of the Tianshan Mountains. We suspect that the specimens of D. variegata from Kyrgyzstan examined by Krivokhatsky et al. (2015) (without figure) are probably conspecific with this new species, considering that the record from Kyrgyzstan belongs to the same fauna as this new species (we also designated a paratype from Kyrgyzstan). Furthermore, Krivokhatsky et al. (2015) and Kerimova et al. (2023) also mentioned that D. variegata occurs in Uzbekistan and Kazakhstan, which might represent the same situation mentioned above. Nikolajev (2006) wrongly reported D. variegata from Takuma sands from the Ili River Valley (Almaty Region). The picture included in the publication confirms that the record most likely refers to D. tengri .</p></div>	https://treatment.plazi.org/id/67F1EA64EACE5BEF8317C0FA5BB2A1AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
0BD298CE074B5AAE8E4458002D7EDDB0.text	0BD298CE074B5AAE8E4458002D7EDDB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deleproctophylla variegata (Klug 1834)	<div><p>Deleproctophylla variegata (Klug in Ehrenberg, 1834)</p><p>Figs 3 E, 3 F, 22, 23, 24 C, 24 D</p><p>Ascalaphus variegatus Klug in Ehrenberg, 1834: 36 (type locality: northern Lebanon; holotype in MfN). Hagen 1860: 53 (Ascalaphus). Hagen 1866: 387 (Ascalaphus). McLachlan 1873: 260 (Ascalaphus). van der Weele 1904: 98 (Theleproctophylla). van der Weele 1908: 287 (Deleproctophylla). Navás 1909: 54 (misidentification, Theleproctophylla). Navás 1911: 92 (misidentification, Theleproctophylla). Navás 1912 a: 124 (misidentification, Theleproctophylla). Navás 1925: 31 (misidentification, Theleproctophylla). Alexandrov-Martynov 1926: 199 (probably misidentification of D. tengri, Ascalaphus). Scott 1929: 80 (Deleproctophylla). Navás 1929: 14 (misidentification, Theleproctophylla). H. Aspöck et al. 1980: 317 (Deleproctophylla). H. Aspöck and Hölzel 1996: 78 (Deleproctophylla). Sziráki 1998: 61 (Deleproctophylla). Sziráki 2000: 91 (Deleproctophylla). Santos 2000: 282 (Theleproctophylla). H. Aspöck et al. 2001: 302 (Deleproctophylla). Whittington 2002: 377 (Deleproctophylla). Koçak and Kemal 2002: 7 (Deleproctophylla). Canbulat 2007: 36 (Deleproctophylla). Dobosz and Ábrahám 2007: 19 (Deleproctophylla). Koçak and Kemal 2012: 17 (Deleproctophylla). Ilyina et al. 2013: 35 (Deleproctophylla). Krivokhatsky et al. 2015: 814 (probably misidentification of D. tengri, Deleproctophylla). Aistleitner 2019: 124 (Deleproctophylla). Monnerat and Ábrahám 2020: 144 (Deleproctophylla). Kerimova et al. 2023: 132 (Deleproctophylla). Poggi 2024: 10 (Deleproctophylla).</p><p>Examined material.</p><p>Type material. Lectotype • Smyrna Leonis col., 129, Type, Syria Ehren., variegatus Klug, variegatus Kl, Theleproctophylla variegata Kl. Soldaniky det. (MfN) (Fig. 24 C, D).</p><p>Other specimens.</p><p>• Cyprus: Umg. v. Paphos ca 350 m, 1980 (27. V – 8. VI), H. u. L. Hölzel, 27 ♂ 27 ♀ (NHMW) ; • Cyprus: 6 km Z v. Kyrenia, 300 m, 15. VI. 1971, M. J. and J. P. Duffels, 1 ♀ (RMNH); • Cyprus: Larnika Salt Lake, 1–16. VI. 1982, M. C. and G. Kruseman, 2 ex (RMNH) ; • Cyprus: Mamonia, N of Kouklia, 7–14 and 18. VI. 1982, M. C. and G. Kruseman, 1 ♀ (RMNH) ; • Greece: Rhodes, Mount Profitis Ilias, 22. VII. 2009, 1 ♂ 1 ♀ D. Badano, (DB) ; • Iran: Lorestan, Zagros Mts. near Borujerd, Vil. Vaenna’Ialt. 2400–3000 m, 29. VII. 2014, E. Rutjan, 1 ♀ (DB) ; • Iran: Tehran, Qolhak, 1700 m, 1961. VI. 1923, J. Klapperich, 1 ♂ 1 ♀ (HUAC) ; • Iran: N Iran, Abyek, Loc. No. 30, 24.06. [19] 70, Exp. Nat. Mus. Praha, 1 ♂ (NMPC) ; • Iran: S Iran, 30 km E Kazerun, 1300 m, Loc. No. 229, 08–10.06. 1973, Exp. Nat. Mus. Praha, 1 ♀ (NMPC) ; • Iran: S Iran, 28 km N Masiri, 1650 m, Loc. No. 236, 12. 06. 1973, Exp. Nat. Mus. Praha, 2 ♀♀ (NMPC) ; • Iran: Qolhak, B. Tehran, 1700 m, 09. 06. 1961, J. Klapperich, 1 ♂ (SCM) ; • Iran: Qolhak, B. Tehran, 1700 m, 23. 06. 1961, J. Klapperich, 1 ♀ (SCM) ; • Iran: Province Fars, Persepolis, Zagros Mts., 1200 m, 25–26.05. 1999, Hácz-Kőszegi, 1 ♂ (SCM) ; • Iran: Province Lorestan, Ostoran Mt., 17 km E from Dorud, 2300–2700 m, 22. 06. 2000, Gy. Fábián, L. Szécsényi and K. Székely, 1 ♀ (SCM) ; • Iran: Province Hamadan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.3769&amp;materialsCitation.latitude=34.045933" title="Search Plazi for locations around (long 48.3769/lat 34.045933)">Nahavand</a> 34°02.756'N, 48°22.614'E, 1851 m, 26. 06. 2005, L. Ábrahám, 4 ♂♂ 14 ♀♀ (SCM) ; • Iran: Province Fars, Kuh-e Kum Mts., Hasanabad vill. env., 2100 m, 27. 05. 2000, M. Kalabza, 1 ♀ (SCM) ; • Iran: E Téhrán, 2200 m, 18. 05. 2000,?, 1 ♀ (SCM); • Iran: Province Hamadan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=48.3769&amp;materialsCitation.latitude=34.045933" title="Search Plazi for locations around (long 48.3769/lat 34.045933)">Nahavand</a> 34°02.756'N, 48°22.614'E, 1851 m, 26. 06. 2005, L. Ábrahám, 1 ♀ (USMB) ; • Iran: Province Fars, S of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=51.53389&amp;materialsCitation.latitude=30.210556" title="Search Plazi for locations around (long 51.53389/lat 30.210556)">Nüräbäd</a>, 30°12'38"N, 51°32'02"E, 900 m, 29. 04. 2016, netting, R. Królik, 1 ♀ (USMB) ; • Israel: Tabgha 10 km N. v. Tiberias, 25. V. 1967, C. A. W. Jeekel, 1 ♂ 2 ♀ (RMNH) ; • Jordan: Oberes Jordantal, 06.1999, G. Müller, 1 ♂ (SCM) ; • Syria: Damascus, Mts. Hermon, 1 km E <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.027267&amp;materialsCitation.latitude=33.471016" title="Search Plazi for locations around (long 36.027267/lat 33.471016)">Burqus</a>, 33°28.261'N, 36°01.636'E, 1140 m, 12. 06. 2006, N. Rahmé, A. Kotán, A. Márkus, D. Szalóki and K. Székely, 1 ♀ (SCM) ; • Turkey: Anatolia orient., vil. Elâzıg ˘, valico Gözeli, m 900, 24. VI. 1968, V. Sbordoni, 1 ♂ (MNSG) ; • Turkey: Bingöl, 38°55'11.3"N, 40°20'8.3"E, T. and W. Garrevoet and N. Vandorpe, 1 ♀ (RMNH); • Turkey: Bitlis, Kuzkun Kiran Gecidi, 2100 m, 28. VII. 1992, J. A. W. Lucas, 1 ♀ (RMNH) ; • Turkey: Hakkâri, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.372166&amp;materialsCitation.latitude=37.49714" title="Search Plazi for locations around (long 43.372166/lat 37.49714)">Süvarihalil Geçidi</a>, 1800–2400 m, 37°29'49.7"N, 43°22'19.8"E, 11. VII. 2004, Dils J. and Faes J., 4 ♀ (RMNH) ; • Turkey: İçel (present Mersin), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.164448&amp;materialsCitation.latitude=38.698334" title="Search Plazi for locations around (long 34.164448/lat 38.698334)">Güzeloluk</a>, 950 m, 38°41'54"N, 34°09'52"E, 15. VI. 1999, Dils J. and Faes J., 1 ♂ 3 ♀ (RMNH) ; • Turkey: Izmir, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.821112&amp;materialsCitation.latitude=38.65917" title="Search Plazi for locations around (long 26.821112/lat 38.65917)">Foça</a>, 50 m, 38°39'33"N, 26°49'16"E, 9. VI. 1999, Dils J. and Faes J., 1 ♂ 1 ♀ (RMNH) ; • Turkey: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.064445&amp;materialsCitation.latitude=38.275276" title="Search Plazi for locations around (long 43.064445/lat 38.275276)">Gevaş</a> 1870 m, 38°16'31"N, 43°03'52"E, 16. VII. 2006, Dils J. and Faes J., 2 ♀ (RMNH) ; • Turkey: Mut, VI – VII. 1966, H. H. F. Hamann, 1 ♂ 1 ♀ (HUAC) ; • Turkey: Mut, 5–8. VI. 1968, Josef Schmidt, 1 ♀ (HUAC) ; • Turkey: Mut, 8. VI. 1968, Jos. Schmidt, 1 ♂ (HUAC) ; • Turkey: Mut, 12. VI. 1965, Jos. Schmidt, 4 ♀ (HUAC) ; • Turkey: Urfa, 30–31. V. 1978, J. Schmidt, 1 ♂ 1 ♀ (HUAC); • Turkey: Urfa, 10. VI. 1988, J. Schmidt 1 ♂ 1 ♀ (HUAC); • Turkey: Province Agri, 7 km W of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=42.5&amp;materialsCitation.latitude=39.816666" title="Search Plazi for locations around (long 42.5/lat 39.816666)">Aydintepe</a>, 39°49'N, 42°30'E, 1800 m, 05–06.08. 1988, Gyulai, Hreblay, Ronkay and Ronkay, 2 ♂♂ (SCM) ; • Turkey: Kusgunkiran Gecidi, 12. 07. 1989,?, 1 ♀ (SCM) ; • Turkey: mer., village Icel Orta Toroslar, Hocali, 500 m, 03–04.06. 2004, Z. Rahmé, L. Nádai and K. Székely, 2 ♀♀ (SCM) ; • Turkey: Province Mersin, 10 km E of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.566666&amp;materialsCitation.latitude=37.133335" title="Search Plazi for locations around (long 36.566666/lat 37.133335)">Aydincik</a>, 37°08'N, 36°34'E, in meadows, 01. 06. 2001, R. Dobosz, 3 ♂♂ 4 ♀♀ (USMB) ; • Turkey: Province Mersin, 10 km E of Aydincik, 37°08'N, 36°34'E, in meadows, 01. 06. 2001, J. Kurzawa, 1 ♂ 2 ♀♀ (USMB); • Turkey: Province Mersin, 3 km N of Erdemli, 36°37'N, 34°19'E, oak woods, 350 m, A. Lasoń, 1 ♀ (USMB); • Turkey: Province Mersin, 3 km N of Erdemli, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=34.25&amp;materialsCitation.latitude=36.616665" title="Search Plazi for locations around (long 34.25/lat 36.616665)">Taşeli Platosu</a>, 36°37'N, 34°15'E, 05. 06. 2011, A. Lasoń, 1 ♂ (USMB) ; • Turkey: Province Şanliurfa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.883335&amp;materialsCitation.latitude=37.233334" title="Search Plazi for locations around (long 37.883335/lat 37.233334)">Yeni Halfeti</a>, 37°14'N, 37°53'E, 08. 06. 2011, A. Lasoń, 1 ♂ (USMB) ; • Turkey: Province Şanliurfa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.883335&amp;materialsCitation.latitude=37.233334" title="Search Plazi for locations around (long 37.883335/lat 37.233334)">Yeni Halfeti</a>, 37°14'N, 37°53'E, 08. 06. 2011, R. Królik, 1 ♂ 1 ♀ (USMB) ; • Turkmenistan: Turkmenia SW, КАЛА-Kala, V. 1993, [S.] Naglis, 1 ♂ (ex coll. Gallo, 2000), det. Badano, 2012 (MSNG) .</p><p>Diagnostic characters.</p><p>Forewing unmarked; hindwing posterior margin curved, convex; hindwing with faint marking below pterostigma, usually just a shade around crossveins; male ectoproct with branch stout, in apical third, well after mid-length, after branch ectoproct narrowing and progressively broadening caudally; setae on branch for its entire length; setae on branch and tip of ectoproct long and thin.</p><p>Redescription.</p><p>Size (mm, based on 5 specimens). Head + thorax length 6.01–7.04; antenna length 10.83–12.78; fore wing length 16.30–17.87; fore wing width 4.14–5.38; hindwing length 10.61–13.02; hindwing width 3.84–4.78.</p><p>Head. Vertex yellowish brown; setae dense, long, pale yellow (Fig. 22 C). Para-ocular band yellow. Frons yellowish brown; setae dense, long, pale yellow. Clypeus and labrum yellowish brown; setae long, pale. Mandible yellowish brown, darker apically. Labium and palp yellowish brown, of the same color of mandible (Fig. 22 B). Antenna with pedicel with dense pale and dark setae, flagellomeres brown, nodes slightly darker; club brown, paler anteriorly (Fig. 22 A).</p><p>Thorax. Cervical sclerite yellowish brown. Pronotum brown; setae long, pale yellow. Mesonotum brown with yellowish brown markings: an anterior stripe-like pair, a median triangular pair, and a posterior rounded pair; mesoscutellum dark brown, with yellowish brown posterior margins. Metanotum yellowish brown, with dark brown median marking; setae pale yellow (Fig. 22 B). Pleurae yellowish; setae pale yellow (Fig. 22 D).</p><p>Legs. Pro- and mesothoracic legs entirely yellowish brown. Metathoracic leg yellowish brown, usually unmarked, occasionally with a faint brown marking. Setae of coxae pale yellow setae of femora and tibiae thin and pale yellow dorsally, thicker and blackish ventrally; tibial spurs narrow; setae of tarsi blackish (Fig. 22 D).</p><p>Wings. Forewing long, apex rounded. Pterostigma with ~ 4 unforked brown veinlets; membrane brown. Presectoral area with ~ 6 crossveins. RP with four branches. MA gradually curving toward hind margin in distal portion. MP - CuA area with ~ 3 irregular rows of cells. Anal area hind margin straight; axillary lobe slightly prominent. Forewing venation brown, paler toward wing base and at tip; Sc and R yellowish. Forewing membrane hyaline, unmarked. Hindwing broad, wider at RP fork; hind margin markedly convex, with a pronounced curve. Presectoral area with three crossveins. RP with three branches. MA largely straight, slightly curved toward hind margin distally. MP - CuA area with ~ 2 irregular rows of cells. Hindwing venation brown, paler near wing base; Sc yellowish (Fig. 3 E, F, 22 A). Hindwing membrane hyaline, distal section with an inconspicuous brown marking, as a shade bordering crossveins from pterostigma to first branch of RP (Fig. 3 E, F, 22 A).</p><p>Abdomen. Tergites pale brown, each with a caudal pair of yellowish markings. Sternites pale brown, paler than tergites. Pleural membrane pale brown.</p><p>Male genitalia. Ectoproct with branch bifurcating at two-third of length; ectoproct abruptly narrowing after branching and gradually expanding caudally. Setae of ectoproct long, dark brown; setae of branch arranged on entire length, sparse and gradually denser toward apex, black, long, and slightly curved; setae on tip of ectoproct, black, long, and slightly curved, denser apically (Fig. 23 A – C). Tergite 9 sub-rectangular in lateral view, with large ventral thumb-like projection; setae long and black, longer and denser on projection. Sternite 9 with three lobes, lateral pair slightly prominent, median pair large; setae thin, thicker on median lobe (Fig. 23 A – C). Gx 11 slightly curved downward; gp 11 as wide as gx 9, forming a furrow, with short setae; gx 9 set apart, relatively broad, almost subrectangular, strongly sclerotized, folded downward (Fig. 23 F – J).</p><p>Female genitalia. Ectoproct elongated, with ventral projection, setae long, black. Tergite 9 sub-rectangular. Gx 9 wider than long, wider than tergite; setae long. Tergite 8 subrectangular. Gx 8 wider than long, with long setae. Gp 8 ventrally prominent; setae thin (Fig. 23 D, E).</p><p>Comparative notes.</p><p>Deleproctophylla variegata differs from most species of Deleproctophylla due to the highly reduced hindwing pigmentation, which is often limited to a slight shading around the crossveins in the radial area (Fig. 3 E, F). Deleproctophylla variegata is easily distinguishable by the male ectoproct with branch arising at two-third of length — the most apical position among Deleproctophylla species (Fig. 23 A – C). Additionally, the caudal narrowing of ectoprocts after the branch is unique to this species.</p><p>Distribution.</p><p>Cyprus, Greece (Aegean and Dodecanese islands), Iran, Iraq, Israel, Jordan, Kazakhstan?, Kyrgyzstan?, Lebanon, Russia? Turkey, Turkmenistan (NEW RECORD), Uzbekistan (H. Aspöck et al. 2001; Monnerat and Ábrahám 2020; Oswald 2025) (Fig. 25).</p><p>Remarks.</p><p>Deleproctophylla variegata is a species with a wide distribution (Oswald 2025), but it is likely that some old records belong to other species (see D. tengri). The specimen reported as D. variegata from Azerbaijan by Kerimova et al. (2022) is D. australis, as shown by molecular data (Fig. 1) and confirmed by examination of the voucher photograph by RD. Records of D. variegata from Kazakhstan, Kyrgyzstan, and Russia are also likely misidentifications and require verification.</p></div>	https://treatment.plazi.org/id/0BD298CE074B5AAE8E4458002D7EDDB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Badano, Davide;Zheng, Yuchen;Aspöck, Ulrike;Aspöck, Horst;Dobosz, Roland;Funari, Rebecca;Pantaleoni, Roberto Antonio;Ábrahám, Levente;Liu, Xingyue	Badano, Davide, Zheng, Yuchen, Aspöck, Ulrike, Aspöck, Horst, Dobosz, Roland, Funari, Rebecca, Pantaleoni, Roberto Antonio, Ábrahám, Levente, Liu, Xingyue (2026): Integrative revision of the Palaearctic owlfly genus Deleproctophylla Lefèbvre (Neuroptera, Myrmeleontidae, Ascalaphinae). ZooKeys 1267: 197-254, DOI: 10.3897/zookeys.1267.173792
