taxonID	type	description	language	source
E025B554FFED6D056A71F9CBFCE6AFD0.taxon	type_taxon	Type species: Carabodes fraterculus Balogh, 1963	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFED6D0169B8F967FD2CADDB.taxon	materials_examined	Type material Holotype (male) and one paratype (one male): Republic of Zambia, North of Ndole at Lake Tanganyika, Ndole Bay Lodge (garden), 8 ° 28 ′ 39.5 ″ S, 30 ° 26 ′ 56.8 ″ E, 785 m a. s. l., rotten wood and humus at the basis of the trunk of a dead tree (Fig. 1), 04. x. 2021 (F. D. B. Schedel). Type deposition The holotype and one paratype are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol. Diagnosis Body length: 330. Prodorsum and centrodorsal part of notogaster foveolate; humeral, lateral and posterior part of notogaster tuberculate. Rostral margin concave. Basal part of prodorsum with depression. Rostral and lamellar setae short, setiform, roughened; interlamellar seta minute, acicular; bothridial seta medium-sized, clavate, with slightly flattened, barbed head. All notogastral, epimeral, and anogenital setae minute, acicular. Palp tibia with two setae. Aggenital setae present. Leg seta l' absent on tibiae I, II; all tarsal setae u modified. Description	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFED6D0169B8F967FD2CADDB.taxon	description	Measurements – Body length: 330 (holotype), 330 (paratype); body width: 180 (holotype), 173 (paratype). Integument (Figs. 3 – 12) – Body color light brown. Body with dense, microgranulate microsculpturing; additionally: prodorsum, subcapitular mentum, anal plate, centrodorsal part of notogaster, anterolateral part of epimere I foveolate (diameter of foveola up to 7); basal part of prodorsum with slight transverse ridges bordering depression; humeral, lateral, and posterior part of notogaster tuberculate (diameter of tubercle up to 7); lateral part of anogenital region partially with slight foveolae and depressions; antiaxial side of leg femora I – IV with some large foveolae and small, dense, simple and elongate tubercles versus trochanters III, IV only with tubercles. Prodorsum (Figs. 3, 4, 6) – Rostral margin slightly concave medially (visible in anterior aspect). Distal tooth-like process of lamella long, almost reaching rostral margin; translamella present (visible in dorsal aspect). Basal part of prodorsum with distinct depression. Lateral part of prodorsum with thin longitudinal ridge bearing some small teeth. Rostral and lamellar setae (15) setiform, roughened; interlamellar seta (4) acicular; bothridial seta (34 – 37) clavate, with slightly flattened, barbed head; exobothridial seta absent. Notogaster (Figs. 3, 6, 7) – Anterior notogastral margin nearly straight. Humeral process distinctly developed, anteriorly triangular (visible in lateral aspect). All notogastral setae (4) acicular. Opisthonotal gland opening and lyrifissures ia, im, ip distinct; ih, ips not observed. Gnathosoma (Fig. 8) – Generally, similar to Gymnobodes (Ermilov and Yurtaev 2023). Subcapitulum size: 79 × 60; subcapitular setae a (11) and m (7) setiform, roughened; h (4) acicular. Palp length: 49; setation: 0 – 2 – 1 – 2 – 8 (+ ω); postpalpal seta (7) spiniform, roughened. Chelicera length: 94; setae (cha: 26; chb: 15) setiform, barbed. Epimeral and lateral podosomal regions (Figs. 5, 6) – Sternal region slightly depressed. Epimeral setal formula: 1 – 1 – 3 – 3; all setae (4) acicular. Discidium tubercle-like. Anogenital region (Figs. 5 – 7) – Region between epimere IV and genital aperture depressed. Anogenital formula: 4 – 1 – 2 – 3; all setae (4) acicular. Adanal lyrifissure not observed. Legs (Figs. 9 – 12) – Claw of each leg strong, smooth. Femora I – IV with dorsoparaxial porose area. Formulas of leg setation and solenidia: I (1 – 4 – 3 – 3 – 16) [1 – 2 – 2], II (1 – 4 – 3 – 2 – 15) [1 – 1 – 2], III (2 – 3 – 1 – 2 – 15) [1 – 1 – 0], IV (1 – 2 – 2 – 2 – 12) [0 – 1 – 0]; homology of setae and solenidia indicated in Table 1; seta l' absent on tibiae I, II; seta s of tarsus I eupathidial; all tarsal setae u modified (with cylindrical mediobasal part, distal spike and ventral tooth). Comparison Gymnobodes schedeli sp. nov. differs from all species of the genus in having of a strong depression in the basal part of the prodorsum (versus depression absent in other species), and the presence of foveolae in centrodorsal part of the notogaster (versus notogaster entirely tuberculate in other species). Etymology This species is named in honour of Dr. Frederic D. B. Schedel, the German zoologist and ichthyologist, who collected this new species.	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFE96D0169A0FAD0FD2EACBE.taxon	type_taxon	Type species: Notaspis alatus Hermann 1804	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFE96D0D6983FA4DFD0DADF5.taxon	materials_examined	Type material Holotype (male) and two paratypes (two males): Uganda, Buvi, Nortwest of Buvi Lodge, wetland at the shore of Lake Victoria, 0 ° 07 ′ 43.0 ″ N, 32 ° 26 ′ 44.0 ″ E, 1146 m a. s. l., detritus among Cyperus papyrus (Fig. 2), 26. vii. 2022 – 1. viii. 2022 (D. Georgiev). Type deposition The holotype and two paratypes are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol. Diagnosis Body length: 495 – 525. Rostrum protruding, rounded. Rostral region with strong bulge. Lamellar and sublamellar lines parallel, curving backwards at ventral end. Rostral, lamellar, and interlamellar setae long, setiform, barbed; in ˃ le ˃ ro; bothridial seta long, lanceolate, with unilaterally dilated, barbed head having distal spike. Dorsosejugal porose area present. Dorsosejugal suture complete. Anteromedial part of notogaster with long longitudinal fold having short lateral branches. Four pairs of porose areas developed: Aa elongate oval, longitudinally oriented, located close to pteromorphal hinge; A 1 and A 2 nearly circular; A 3 elongate oval. Median pore absent. Opisthonotal gland opening located close and lateral to A 1; lyrifissure im located between setal alveoli lm and lp. Epimeral setal formula: 1 - 0 - 1 - 2. All epimeral and anogenital setae short, setiform, roughened. Circumpedal carina medium-sized. Postanal porose area elongate oval. Leg famulus straight, inserted close and anterolateral to solenidion ω 1; solenidion of tibia IV inserted in anterior part of dorsal side of segment. Description	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFE96D0D6983FA4DFD0DADF5.taxon	description	Measurements – Body length: 495 (holotype), 525, 525 (paratypes); body width: 315 (holotype), 300, 330 (paratypes). Integument (Figs. 13, 14, 16, 18, 19, 22) – Body color brown. Body slightly microgranulate microsculpturing, tubercles in rostral region and lateral part of epimere I larger; subcapitular mentum and antiaxial side of leg femora I – IV and trochanters III, IV partially with simple and elongate tubercles; laterobasal part of prodorsum slightly striate; anteromedial part of notogaster with long longitudinal fold having short lateral branches. Prodorsum (Figs. 13, 14, 16) – Rostrum protruding, rounded. Rostral region with strong bulge. Lamellar and sublamellar lines parallel, curving backwards at ventral end; L thicker than S. Rostral (64 – 67), lamellar (90 – 97), and interlamellar (108 – 116) setae setiform, barbed; bothridial seta (108 – 116) lanceolate, with long, roughened stalk and short, unilaterally dilated, barbed head having distal spike. Dorsosejugal porose area elongate oval (22 × 7), transversely oriented, located posterior to insertion of interlamellar seta. Dorsophragma comparatively long, elongated longitudinally. Notogaster (Figs. 13, 16, 17) – Dorsosejugal suture complete, simple. All notogastral setae represented by setal alveoli. Four pairs of porose areas developed: Aa (34 – 41 × 15 – 19) elongate oval, longitudinally oriented; A 1 (19 – 22) and A 2 (15 – 19) nearly circular; A 3 elongate oval (34 – 41 × 19 – 22); Aa close to pteromorphal hinge, located anterior to la. Median pore absent. Opisthonotal gland opening and all lyrifissures distinct: gla close and lateral to A 1; im between lm and lp; ip lateral to p 1; ih and ips anterior to p 3. Gnathosoma (Fig. 18) – Similar to Galumna (Ermilov 2022). Subcapitulum size: 127 – 131 × 105 – 112; subcapitular setae (a: 22 – 26; m: 13 – 15; h: 19 – 22) and both adoral setae (13 – 15) setiform, slightly barbed; m thinner than a and h. Palp length: 94; setation: 0 - 2 - 1 - 3 - 9 (+ ω); postpalpal seta (7) spiniform, smooth. Chelicera length: 150; setae (cha: 49; chb: 34) setiform, barbed. Epimeral and lateral podosomal regions (Figs. 15, 16) – Epimeral setal formula: 1 - 0 - 1 - 2; all setae (1 b, 3 b: 26 – 30; 4 a, 4 b: 19 – 22) setiform, roughened. Circumpedal carina directed to insertion of seta 3 b, bit not reaching it. Anogenital region (Figs. 15 – 17) – Anogenital formula: 6 - 1 - 2 - 3; genital (g 1: 22; g 2: 17 – 19; others: 13 – 15), aggenital (19 – 22), anal (19 – 22), and adanal (19 – 22) setae setiform, roughened; anterior edge of genital plate with three setae; aggenital setae equally distanced from genital and anal apertures; ad 1 and ad 2 posterior, ad 3 lateral to anal plate; distance ad 1 – ad 2 shorter than ad 2 – ad 3. Adanal lyrifissure located close and parallel to anal plate. Postanal porose area elongate oval (56 – 60 × 15 – 19). Legs (Figs. 19 – 22) – Median claw distinctly thicker than lateral claws; all claws slightly barbed on dorsal side. Porose area on femora I – IV and on trochanters III, IV distinct; proximoventral porose area on all tarsi and distoventral porose area on all tibiae not observed. Formulas of leg setation and solenidia: I (1 - 4 - 3 - 4 - 20) [1 - 2 - 2], II (1 - 4 - 3 - 4 - 15) [1 - 1 - 2], III (1 - 2 - 1 - 3 - 15) [1 - 1 - 0], IV (1 - 2 - 2 - 3 - 12) [0 - 1 - 0]; homology of setae and solenidia indicated in Table 2; famulus straight, inserted close and anterolateral to solenidion ω 1; seta s of tarsus I eupathidial; solenidion of tibia IV inserted in anterior part of dorsal side of segment. Comparison	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
E025B554FFE96D0D6983FA4DFD0DADF5.taxon	etymology	Etymology This species is named in honour of the late Prof. Dr. Luis S. Subías, the Spanish acarologist, who previously identified this new species.	en	Ermilov, Sergey G., Friedrich, Stefan, Georgiev, Dilian (2025): Two new species of oribatid mites (Acari, Oribatida) from the Afrotropical region. Persian Journal of Acarology 14: 345-356, DOI: 10.22073/pja.v14i2.86976
