identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
27059D55ED515430BB87B0058DB58119.text	27059D55ED515430BB87B0058DB58119.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Echiniscus lineatus Pilato, Fontoura, Lisi and Beasley 2008	<div><p>Echiniscus lineatus Pilato, Fontoura, Lisi and Beasley, 2008</p><p>Material examined.</p><p>One specimen; UGANDA; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.37629&amp;materialsCitation.latitude=0.5670147" title="Search Plazi for locations around (long 30.37629/lat 0.5670147)">Kibale NP</a>, Kabarole District; 00°34'1.253"N, 30°22'34.642"E; ca. 1535 m asl; 4 Jul. 2022; coll. Barbara Wiśniewska; moss on tree trunk; barcodes (GenBank: PZ 120891, GenBank: PZ 120889, GenBank: PZ 120886, GenBank: PZ 124266 -7); sample code (UG 77 L) . — One specimen was examined for DNA isolation. The exoskeleton after DNA extraction was mounted in Hoyer’s medium.</p><p>Specimen depository.</p><p>The slide UG 77 / 19 with one individual is deposited in the Department of Animal Taxonomy and Ecology, Institute of Environmental Biology, Adam Mickiewicz University, Poznań, Uniwersytetu Poznańskiego 6, 61-614 Poznań, Poland.</p><p>Morphological identification.</p><p>The morphological characteristics fully correspond to the original species description.</p><p>DNA sequences.</p><p>The sequences obtained from one specimen for all five molecular markers analysed in this study were of good quality. 28 S rRNA sequence (GenBank: PZ 120891) 710 bp long; 18 S rRNA sequence (GenBank: PZ 120889) 714 bp long; COI sequence (GenBank: PZ 120886) 619 bp long; ITS- 1 sequence (GenBank: PZ 124266) 613 bp long; ITS- 2 sequence (GenBank: PZ 124267) 387 bp long.</p><p>Haplotype distribution.</p><p>The analysis included the following numbers of sequences: 30 sequences for COI, nine sequences for ITS- 1, and ten sequences for ITS- 2. The total number of countries from which the material originated was nine for COI (Brazil, the United States, Indonesia, Uganda, Seychelles, Taiwan, Vietnam, Madagascar and Tanzania; Fig. 12 A), five for ITS- 1 (Indonesia, Uganda, Seychelles, Taiwan and Brazil; Fig. 12 B), and five for ITS- 2 (Uganda, Seychelles, Indonesia, Brazil and USA; Fig. 12 C). Results of the analysis revealed no discernible geographic patterns, as haplotype clustering was random and individuals from similar geographic regions were often distantly related in the presented networks.</p></div>	https://treatment.plazi.org/id/27059D55ED515430BB87B0058DB58119	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Warguła, Jędrzej;Stec, Daniel;Dmuchowska, Wiktoria;Krakowiak, Michalina;Polishchuk, Anastasiia;Gawlak, Magdalena;Kaczmarek, Łukasz	Warguła, Jędrzej, Stec, Daniel, Dmuchowska, Wiktoria, Krakowiak, Michalina, Polishchuk, Anastasiia, Gawlak, Magdalena, Kaczmarek, Łukasz (2026): Integrative description of two new tardigrade species from the rainforests of Kibale National Park, Uganda. Arthropod Systematics & Phylogeny 84: 423-446, DOI: 10.3897/asp.84.e185344
F61C053F1D48521EBDE0E7ECEF7EE49E.text	F61C053F1D48521EBDE0E7ECEF7EE49E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Echiniscus osiejuki Warguła, Dmuchowska, Polishchuk, Stec, Kaczmarek 2026	<div><p>3.2. 1. Echiniscus osiejuki sp. nov. Warguła, Dmuchowska, Polishchuk, Stec, Kaczmarek</p><p>Figures 5, 6, 7, 8, 9; Tables 2, 3</p><p>Material examined.</p><p>Twelve specimens; UGANDA; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.37629&amp;materialsCitation.latitude=0.5670147" title="Search Plazi for locations around (long 30.37629/lat 0.5670147)">Kibale NP</a>, Kabarole District; 00°34'1.253"N 30°22'34.642"E; ca. 1535 m asl; 4 Jul. 2022; coll. Barbara Wiśniewska; moss on tree trunk; barcodes at GenBank: PZ 127154 -5, PZ 129998, PZ 127159, PZ 127161; specimen code UG 77 .</p><p>Type repository.</p><p>The holotype (slide: UG 77 / 13 and 8 paratypes (slides: UG 77 / *, where the asterisk can be substituted by any of the following numbers: 11, 13, 14, 15, 16, 17) are deposited in the Department of Animal Taxonomy and Ecology, Institute of Environmental Biology, Adam Mickiewicz University, Poznań, Uniwersytetu Poznańskiego 6, 61-614 Poznań, Poland. Two paratypes (slide: UG 79 / 12) are deposited in Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Sławkowska 17, 31-016, Kraków, Poland.</p><p>Etymology.</p><p>We dedicate this species to our friend, Professor Tomasz Osiejuk from the Department of Behavioural Ecology, Faculty of Biology, Adam Mickiewicz University in Poznań, who initiated the Tropical Biology course series at the Faculty of Biology.</p><p>Description.</p><p>Adults (measurements and statistics in Tables 2, 3). Before mounting, the body orange and plump (Figs 5, 6 A, B), with red eyespots. Bulbous cirrophores are present at the base of all cirri (Fig. 7 A, B). Dactyloid / ovoid cephalic papillae (secondary clavae) and minute (primary) clavae present (Fig. 7 A, B). Cirrus A short. The spines configuration is A- (B) - C-C d - (D) - D d - E (Figs 5 A, 5 B, 6 A, 6 B, 8). Spines D and E may have serrated margins (Fig. 6 A). Asymmetry in the development of appendages is frequently observed, with one of the spines B or D, which are sometimes absent. Specifically, the absence of spine B was noted in one of the eight specimens examined, while spine D was absent in three of the eight specimens. Considerable variation in the length of all spines is observed across all examined individuals. The female gonopore is rosette-shaped (Fig. 9 D).</p><p>The dorsal plates with spinulosus type of sculpture, characterized by irregularly distributed pores, across all plates (Figs 5 A, 5 B, 6 A). However, pores on anterior portion of the first paired plate often gradually become smaller in the direction from the head to the caudal region. Most of pores on the dorsal plate have a diameter ranging from 1.1–2.0 μm in all measured females, 0.9–2.2 μm in all measured males. Dark central rings in pores absent (Figs 5, 7 A). Intracuticular pillars not visible under PCM. The ventral sculpture is visible as tiny granulation, but only in PCM. The cephalic plate is narrow and with an anterior incision (Fig. 7). The cervical (neck) plate is also narrow and formed as a thin dark belt without pores (Figs 5 B, 7 A). A smooth, thin transverse stripe divides the first and second paired plates into smaller, narrower anterior and larger posterior parts (Figs 5 B, 6 A). Median plates I and III are unipartite, whereas median plate II is divided into a narrower anterior and a wider posterior part (Figs 5 B, 6 A). Median plate III is tiny, but densely covered by pores, often partially hidden under the paired plate II and the caudal plate (Figs 5 B, 6 A). The caudal plate with well visible incisions (Fig. 5 B). Pedal plates absent but pulvini present (Fig. 6 B). The first pair of legs with small spine, the fourth pair of legs with papilla (Fig. 9 A). Dentate collar with 7–12 large teeth, present on leg IV (Fig. 9 C). All claws of similar length. External claws smooth. Internal claws with small, slender spurs located close to the claw bases (Fig. 9 B).</p><p>One juvenile (identified by the absence of reproductive organs) was found in the sample and its morphology is similar to that of adult females, including appendage configuration and sculpturing (measurements and statistics in Table 3). Pores on dorsal plates are well visible. Gonopore absent.</p><p>No eggs and larvae were found.</p><p>DNA sequences.</p><p>The sequences obtained from one specimen for all five molecular markers analysed in this study were of good quality. The 28 S rRNA sequences (GenBank: PZ 127155) 711 bp long; 18 S rRNA sequences (GenBank: PZ 127154) 758 bp long; COI sequences (GenBank: PZ 129998) 650 bp long; ITS- 1 sequences (GenBank: PZ 127159) 607 bp long; ITS- 2 sequences (GenBank: PZ 127161) 380 bp long.</p><p>Genetic distances.</p><p>The ranges of uncorrected genetic p - distances between the molecular markers of Ech. osiejuki sp. nov. obtained in our study and the sequences of all species of the genus Echiniscus available in GenBank are as follows (File S 3):</p><p>28 S rRNA: 0.01–0.6 % (0.3 % on average), with the most similar being Ech. manuelae Gąsiorek et al., 2019 (GenBank: MK 529708) and the least similar being Ech. perarmatus Gąsiorek et al., 2022 (GenBank: OM 517009).</p><p>18 S rRNA: 0.0–3.9 % (1.8 % on average), with the most similar being Ech . succineus Gąsiorek &amp; Voncina, 2019 (GenBank: MK 675903) and the least similar being Ech. evelinae Gąsorek et al., 2021 (GenBank: MZ 467757).</p><p>COI: 16.3–25.3 % (19.2 % on average), with the most similar being Ech. tristis Bochnak et al., 2020 (Genbank: MT 374161) and the least similar being Ech . tantulus Bochnak et al., 2020 (GenBank: MT 107427).</p><p>ITS 1 rRNA: 1.9–13.8 % (10.64 % on avarage), with the most similar being Ech. cavagnaroi Gąsiorek et al., 2022 (Genbank: OM 516846) and the least similar being Ech. spinulosus Gąsiorek &amp; Voncina, 2023 (GenBank: PQ 283268).</p><p>ITS 2 rRNA: 1.4–14.9 % (14.2 % on average), with the most similar being Ech. tristis Bochnak et al., 2020 (GenBank: MT 374184) and the least similar being Ech . aonikenk Gąsiorek et. al., 2021 (GenBank: MZ 467833).</p><p>Differential diagnosis.</p><p>Echiniscus osiejuki sp. nov. is diagnosed by the combination of the following morphological characters: a spines configuration [A – (B) – C – C d – (D) – D d – E], lacking internal dark rings inside pores and pores not connected by striae, and the presence of spurs on internal claws of all legs. Taking these diagnostic features the new species is similar to nine Echiniscus species, but it differs specifically from:</p><p>(1) Echiniscus dikenli Maucci, 1972, known only from the type locality in Turkey (Maucci 1972), by: the presence of median plate III, shorter cirrus A (15.6–23.5 μm in Ech. osiejuki sp. nov. vs. 60.0–115.0 μm in Ech. dikenli), shorter spine C d (3.5–15.2 μm in Ech. osiejuki sp. nov. vs. 22.0–34.0 μm in Ech. dikenli) and shorter spine E (5.2–15.5 μm in Ech. osiejuki sp. nov. vs. 42.0–90.0 μm in Ech. dikenli).</p><p>(2) Echiniscus kosickii Kaczmarek and Michalczyk, 2010, known only from the type locality in Costa Rica (Kaczmarek and Michalczyk 2010), by: a longer cirrus internus (9.5–14.5 μm in Ech. osiejuki sp. nov. vs. 8.2–9.1 μm in Ech. kosickii), shorter cephalic papilla (4.6–6.2 μm in Ech. osiejuki sp. nov. vs. 6.3–6.8 μm in Ech. kosickii), a cirrus A / body length ratio (11–13 % in Ech. osiejuki sp. nov. vs. 14–17 % in Ech. kosickii) and a longer spurs on claws I and IV (I: 1.3–2.2; IV: 1.6–2.4, respectively in Ech. osiejuki sp. nov. vs. I and IV: ca. 1.0 in Ech. kosickii).</p><p>(3) Echiniscus marcusi Pilato, Claxton and Binda, 1989, known only from the type locality in Australia (Pilato et al. 1989; Claxton 2004), by: the absence of granulation on legs I – III, shorter cephalic papilla (4.6–6.2 μm in Ech. osiejuki sp. nov. vs. ca. 10.2 μm in Ech. marcusi), a shorter cirrus A (15.6–23.5 μm in Ech. osiejuki sp. nov. vs. ca. 37.8 μm in Ech. marcusi), shorter spine C d (3.8–15.2 μm in Ech. osiejuki sp. nov. vs. ca. 22.5 μm in Ech. marcusi) and longer spine D (12.9–22.0 μm in Ech. osiejuki sp. nov. vs. ca. 8.6 μm in Ech. marcusi).</p><p>(4) Echiniscus minutus Gąsiorek and Michalczyk, 2024, known only from the type locality in Indonesia (Gąsiorek and Michalczyk 2024), by the absence of pedal plates on legs I – III, longer spine D d (12.9–22.0 μm in Ech. osiejuki sp. nov. vs. 3.3–9.8 in Ech. minutus) and a longer spurs on claws III (1.5–1.8 μm in Ech. osiejuki sp. nov. vs. 1.0–1.4 μm in Ech. minutus).</p><p>(5) Echiniscus pooensis Rodriguez-Roda, 1947 known only from the type locality in Ecquatorial Guinea (Rodriguez-Roda 1947), by the orange body color, the presence of papilla on leg IV, a shorter cirrus A (15.6–23.5 μm in Ech. osiejuki sp. nov. vs. ca. 30.0 μm in Ech. pooensis), shorter spine B (3.8–12.7 μm in Ech. osiejuki sp. nov. vs. ca. 22.0 μm in Ech. pooensis), shorter spine C (6.6–16.4 μm in Ech. osiejuki sp. nov. vs. ca. 28.0 μm in Ech. pooensis), shorter spine C d (3.8–15.2 μm in Ech. osiejuki sp. nov. vs. ca. 20.0 μm in Ech. pooensis), shorter spine D (5.9–15.3 μm in Ech. osiejuki sp. nov. vs. ca. 24.0 μm in Ech. pooensis) and shorter spine E (5.2–15.5 μm in Ech. osiejuki sp. nov. vs. ca. 22.0 μm in Ech. pooensis).</p><p>(6) Echiniscus rugospinosus Marcus, 1927 known from Uganda and Kenya (McInnes et al. 2017), by the presence of median plate III. However, considering that Ech. rugospinosus is regarded in the literature as a nomen inquirendum (Gąsiorek and Michalczyk 2024), it is impossible to carry out a detailed morphological comparison between these two species.</p><p>(7) Echiniscus siticulosus Gąsiorek and Michalczyk, 2020, known only from the type locality in Australia, by: the lack of pedal plates, the lack of caudal plate faceting, a larger body size (144–202 μm in Ech. osiejuki sp. nov. vs. 110–132 μm in Ech. siticulosus), longer cirrus internus (9.5–14.5 μm in Ech. osiejuki sp. nov. vs. 4.9–5.1 μm in Ech. siticulosus), longer clava (4.2–6.7 μm in Ech. osiejuki sp. nov. vs. 2.2–3.6 μm in Ech. siticulosus) and longer papilla on leg IV (2.4–4.2 μm in Ech. osiejuki sp. nov. vs. 1.9–2.3 μm in Ech. siticulosus).</p><p>(8) Echiniscus spinulosus (Doyère, 1840), known from Asia, Europe and Hawaiian Islands (McInnes 1994; Gąsiorek and Vončina 2023), by: the presence of pedal plates on legs I – III, the lack of granulation on legs, a smaller body size (144–202 μm in Ech. osiejuki sp. nov. vs. 236–314 μm in Ech. spinulosus), narrower scapular plate (25.6–36.5 μm in Ech. osiejuki sp. nov. vs. 54.3–64.8 μm in Ech. spinulosus), shorter cephalic papilla (4.6–6.2 μm in Ech. osiejuki sp. nov. vs. 8.6–10.5 μm in Ech. spinulosus), a shorter cirrus externus (11.8–15.9 μm in Ech. osiejuki sp. nov. vs. 20.2–29.1 μm in Ech. spinulosus), shorter cirrus A (15.6–23.5 μm in Ech. osiejuki sp. nov. vs. 117.1–166.6 μm in Ech. spinulosus), lower cirrus A / body length ratio (11–13 % in Ech. osiejuki sp. nov. vs. 44–56 % in Ech. spinulosus), longer spine C (6.6–16.4 μm in Ech. osiejuki sp. nov. vs. 2.2–5.2 μm in Ech. spinulosus), shorter spine C d (3.8–15.2 μm in Ech. osiejuki sp. nov. vs. 48.7–62.2 μm in Ech. spinulosus), shorter spine D d (12.9–22.0 μm in Ech. osiejuki sp. nov. vs. 43.6–56.1 μm in Ech. spinulosus), shorter spine on leg I (1.1–2.7 μm in Ech. osiejuki sp. nov. vs. 4.2–6.1 μm in Ech. spinulosus), smaller papilla on leg IV (2.4–4.2 μm in Ech. osiejuki sp. nov. vs. 4.5–5.5 μm in Ech. spinulosus), shorter claws on legs I – IV (I: 7.6–10.8 μm; II: 7.1–10.5 μm; III: 7.1–10.2 μm; IV: 8.3–12.4 μm, respectively in Ech. osiejuki sp. nov. vs. I: 19.1–22.4 μm; II: 16.2–21.5 μm; III: 15.6–22.6 μm; IV: 19.5–28.5 μm, respectively in Ech. spinulosus) and shorter spurs on legs I – IV (I: 1.3–2.2 μm; II: 1.4–1.9 μm; III: 1.5–1.8 μm; IV: 1.6–2.4 μm, respectively in Ech. osiejuki sp. nov. vs. I: 2.4–3.3 μm; II: 2.3–3.4 μm; III: 2.6–3.4 μm; IV: 3.0–4.5 μm, respectively in Ech. spinulosus).</p><p>(9) Ech. tropicalis Binda and Pilato, 1995, known from (Indonesia, Malaysia and Seychelles (type locality) Singapore (Binda and Pilato 1995; Kiosya et al. 2021), by: the presence of pores on the entire median plates I and II (lack of pores on anterior part of median plates I and II in Ech. tropicalis) and a shorter spine D d (12.9–22.0 μm in Ech. osiejuki sp. nov. vs. 5.6–6.6 μm in Ech. tropicalis).</p></div>	https://treatment.plazi.org/id/F61C053F1D48521EBDE0E7ECEF7EE49E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Warguła, Jędrzej;Stec, Daniel;Dmuchowska, Wiktoria;Krakowiak, Michalina;Polishchuk, Anastasiia;Gawlak, Magdalena;Kaczmarek, Łukasz	Warguła, Jędrzej, Stec, Daniel, Dmuchowska, Wiktoria, Krakowiak, Michalina, Polishchuk, Anastasiia, Gawlak, Magdalena, Kaczmarek, Łukasz (2026): Integrative description of two new tardigrade species from the rainforests of Kibale National Park, Uganda. Arthropod Systematics & Phylogeny 84: 423-446, DOI: 10.3897/asp.84.e185344
79AADA4658185487B1AF45F96F75F2E0.text	79AADA4658185487B1AF45F96F75F2E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parahypsibius arletae Warguła, Dmuchowska, Stec, Kaczmarek 2026	<div><p>3.1. 2. Parahypsibius arletae sp. nov. Warguła, Dmuchowska, Stec, Kaczmarek</p><p>Figures 1, 2, 3, 4; Table 1</p><p>Material examined.</p><p>Seven specimens; UGANDA; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=30.353437&amp;materialsCitation.latitude=0.5673889" title="Search Plazi for locations around (long 30.353437/lat 0.5673889)">Kibale NP</a>, Kabarole District; 00°34'2.600"N 30°21'12.373"E; ca. 1535 m asl; 5 Jul. 2022; coll. Zuzanna Kudelska, Bogna Malinowska, Marta Janecka; lichen on fallen tree branch; barcodes (GenBank: PZ 129105 –6, PZ 129999 –30000); sample code (UG 79) .</p><p>Type repository.</p><p>The holotype and two paratypes (slide: UG 79 / 1) and one paratype (slide: UG 79 / 2) and one exoskeleton after DNA extraction (slide: UG 79 / 5) are deposited in the Department of Animal Taxonomy and Ecology, Institute of Environmental Biology, Adam Mickiewicz University, Poznań, Uniwersytetu Poznańskiego 6, 61-614 Poznań, Poland. Two paratypes (slide: UG 79 / 1-2) are deposited in Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Sławkowska 17, 31-016, Kraków, Poland.</p><p>Etymology.</p><p>We dedicate this species to our friend, Professor Małgorzata Arlet from the Institute of Human Biology and Evolution, Faculty of Biology, Adam Mickiewicz University in Poznań, who initiated the Tropical Biology course series at the Faculty of Biology.</p><p>Description.</p><p>Body cylindrical, measurements in Table 1. Eyespots absent after fixing in Hoyer’s medium. The cuticle exhibits a distinct sculpture composed of irregular, roughly polygonal protuberances (2.5–3.7 μm in diameter) arranged in seven transverse bands (Fig. 1 A, B). The first two bands are situated at the level of the first pair of legs and are considerably less distinct than the remaining ones, whereas the subsequent bands are arranged progressively along the body. The seventh and widest band is situated in the caudal region and extends then uniformly as far as the caudal end, which itself is entirely covered by an irregular cuticular pattern of sculpturing composed of irregular protuberances (Figs 1 A, 1 B, 2 A). Areas between bands are smooth. Protuberances gradually increase in size along the rostro-caudal axis, reaching their largest dimensions in the caudal region. Delicate sculpture is present also on the external surface of leg pair IV (Fig. 2 A).</p><p>A pair of elliptical organs (Kihm et al. 2023) present (Fig. 2 B). Peribuccal structures not visible under PCM. Bucco-pharyngeal apparatus of the Ramazzottius type (Fig. 3 A – C). The oral cavity armature absent or not visible under PCM. Apophyses for the insertion of stylet muscles (AISMs) asymmetrical with respect to the frontal plane: dorsal apophysis is shorter, higher, and stumpy, with a prominent caudal apex (Fig. 3 A, C). Ventral apophysis slenderer, with more developed caudal processes (Fig. 3 D). Furcae unmodified, of the Hypsibius type, with well-developed condyles (Fig. 3 A). Buccal tube straight and narrow (Fig. 3 A – C). Pharynx circular, with only two roundish granular macroplacoids, similar in size (Fig. 3 A). Constrictions in macroplacoids absent or not visible under PCM (Fig. 3 B). Microplacoid absent (Fig. 3 A).</p><p>Claws of Ramazzottius - type (sensu Tumanov 2020), with elongated primary branches of external / posterior claws and thinned connective portions in the form of light-refracting units (Fig. 4 A – D). Posterior claws IV always longer than external claws in legs I – III. Accessory points on all primary branches evident and clearly divergent (Fig. 4 A – D). Pseudolunulae at all claws bases present (Fig. 4 B), but often poorly visible, and most prominent under claws IV. Pulvini and cuticular bars on legs absent. Eggs not found.</p><p>DNA sequences.</p><p>The sequences obtained from one specimen for two out of four molecular markers analysed in this study were of good quality. The amplification of 28 S rRNA and ITS- 2 was not successful. The 18 S rRNA sequences (GenBank: PZ 129105 –6) 878 bp long; COI sequences (GenBank: PZ 129999 –30000) 641 bp long.</p><p>Differential diagnosis.</p><p>Currently there are nine species included in the genus Parahypsibius: Pap. biscuitiformis (Bartoš, 1960); Pap. calcaratus (Bartoš, 1935); Pap. camelopardalis (Ramazzotti and Maucci, 1983); Pap. macrocalcaratus (Beasley, 1988); Pap. ragonesei (Binda and Pilato, 1985); Pap. roanensis (Nelson and McGlothin, 1993); Pap. runae (Bartoš, 1941); Pap. scabropygus (Cuénot, 1929), and Pap. stiliferus (Abe, 2004) . The new species differs specifically from:</p><p>(1) Pap. biscuitiformis, known only from the type locality in the Czech Republic (Bartoš 1960), by: the absence of sculpture on the head region (the entire body covered with uniform granular sculpture in Pap. biscuitiformis), the absence of the subcentral constriction on macroplacoid I, shorter macroplacoid I (1.5–2.3 μm in Pap. arletae sp. nov. vs. ca. 4.05 μm in Pap. biscuitiformis) and reticulated cuticle on legs IV.</p><p>(2) Pap. calcaratus, known only from its type locality in Slovakia (Bartoš 1935), by: the absence of sculpture on the head region (the entire body is covered with an irregular sculpturing in Pap. calcaratus).</p><p>(3) Pap. camelopardalis, known only from type locality in Portugal and Spain (Ramazzotti and Maucci 1983), by: the absence of sculpture on the head region (the entire body covered with granular sculpture in Pap. camelopardalis), lack of the sculpture on the legs I – III, wider buccal tube (1.0–1.8 μm in Pap. arletae sp. nov. vs. 4.0 μm in Pap. camelopardalis) and smaller protuberances on the dorsal side of the body (2.5–3.7 μm in Pap. arletae sp. nov. vs. ca. 6.0 μm in Pap. camelopardalis).</p><p>(4) Pap. macrocalcaratus, known only from USA (Beasley 1988, 1990), by: the absence of sculpture on the head region (the entire body covered with an irregular sculpturing in Pap. macrocalcaratus) and larger protuberances on the dorsal side of the body (2.5–3.7 μm in Pap. arletae sp. nov. vs. ca. 2.0 μm in Pap. macrocalcaratus).</p><p>(5) Pap. ragonesei, known only from the type locality in Italy (Binda and Pilato 1985), by: the absence of sculpture on the head region (the entire body covered with an irregular sculpturing in Pap. ragonesei) and the absence of cuticular bars between claws on all the legs.</p><p>(6) Pap. roanensis, known only from the type locality in USA (Nelson and McGlothin 1993), by: the absence of sculpture on the head region (the entire body covered with an irregular sculpturing in Pap. roanensis).</p><p>(7) Pap. runae, known only from the type locality in Ukraine (Bartoš 1941), by: the absence of sculpture on the head region (the entire body covered with an irregular sculpturing in Pap. runae).</p><p>(8) Pap. scabropygus, known from Denmark, France (type locality), Poland and Scotland (Cuénot 1929; Gąsiorek et al. 2024 a), by: the different dorsal sculpture (sculpture composed of seven bands extending from the first pair of legs to the caudal region in Pap. arletae sp. nov. vs. sculpture present only on caudal region in Pap. scabropygus).</p><p>(9) Pap. stiliferus, known only from the type locality in Russia (Abe 2004), by: the absence of sculpture on the head region (the entire body covered with sculpturing in Pap. stiliferus).</p><p>Remarks.</p><p>Due to the very limited number of sequences currently available in public databases for the genus Parahypsibius, p - distances were not calculated. At present, sequences for only three taxa have been published, which is insufficient to perform a meaningful comparison.</p></div>	https://treatment.plazi.org/id/79AADA4658185487B1AF45F96F75F2E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Warguła, Jędrzej;Stec, Daniel;Dmuchowska, Wiktoria;Krakowiak, Michalina;Polishchuk, Anastasiia;Gawlak, Magdalena;Kaczmarek, Łukasz	Warguła, Jędrzej, Stec, Daniel, Dmuchowska, Wiktoria, Krakowiak, Michalina, Polishchuk, Anastasiia, Gawlak, Magdalena, Kaczmarek, Łukasz (2026): Integrative description of two new tardigrade species from the rainforests of Kibale National Park, Uganda. Arthropod Systematics & Phylogeny 84: 423-446, DOI: 10.3897/asp.84.e185344
