identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E4BD862C75C65EE58E11376337B92DB8.text	E4BD862C75C65EE58E11376337B92DB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tichostephanus Engel 2019	<div><p>Genus † Tichostephanus Engel, 2019</p><p>Type species.</p><p>† Tichostephanus hui Engel, 2019 .</p><p>Differential diagnosis.</p><p>Head with distinct coronal tubercles. Fore wing with first discal cell absent. Ovipositor sheath elongated, with “ bottle-brush-like ” bristles densely developed apically.</p><p>Diagnosis.</p><p>Antenna in males short and robust, with 14 antennomeres; while in females comparatively long and slender, with 13 antennomeres. Head with two unpaired anterior tubercles situated at frons, and three pairs of distinct coronal teeth situated around the ocellus; veins of fore wing strongly reduced; pterostigma distinct elongate and acute apically; vein 1 Rs / 1 M straight, vein 2 Rs + M and 1 m-cu continuous, forming straight elongated vein between anterior apex of subdiscal cell to origin of 2 - Rs, forming large submarginal cell; vein M does not reach the wing margin; vein A extended beyond 1 cu-a and almost reaching 2 Cua; hindwing with cu-a absent. Hind femur slender; hind tibia strongly swollen-appearing distally; hind tarsus 5 - segmented. Metasomal articulation situated slightly lower on the propodeal dorsum; first metasomal sternite not fused with first tergite. Ovipositor slightly longer than metasoma. Ovipositor sheath elongated, with “ bottle-brush-like ” bristles densely developed apically. Male with genitalia protruding. Parameres tapering apically to broadly rounded apex, with fringe of setae developed along inner margin; gonostipes broadly attached to paramere except apically distinct and tapering to narrowly rounded apex; volsella with long, curved, apical lobe approaching apex of Gonostipex; aedeagus tubular, slightly narrower than paramere, not extending beyond paramere apex.</p></div>	https://treatment.plazi.org/id/E4BD862C75C65EE58E11376337B92DB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ge, Si-Xun;Ren, Li-Li;Tan, Jiang-Li	Ge, Si-Xun, Ren, Li-Li, Tan, Jiang-Li (2026): Description of two new species of the mid-Cretaceous genus † Tichostephanus Engel (Hymenoptera, Gasteruptiidae). Journal of Hymenoptera Research 99: 47-57, DOI: 10.3897/jhr.99.175957
A7C571B74CB650CBAEEECCACF1EECD5E.text	A7C571B74CB650CBAEEECCACF1EECD5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tichostephanus qinshubao Ge & Tan 2026	<div><p>† Tichostephanus qinshubao Ge &amp; Tan sp. nov.</p><p>Figs 1, 2</p><p>Holotype.</p><p>BFU, Myanmar Amber, Cretaceous. Part of Si-Xun Ge’s collection. Sex unknown.</p><p>This specimen was previously included in the phylogenetic analysis of Ge et al. (2025) as ‘ † Tichostephanus sp 1 ’.</p><p>Etymology.</p><p>The genus-group name is a combination of the Greek words teichos (meaning ‘ city wall’). The new species was named after a famous minister of the Tang Dynasty (who is also one of the door gods in Chinese culture).</p><p>Diagnosis.</p><p>The new species can be distinguished from all other congeners by the following combination of character states: Neck (modified and more or less narrowed anterior part of pronotum) with two distinct carinae and at distinct lower level than middle part of pronotum from lateral view. The posterior part of the pronotum is slightly ascending above the middle part in lateral view. Pterostigma elongate and narrow, about 12 × as long as its maximum width. Vein 1 Cu slightly curved distally, curvature directed towards the posterior margin; first subdiscal cell approximately rectangular.</p><p>Description.</p><p>Sex unknown because antenna and genitalia are not preserved. Body length (only partially preserved part because of the destroyed metasomal apex) 6.2 mm; fore wing length ca. 2.4 mm.</p><p>Head (Fig. 2 A): Antennae missing, head globular, compound eyes occupying about 2 / 3 of lateral surface; vertex with two unpaired anterior teeth distinctly developed, followed with three pairs of distinct coronal teeth on upper frons and vertex around ocelli; temple more or less bulging; occipital carina distinctly developed. Maxillary and labial palpi not observed.</p><p>Mesosoma (Fig. 2 B): Pronotum robust with neck differentiated; neck at lower level than middle part of pronotum posterodorsally; neck with two distinct carinae, middle and posterior part of pronotum smooth (as laterally); middle part of pronotum weakly differentiated from posterior part; posterior part of pronotum and mesonotum with sparse setosity; scutellum slightly raised above mesonotum, at least partially smooth.</p><p>Wings (Fig. 2 C, D): Fore wing with vein 1 Rs and 1 M merged and straight; 2 - Rs, 2 Rs + M and 1 m-cu merged as continuous (i. e. vein 2 Rs + M + 1 m-cu) vein; pterostigma elongate and narrow, about 12 × as long as its maximum width; vein Rs complete and sclerotized; vein M does not reach the wing margin; subdiscal cell parallel-sided with vein 1 Cu parallel to vein A; vein A extended to 2 Cu-a; 2 Cu-b entirely absent. Hind wing hyaline and clear with only Sc + R present.</p><p>Legs (Fig. 2 E, F): All legs elongate and comparatively slender, without distinct sculpturing; hind coxa comparatively robust, spindle shaped without distinct striae; hind femur slender without ventral dentition, about 8.6 × as long as its maximum width; hind tibia distinct flattened apically; inner side of widened part not concave basally; hind tibia about 1.2 × longer than hind femur, with maximum width of apical widened part more than 6 × as wide as minimum width of basal narrow part; hind tarsus with five tarsomeres; basitarsus 7.5 × as long as wide.</p><p>Metasoma (Fig. 2 E): Metasoma incompletely preserved. First tergum and sternum not fused laterally; tergite I slender but with apical part more or less swollen, more or less conical shaped; tergite I indistinctly narrower than tergite II.</p></div>	https://treatment.plazi.org/id/A7C571B74CB650CBAEEECCACF1EECD5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ge, Si-Xun;Ren, Li-Li;Tan, Jiang-Li	Ge, Si-Xun, Ren, Li-Li, Tan, Jiang-Li (2026): Description of two new species of the mid-Cretaceous genus † Tichostephanus Engel (Hymenoptera, Gasteruptiidae). Journal of Hymenoptera Research 99: 47-57, DOI: 10.3897/jhr.99.175957
67FD9A99C9FC55A18243B9B1A37E1C66.text	67FD9A99C9FC55A18243B9B1A37E1C66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tichostephanus yuchijingde Ge & Tan 2026	<div><p>† Tichostephanus yuchijingde Ge &amp; Tan sp. nov.</p><p>Figs 3, 4</p><p>Holotype.</p><p>♂; BFU, Myanmar Amber, Cretaceous. Part of Si-Xun Ge’s collection.</p><p>This specimen was previously included in the phylogenetic analysis of Ge et al. (2025) as ‘ † Tichostephanus sp 2 ’.</p><p>Etymology.</p><p>The new species was named after a famous minister of the Tang Dynasty (another door god besides Shubao Qin in Chinese culture).</p><p>Diagnosis.</p><p>The new species can be distinguished from all other congeners by the following combination of character states: Neck with at least two distinct carinae, at slightly lower level than middle part of pronotum from lateral view. Pronotum with posterior part distinctly ascending at a higher level than the middle part from the lateral view. Pterostigma elongated but comparatively broad, less than 10 × as long as its maximum width. Vein 1 Cu straight; first subdiscal cell parallelogram shaped; vein 2 Cu-a oblique to vein A.</p><p>Description.</p><p>Male. Body length (except antenna) 6.4 mm; forewing length 2.1 mm.</p><p>Head (Fig. 4 A): Antenna comparatively short and robust, with 12 flagellomeres; scape robust; pedicel short, about as long as wide. Head globular, compound eyes occupying much of anterior portion of lateral surface; vertex with two unpaired triangular tubercles developed anteriorly, followed by three pairs of coronal teeth; Gena slightly bulging; occipital carina distinctly developed; hypostomal carina large, without distinct rugosity but some punctures. Maxillary and labial palpi mostly invisible (basal part concealed).</p><p>Mesosoma (Fig. 4 B): Pronotum robust with neck differentiated; neck at slightly lower level than middle part of pronotum; neck with at least two carinae, middle and posterior part of pronotum coriaceous; posterior part of pronotum distinctly ascending to higher level than middle part in lateral view.</p><p>Wings (Fig. 4 C, D): Forewing with vein 1 Rs and 1 M continuous and straight; 2 - Rs, 2 Rs + M and 1 m-cu continuous vein (i. e. vein 2 Rs + M + 1 m-cu) with individual veins indistinguishable; pterostigma very elongate but comparatively wide, less than 10 × as long as maximum width; vein Rs complete and sclerotized, reaching distal part of costal margin; vein M does not reach the wing margin; subdiscal cell parallel-sided with vein 1 Cu parallel to vein A; vein A extended to 2 Cu-a; 2 Cu-b entirely absent. Hind wing clear, with only Sc + R present.</p><p>Legs (Fig. 4 F): all legs elongate and slender, without distinct sculpture; hind coxa robust, spindle shaped without striae; hind femur slender without ventral dentition, about 7.2 × as long as its maximum width; hind tibia with its apical part distinctly flattened; inner side of the apical part without concavity basally; hind tibia about 1.5 × longer than hind femur, with maximum width of apical widened part up to 5.3 × as wide as minimum width of basal narrow part; hind tarsus with five tarsomeres; basitarsus 11.5 × as long as wide.</p><p>Metasoma (Fig. 4 E): First tergum and sternum not fused laterally, tergite I slender but with apical part more or less swollen, about 1.2 × as long as tergite II and 0.2 × as long as remainder of metasoma; tergite I indistinctly narrower than tergite II. Genitalia only partly exserted, parameres long and broad, with its apex rounded.</p><p>Female. Unknown.</p></div>	https://treatment.plazi.org/id/67FD9A99C9FC55A18243B9B1A37E1C66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ge, Si-Xun;Ren, Li-Li;Tan, Jiang-Li	Ge, Si-Xun, Ren, Li-Li, Tan, Jiang-Li (2026): Description of two new species of the mid-Cretaceous genus † Tichostephanus Engel (Hymenoptera, Gasteruptiidae). Journal of Hymenoptera Research 99: 47-57, DOI: 10.3897/jhr.99.175957
