identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E13FD4141724D3154C8AFBA07CCEFA89.text	E13FD4141724D3154C8AFBA07CCEFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Archicupes Rohdendorf 1961	<div><p>Genus Archicupes Rohdendorf, 1961</p><p>Type species. Archicupes jacobsoni Rohdendorf, 1961 .</p><p>Species included. Type species? A. reichardti Rohdendorf, 1961 (could be indeed a member of the genus Kaltanicupes rather than of Archicupes; Kirejtshuk, 2020), and Archicupes storozhenkoi Yan &amp; Strelnikova, sp. nov.</p></div>	https://treatment.plazi.org/id/E13FD4141724D3154C8AFBA07CCEFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141724D3164C8AFA8478E7FEC0.text	E13FD4141724D3164C8AFA8478E7FEC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Archicupes storozhenkoi Yan & Felker & Strelnikova 2025	<div><p>Archicupes storozhenkoi Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: 058489FC-A200-485E-AF3A-798B4CFA9F49</p><p>(Fig. 2)</p><p>Material. Holotype — PIN, No. 5890/8. Part and counterpart of isolated elytron, left.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Epipleural margin with small cells, forming two parallel rows near the humerus, at midlength become a singular row. Elytron with four visible veins, vein 3 reaching at least at elytral distal fourth, vein 6 is stemming from 4 at the level of humerus, enclosing the vein 5, all reaching beyond elytral midlength. Field III is widest at the elytral base.</p><p>Description. Elytron with maximum width at distal third, apex pointed, asymmetrical, not elongated into thin, tail-like extension. Cells rather large, rounded, organized irregularly, without maculae. Cells of different shapes, mostly round, or rounded-oval, some deformed and concave, usually smaller at the elytral base. Cuticular surface between cells glabrous. Singular row of cells between first vein and epipleural carina, field III is extended at the base and has up to 4 cells, then reduced to two rows. Number of cells varies from single to double in fields IV and V. There are 3–4 cells row in the field VI. Cells somewhat organize uniformly at elytron`s apical fourth.</p><p>Measurements (in mm). Elytron length 5.8, width 1.8.</p><p>Etymology. After prominent entomologist Sergei Yu. Storozhenko.</p><p>Remarks. Species below were attributed to Permocupedidae for having more than three veins, lattice elytra cell pattern and small overall size. The specimen was attributed to the genus Archicupes for having two rows of cells in a first field and three central veins having a common base, also for the lack of vein closest to sutural margin (A2 sensu Ponomarenko 1969).</p><p>Comparison. Differs from most other species in having elytral apex not extended into tail-like structure, also in three rather than two central veins having a common base and field III being widest at the base.</p></div>	https://treatment.plazi.org/id/E13FD4141724D3164C8AFA8478E7FEC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3164C8AFA257877F9C5.text	E13FD4141727D3164C8AFA257877F9C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cytocupes Rohdendorf 1961	<div><p>Genus Cytocupes Rohdendorf, 1961</p><p>Type species. Cytocupes angustus Rodendorf, 1961 Species included. Type species and Cytocupes magnus Yan &amp; Strelnikova, sp. nov.</p></div>	https://treatment.plazi.org/id/E13FD4141727D3164C8AFA257877F9C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141727D3174C8AF94D7FA9FD99.text	E13FD4141727D3174C8AF94D7FA9FD99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cytocupes magnus Yan & Felker & Strelnikova 2025	<div><p>Cytocupes magnus Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:pub: E08412CD-3FDD-412A-A821-D32E712957DF</p><p>(Fig. 3)</p><p>Material. Holotype — PIN, No. 5890/10. Isolated elytron, left.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Veins 4 and 5 are very closely spaced. Veins 2 and 3 are merging at the level of elytral distal 1/5. Veins 4 and 5 half as long as the elytron. Double rows of cells nearly touching each other’s borders, in the field III, IV and VI.</p><p>Description. Elongated elytron, 4 times as long as wide, carinated, apex symmetrical, rounded. Epipleural and sutural margins lacks cells or punctures. Humeral bulge weakly defined. Veins 2 and 3 are merging at the level of elytral distal 1/5. Veins 4 and 5 are close to each other, half as long as the elytron. Fields III, V and VI has straight double row of cells, field IV have 1–2 rows of cells. Field IV are expanded at the base and has up to 3 cells. Cells roundish, rarely oval-shaped, without encrusted maculae. Cells larger near humeral area, size decrease very slightly towards the apex.</p><p>Measurements (in mm). Elytron length 11.6, width 2.9.</p><p>Etymology. After Latin “ large ”, due to considerable elytron size.</p><p>Remarks. The specimen was attributed to the genus Cytocupes due to having veins 2 and 3 merging near elytral apex; having short vein 4 ( CuA sensu Ponomarenko 1969) near elytral base; lack of cells on epipleural margin.</p><p>Comparison. Differs from other congener species in large size; having elytral apex not elongated into tail-like extension; field IV being wider than V, having 3 cells in it instead of 6.</p></div>	https://treatment.plazi.org/id/E13FD4141727D3174C8AF94D7FA9FD99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3174C8AF97878D6F8A8.text	E13FD4141726D3174C8AF97878D6F8A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tatarocupes Ponomarenko 2004	<div><p>Genus Tatarocupes Ponomarenko, 2004</p><p>Type species. Tatarocupes granulatus Ponomarenko, 2004 Species included. Type species and Tatarocupes brevicellularis Yan &amp; Strelnikova, sp. nov.</p></div>	https://treatment.plazi.org/id/E13FD4141726D3174C8AF97878D6F8A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141726D3104C8AF8A07961FD55.text	E13FD4141726D3104C8AF8A07961FD55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tatarocupes brevicellularis Yan & Felker & Strelnikova 2025	<div><p>Tatarocupes brevicellularis Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: 095094C5-0555-422B-B891-36381E9923E7</p><p>(Fig. 4)</p><p>Material. Holotype — PIN, No. 5890/13. Isolated elytron, left . Paratypes: PIN, No 5890/12, part and counterpart, and PIN, No 5890/15, partially preserved isolated elytra.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Elytron of middle size, carinated, bearing at least 5 veins and one scutellar striole. Cells roundish, or rarely slightly oval-shaped, reinforced with dark maculae. Elytral surface covered with dense tubercles. Vein 2 drows close to epipleural carina near humerus; Vein 3 very short, as long as combined length of 4 cells. Cells between veins are placed unevenly, without forming well-defined longitudinal rows.</p><p>Description. Epipleural carina widening at basal third, then narrowing towards elytral apex. Vein 2 drows located close to epipleural carina near humerus then widens posteriorly; Vein 3 very short, as long as combined length of 4 cells, veins 3 and 4 are close together. Epipleuron without cells, in the field II up to 3 rows of cells, field III — 3 in central part and 4 in basal part, IV —1 row of cells. There are more than 34 cells in a row.</p><p>Measurements (in mm). Holotype —Elytron length 8.6, width 3; paratype PIN, No. 5890/15—length of the preserved part of the elytron 6.9, width 2.6.</p><p>Etymology. After Latin “ brevis ”—small, and “ cellularis ”—due to lattice pattern formed by cells.</p><p>Remarks. The specimen was attributed to the genus Tatarocupes for having at least 6 rows of cells between two veins, closest to sutural margin.</p><p>Comparison. Differs in having a single row of cells between veins 3 and 4 (at least near the elytral base).</p></div>	https://treatment.plazi.org/id/E13FD4141726D3104C8AF8A07961FD55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141721D3114C8AF94C7D4DFEC1.text	E13FD4141721D3114C8AF94C7D4DFEC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alisacupes Yan & Strelnikova 2025	<div><p>Genus Alisacupes Yan &amp; Strelnikova, gen. nov.</p><p>urn:lsid:zoobank.org:act: 69841F08-2024-4094-B609-EF8AEA8953E8</p><p>Type species. Alisacupes dunaevi Yan &amp; Strelnikova, sp. nov.</p><p>Diagnosis. Elytron with 4 veins, at least two merging at the elytral apex. Epipleuron covered with tubercles.</p><p>Composition. Monotypic.</p><p>Description. Large elytra, elongated, apex slightly pointed, assymetrical. Epipleural flange wide, at least as wide as several cell`s width combined. Elytral cells roundish, walls reinforced with maculae, cuticular surface between cells covered with tubercles. Сells are surrounded by macula.</p><p>Etymology. After Alisa quarry, source locality of the described material. Gender: masculine.</p><p>Remarks. Species below was attributed to Asiocoleidae for having four veins, with three of them being on the elytral disc; large size of the elytron.</p><p>Comparison. Differs in moderately large size; in having tubercles at proximal half of epipleron`s length. Similar to Loculitricoleus but differs from it in larger size, tubercles sculpture on elytron and by four main veins instead of three.</p></div>	https://treatment.plazi.org/id/E13FD4141721D3114C8AF94C7D4DFEC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141720D3114C8AFE557E2DFC1E.text	E13FD4141720D3114C8AFE557E2DFC1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alisacupes dunaevi Yan & Felker & Strelnikova 2025	<div><p>Alisacupes dunaevi Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: 96B08ED0-B5A4-4E58-A2B7-65B56A1688FA</p><p>(Fig. 5)</p><p>Material. Holotype — PIN, No. 5890/16. Part and counterpart of isolated elytron, left.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. As for the genus.</p><p>Description. Elytron with 4 veins, at least 2 nd and 3 rd are merging at elytral apex. Epipleural flange as wide as combined width of two cells near elytral base, slightly widens near the apex. Number of cell rows in the fields: II —2, field III — 2 in central part and 3 near elytral basal, field IV —4, field V —3, field VI —1. There are up to 43 cells in a row.</p><p>Measurements (in mm): eytron length 13.6, elytron width 4.</p><p>Etymology. After prominent naturalist, teacher and herpetologist E.A. Dunaev.</p><p>Remarks. The specimen was attributed to the genus Tatarocupes for having at least 6 rows of cells between two veins, closest to sutural margin.</p></div>	https://treatment.plazi.org/id/E13FD4141720D3114C8AFE557E2DFC1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3124C8AFF6978AEFEB9.text	E13FD4141723D3124C8AFF6978AEFEB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizocoleidae Rohdendorf 1961	<div><p>Family Schizocoleidae Rohdendorf, 1961</p><p>Species below attributed to Schizocoleidae due to lack of visible elytral veins and cells, but having an interlocking longitudinal protrusion (“schiza”) on the underside of the elytron, close to epipleural margin.</p></div>	https://treatment.plazi.org/id/E13FD4141723D3124C8AFF6978AEFEB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3124C8AFE957EB2FCB5.text	E13FD4141723D3124C8AFE957EB2FCB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Uskatocoleus Rohdendorf 1961	<div><p>Genus Uskatocoleus Rohdendorf, 1961</p><p>Type species. Uskatocoleus zalesskyi Rohdendorf, 1961</p><p>Species included. In total, 19 species: type species, U. artus Ponomarenko, 2015, U. avitus Rohdendorf, 1961, U. convexus Rohdendorf, 1961, U. convexus Ponomarenko, 2013, U. eurypygus Ponomarenko, 2011, U. foveolatus Rohdendorf, 1961, U. hirsutus Ponomarenko, 2015, U. iljinskiensis Rohdendorf, 1961, U. kaltanicus Rohdendorf, 1961, U. kushejakovicus Rohdendorf, 1961, U. lutugini Rohdendorf, 1961, U. micron Ponomarenko, 2013, U. minimus Rohdendorf, 1961, U. minor Ponomarenko, 2015, U. rhynchophorus Ponomarenko, 2011, U. sukhonensis Ponomarenko, 2013, U. surijokovicus Rohdendorf, 1961, and U. uralensis Ponomarenko, 2013 .</p><p>Remarks. U. convexus Rohdendorf, 1961 and U. convexus Ponomarenko, 2013 are homonyms, therefore, in this article a new species name is proposed for the junior homonym— U. isadicus nom. nov. in honor of the location of Isady, from where this species was described.</p></div>	https://treatment.plazi.org/id/E13FD4141723D3124C8AFE957EB2FCB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD4141723D3134C8AFCA17F15FF5D.text	E13FD4141723D3134C8AFCA17F15FF5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Uskatocoleus turensis Yan & Felker & Strelnikova 2025	<div><p>Uskatocoleus turensis Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: CA56BE67-C642-4D1C-B205-16AC6DB67047</p><p>(Fig. 6)</p><p>Material. Holotype — PIN, No. 5890/1, part and counterpart, a pair of interlocked elytra with mesothorax(mesonotum and mesoscutellum), abdominal apex and mesotarsus—partially preserved . Paratypes: PIN, No. 5890/5 and PIN, No. 5890/7, parts and counterparts of isolated elytra, right and left respectively.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Scutellum large, semicircular, only slightly shorter than 1/2 of elytral tongue`s (schiza) length; scutellar incision on the sutural margin near elytral base, forms a weak protrusion. Humerus forms almost right angle. Elytra`s undersides bear very long elytral tongue, as long as 1/5 of elytral overall length, slightly outwardly curved. Elytron`s apices flattened.</p><p>Description. Elongated elytron with round bases, proximal 2/3 of the length—parallel-sided, then gradually narrowing posteriorly, distal third oblique, width to length ratio 1:5. Elytral surface uniformly covered with punctures, inner structures (columellae also visible on the elytral surface as bigger punctures). Humeral bulges indistinct. Elytron`s tongues located at the midlength. Elytron edged, which is slightly thicker around humerus, at basal ¼ and at the rounded apex.</p><p>Remnants of M, Cu veinal systems and bases of A are present.</p><p>Mesocoxae transverse, slightly oblique, contiguous. X-shaped metathoracic scutoscutellar suture is partially visible. Apical abdominal ventrite subtrapezoid, almost as long as penultimate sternite, abdominal length makes less than a half of elytral length.</p><p>Measurements (in mm). Holotype —elytral length 7.1 (measured without elytron`s basal attachment protrusions), maximum width 1.6; paratype PIN, No. 5890/5—length 9.5, width 2.7; paratype PIN, No. 5890/7— length 9.1, width 2.8.</p><p>Etymology. After Tura village, nearest settlement to Alisa quarry, where types specimens were found.</p><p>Remarks. Attributed to Uskatocoleus for having finely sculptured elytral surface, well defined humeral bulge and straight interlocking longitudinal protrusion (“schiza”).</p><p>As an elytra-only family, Schizocoleidae possess very few diagnostic characters, specimens are mainly divided into taxa by locality and age.</p><p>Cuticle on PIN, No. 5890/1 could been rather thin, since remnants of hindwing venation could be observed.</p><p>Comparison. Similar to Uskatocoleus zalesskyi Rohdendorf, 1961 in having long elytral tongue, but differs in lacking of fine longitudinal stria and having twice the length. Similar to U. angustior Rohdendorf, 1961 in having long elytral tongue, but differs in twice the length, having concave elytral tongue and parallel-sided 2/3 of basal elytral length. Differs from Rossocoleus verrucosus Rohdendorf, 1961, R. latus Rohdendorf, 1961 and R. latior Rohdendorf, 1961 in twice the length. Differs from Rhombocoleites uralensis Ponomarenko, 2013 in having longer elytral tongue and apical abdominal sternite half as long.</p></div>	https://treatment.plazi.org/id/E13FD4141723D3134C8AFCA17F15FF5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFE497F2AFBFD.text	E13FD414173DD30C4C8AFE497F2AFBFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizocoleus maximus Yan & Felker & Strelnikova 2025	<div><p>Schizocoleus maximus Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: F33EB2B8-5A3C-411C-AEED-E76147446497</p><p>(Fig. 7)</p><p>Material. Holotype — PIN, No. 5890/2, part and counterpart of left elytron, base and apex partially damaged . Paratypes — PIN, No. 5890/3, 6 and 9 parts and counterparts of left elytra, partially damaged .</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Very large elytron, narrowing anteriorly and posteriorly with maximum width at midlength.</p><p>Description. At least three times as long as wide, maximum width at two thirds of its length; carinated; slightly curved near distal third. Locking tongue straight about ¼ of elytron`s length, located at 1/3 of elytron`s width, closer to epipleural margin at elytron`s basal half.</p><p>Specimens 5890/2 and 6 have remnants of R and M veinal systems visible.</p><p>Measurements (in mm). Holotype’s elytral length 14.5, width 4.5; paratype`s PIN 5890/3 length 12.3, width 4; paratype`s PIN 5890/6 length 11.7, width 4.7; paratype`s PIN 5890/9 length 9.5, width 2.7.</p><p>Etymology. After Latin “ maximus ” due to elytron`s large size, largest among Schizocoleus .</p><p>Remarks. Elytral cuticle was probably rather thin, this allowed folded wings to be partially seen after fossilization, similar to Uskatocoleus turensis sp. nov. Species below attributed to Schizocoleus mainly because of its large size and lack of wrinkled or scaly cuticular sculpture.</p><p>Comparison. Differs from all congeners by large size.</p></div>	https://treatment.plazi.org/id/E13FD414173DD30C4C8AFE497F2AFBFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173DD30C4C8AFF6978D9FEC1.text	E13FD414173DD30C4C8AFF6978D9FEC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizocoleus Rohdendorf 1961	<div><p>Genus Schizocoleus Rohdendorf, 1961</p><p>Type species. Schizocoleus kuznetskiensis Rohdendorf, 1961,</p><p>Species included. In total, 9 species: type species, S. depressus Ponomarenko, 2013, S. glabrus Ponomarenko, 2004, S. longus Ponomarenko, 2004, S. major Ponomarenko, 2013, S. minimus Ponomarenko, 2013, S. robustus Ponomarenko, 2013, S. septentrionalis Ponomarenko, 2008, and S. triassicus Ponomarenko, 2016</p></div>	https://treatment.plazi.org/id/E13FD414173DD30C4C8AFF6978D9FEC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173CD30D4C8AFF697FAAFD55.text	E13FD414173CD30D4C8AFF697FAAFD55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizocoleus frumentum Yan & Felker & Strelnikova 2025	<div><p>Schizocoleus frumentum Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: 4BE4F103-A9EF-44FB-8CA6-3C91CBE0E0A</p><p>(Fig. 8)</p><p>Material. Holotype — PIN, No. 5890/14, right elytron . Paratype — PIN, No. 5890/4, part and counterpart of left elytron .</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Asymmetrical base with flat flange going obliquely towards epipleural margin; interlocking longitudinal protrusion (“schiza”) makes 1/3 of elytral length or slightly less.</p><p>Description. At least three times as long as wide, basal 2/3 of the length—parallel sided, then gradually narrowing towards asymmetrical, pointed apex; finely carinated. Interlocking device located at the midlength level, closer to the epipleural margin, distanced from the elytral edge for 1/3 of maximum elytral width.</p><p>Measurements (in mm). Holotype’s elytral length 8.5, width 2.8; paratype`s PIN, No. 5890/4 length 7.2, width 2.4</p><p>Etymology. After Latin “ frumentum ” meaning “seed” or “grain”, due to elytron`s shape.</p><p>Comparison. Differs from all congeners in having an assymetrical base with flat flange going obliquely towards epipleural margin and rather long interlocking longitudinal protrusion.</p></div>	https://treatment.plazi.org/id/E13FD414173CD30D4C8AFF697FAAFD55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
E13FD414173FD30E4C8AFF697808FD69.text	E13FD414173FD30E4C8AFF697808FD69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Schizocoleus subtilis Yan & Felker & Strelnikova 2025	<div><p>Schizocoleus subtilis Yan &amp; Strelnikova, sp. nov.</p><p>urn:lsid:zoobank.org:act: 003F10FE-560B-4779-A676-7BB4F26B2F9A</p><p>(Fig. 9)</p><p>Material. Holotype — PIN, No. 5890/11, part and counterpart of right elytron.</p><p>Locality and horizon. Alisa locality, Nidym Formation, Lower Triassic.</p><p>Diagnosis. Asymmetrical base with weak scutellar incision; epipleuron wide around humerus at 1/5 of elytron`s proximal lenth, then abruptly narrows towards the apex; interlocking longitudinal protrusion narrow (“schiza”), long, makes 3.5 of elytral length, lies closely to elytron`s midwidth.</p><p>Description. Elytron carinated; weakly narrows from the base; apex asymmetrical, pointed. Epipleural margin obliquely narrows toward the apex at elytron`s distal third; sutural margin weakly curved. Humeral bulge weakly defined.</p><p>Measurements (in mm). Elytral length 6.4, width 2.</p><p>Etymology. After Latin “ subtilis ” meaning “small”, due to elytron`s moderate size.</p><p>Comparison. Asymmetrical base with weak scutellar incision. Interlocking longitudinal protrusion narrow (“schiza”), long, makes 3.5 of elytral length, lies closely to elytron`s midwidth.</p></div>	https://treatment.plazi.org/id/E13FD414173FD30E4C8AFF697808FD69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Yan, Evgeny V.;Felker, Anastasia S.;Strelnikova, Olesya D.	Yan, Evgeny V., Felker, Anastasia S., Strelnikova, Olesya D. (2025): Early Triassic beetle fauna (Insecta: Coleoptera) from the Eastern Siberian locality Alisa quarry showing late Permian taxa composition. Zootaxa 5715 (1): 494-518, DOI: 10.11646/zootaxa.5715.1.43, URL: https://doi.org/10.11646/zootaxa.5715.1.43
