identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
1E31D6CC50405B1790686E39DF856A5F.text	1E31D6CC50405B1790686E39DF856A5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus allochrous Li & Chen 2026	<div><p>3.5. 2. Tricentrus allochrous Li &amp; Chen sp. nov.</p><p>Diagnosis.</p><p>Pronotum rusty orange in female (Fig. S 7 A, B, E); male wine-red with darker margins (except metopidium and callosity, black in both) (Fig. S 7 C, D, F). Forewing smoke-yellow, bearing a transverse light brown band at mid-length (Fig. S 7 G).</p><p>Description.</p><p>Measurements: Body length: males (n = 11): 6.7–6.9 mm, females (n = 12): 7.1–7.3 mm; forewing length: males (n = 11): 6.0– 6.3 mm, females (n = 12): 6.5–6.6 mm; width between humeral angles apices: males (n = 11): 2.3–2.5 mm, females (n = 12): 2.6–2.7 mm; width between suprahumeral horns apices: males (n = 11): 2.7–3.0 mm, females (n = 12): 3.2–3.6 mm. — Coloration: Both sexes body orange-yellow, although males exhibit a darker overall coloration. Head black, eyes yellowish-brown, ocelli light yellow. Thorax black, legs femur with apical, tibia and tarsus orange, claw dark brown. Forewing veins concolorous with the membrane; male veins more deeply pigmented (Fig. S 7 G). — Head: Vertex with dorsal and ventral margins slightly arcuated and wave-shaped respectively (Fig. S 7 E, F). Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex and frontoclypeal lobes distinct not extending to apex of frontoclypeus (Fig. S 7 E, F). — Thorax: Suprahumeral horns base thick and sturdy, the end extends upwards to the side, and apical blunt. Posterior pronotal process straight and extended, apex stays in the vein M 3 + 4 base (Fig. S 7 A – D). Metathoracic leg trochanter with very large spines (Fig. S 7 I). — Male genitalia: Pygofer sub-rectangular in lateral view, tube shaped at the posterodorsal (Fig. S 8 A, B, D). Sternite IX with concave ventral margin in ventral view (Fig. S 8 B). Lateral plate triangular in dorsal half, right portion weakly depressed, margins setose (Fig. S 8 G). Basal more than half of subgenital plate fused (Fig. S 8 B). Style clasp angled dorsally; distally end lateral recurved and slender, inner surface with setae; connective n-shaped (Fig. S 8 C). Aedeagus in lateral view U-shaped, tapered distally with coarse denticles in distal half of dorsal margin; gonopore oval, near apex on posterior surface (Fig. S 8 E, F). — Female genitalia: Pygofer sub-rectangular in lateral view, densely covered with setae, its apex slightly curved ventrally (Fig. S 9 A, B). Sternite VII longer than width in ventral view, with posterior margin deeply concave, accounting for two-thirds of the width (Fig. S 9 C). First valvulae knife-shaped, exhibiting gentle curvature; dorsal margin medially depressed, with fine longitudinal striae proximally and an acuminate apex (Fig. S 9 D). Second valvulae similar in shape to the first valvulae, dorsal margin of the posterior half bearing contiguous denticles, with the base denticle distinctively isolated (Fig. S 9 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 9 F).</p><p>Material examined.</p><p>Holotype: CHINA • ♂; Yunnan Prov., Dehong City, Yingjiang County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.03457&amp;materialsCitation.latitude=25.269285" title="Search Plazi for locations around (long 98.03457/lat 25.269285)">Zhina Town</a> (25°16'9.43"N, 98°2'4.44"E), 1850 m; 25 May 2023; Min Li leg. Paratypes: CHINA • 10 ♂♂ 12 ♀♀; same locality as holotype; Min Li, Guang-Li Gou leg.</p><p>Type deposition statement.</p><p>The holotype and paratype specimens of the new species described herein have been deposited in the Institute of Entomology, Guizhou University, Guiyang, China (GUGC).</p><p>Distribution.</p><p>China (Yunnan).</p><p>Etymology.</p><p>The word “allochrous” is a Latinized adjective, referring to sexual dichromatism, specifically differing pigmentation between males and females.</p><p>Remarks.</p><p>This species is similar to T. albomaculatus Distant, 1907, but differs from this in (1) forewing with a transverse light brown band at mid-length (absent brown band in T. albomaculatus); (2) forewing smoke-yellow (black brown on the terminus in T. albomaculatus); (3) metopidium uniform pigmentation, rusty orange or black (anterior margin of metopidium black and the remaining brown in female, male all black in T. albomaculatus).</p></div>	https://treatment.plazi.org/id/1E31D6CC50405B1790686E39DF856A5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
3599DCBDE7345135AAFE9F3C9F6687A2.text	3599DCBDE7345135AAFE9F3C9F6687A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus colligatoclypei Yuan & Cui 1988	<div><p>3.5. 5. Tricentrus colligatoclypei Yuan &amp; Cui, 1988</p><p>Tricentrus colligatoclypei Yuan &amp; Cui, 1988: 134; Yuan &amp; Chou, 2002: 394–395.</p><p>Diagnosis.</p><p>Suprahumeral horns base thick and sturdy, elongate sideways and upwards; absent in male. Posterior pronotal process straight and extended, apex stays in the tornus (Fig. S 16 A – D). Frontoclypeal margins expanding gradually towards apex, on the same arc with the frontoclypeal lobes apex. The frontoclypeus demonstrates coplanar alignment with the vertex ventral margins (Fig. S 16 E, F).</p><p>Redescription.</p><p>Measurements: Body length: males (n = 2): 4.4–4.5 mm, females (n = 2): 4.9–5.0 mm; forewing length: males (n = 2): 3.4–3.5 mm, females (n = 2): 3.7–4.5 mm; width between humeral angles apices: males (n = 2): 2.4–2.5 mm, females (n = 2): 2.8–3.3 mm; width between suprahumeral horns apices: females (n = 2): 2.2–2.6 mm. — Coloration: Female body black brown, male black (Fig. S 16 A – D). Eyes yellowish brown with brown scattered patches, ocelli yellowish brown (Fig. S 16 E, F). Legs coxae, trochanter, femur black; tibia, tarsus and claw brown. Forewing yellow-brown, tornus neighboring brown; veins R 1 medial, r-m, m-cu, Cu apical, brown; male basal half transparent (Fig. S 16 G, H). — Head: Vertex with dorsal margins convex, slight depression in the center; ventral margins oblique and weakly wavy. Ocelli slightly closer to inner margins of eyes than to each other. — Male genitalia: Pygofer rectangular in lateral view, dorsal margins convex (Fig. S 17 A, B, D). Sternite IX trapezoidal with slightly concave ventral margin in ventral view (Fig. S 17 D). Lateral plate finger-like in dorsal half, right portion weakly depressed, margins setose (Fig. S 17 G). More than basal half of subgenital plate fused (Fig. S 17 D). Style clasp angled dorsally; apex laterally recurved, inner surface with setae; connective n-shaped (Fig. S 17 C). Aedeagus uncinate in lateral view, distally with coarse denticles in distal half of dorsal margin; gonopore oval occupying a quarter of the aedeagus (Fig. S 17 E, F). — Female genitalia: Pygofer with angulate, obvious base and weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. S 18 A, B). Sternite VII with posterior margin deeply concave, accounting for nearly all of the width (Fig. S 18 C). First valvulae knife-shaped, exhibiting weak curvature; dorsal margin depressed, with fine longitudinal striae on the one-third and terminating in an acuminate apex (Fig. S 18 D). Second valvulae similar in shape to the first valvulae, dorsal margin of the posterior third bearing contiguous denticles, with the base denticle distinctively isolated (Fig. S 18 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 18 F).</p><p>Material examined.</p><p>CHINA • 2 ♂♂ 2 ♀♀; Xizang Prov., Linzhi City, Motuo County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.204445&amp;materialsCitation.latitude=29.254128" title="Search Plazi for locations around (long 95.204445/lat 29.254128)">Beibeng Town</a> (29°15'14.857"N, 95°12'16.013"E), 935 m; 30 June 2022; Hong-Li He leg.</p><p>Distribution.</p><p>China (Xizang).</p><p>Remarks.</p><p>This species was initially documented based on a female specimen collected in Beibeng Town, Motuo County, China (Xizang). Following this original discovery, no further records were documented for over a century until our recent recollection at the type locality. Here, we provide the first comprehensive morphological description of male specimens, complemented by complete mitochondrial genome sequencing. These critical additions enable accurate identification and pairing of male specimens within this taxon.</p></div>	https://treatment.plazi.org/id/3599DCBDE7345135AAFE9F3C9F6687A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
5B4F19E5A6C6515FA5372479230B2A12.text	5B4F19E5A6C6515FA5372479230B2A12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus dexingensis Li & Chen 2026	<div><p>3.5. 3. Tricentrus dexingensis Li &amp; Chen sp. nov.</p><p>Diagnosis.</p><p>Suprahumeral horns base thick and sturdy, elongate sideways and upwards, dorsal side flattened; absence in male. Posterior pronotal process weak depression on the base, strongly expanded near middle not exceeding apex of suprahumeral horns (Fig. S 10 A – D). Forewing light brown, bearing two spacers of transparent and tawny band, tornus transparent; veins brown except area of transparent (Fig. S 10 H, I).</p><p>Description.</p><p>Measurements: Body length: males (n = 2): 3.9–4.0 mm, females (n = 2): 4.6–4.7 mm; forewing length: males (n = 2): 3.5–3.7 mm, females (n = 2): 4.0– 4.1 mm; width between humeral angles apices: males (n = 2): 1.9–2.0 mm, females (n = 2): 2.0– 2.1 mm; width between suprahumeral horns apices: females (n = 2): 2.3–2.5 mm. — Coloration: Female body reddish brown with darker margins, male black (Fig. S 10 A – D). Eyes yellowish brown, male darker; ocelli tangerine yellow, male white (Fig. S 10 E, F). Legs coxae, trochanter, femur black; apical of femur, tibia, tarsus rusty brown, and claw yellow (Fig. S 10 G). — Head: Vertex with dorsal margins bow-shaped, slight depression in the center; ventral margins oblique and weakly wavy (Fig. S 10 E, F). Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex in a rectangle, on the same arc with the frontoclypeal lobes apex (Fig. S 10 E, F). — Thorax: Metathoracic leg trochanter with very large spines (Fig. S 10 G). — Male genitalia: Pygofer rectangular in lateral view, dorsal margins base convex (Fig. S 11 A, B, D). Sternite IX rectangular with slightly concave ventral margin in ventral view (Fig. S 11 D). Lateral plate finger-like in dorsal half, base thick and sturdy, right portion weakly depressed, margins setose (Fig. S 11 G). Basal 2 / 3 of subgenital plate fused (Fig. S 11 D). Style clasp angled dorsally; apex laterally recurved, style shank slender, inner surface with setae; connective n-shaped (Fig. S 11 C). Aedeagus uncinate, a quarter apical taper in lateral view, distally with coarse denticles in distal half of dorsal margin; gonopore oval at the apex (Fig. S 11 E, F). — Female genitalia: Pygofer with clearly angulate base and weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. S 12 A, B). Sternite VII U-shaped, triangular shaped on both sides, with posterior margin deeply concave, accounting for nearly all of the width (Fig. S 12 C). First valvulae knife-shaped, exhibiting weak curvature; dorsal margin depressed, base convex, with fine longitudinal striae on about half of its length, and terminating in an acuminate apex (Fig. S 12 D). Second valvulae similar in shape to the first valvulae, dorsal margin of nearly the posterior third bearing contiguous denticles (Fig. S 12 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 12 F).</p><p>Material examined.</p><p>Holotype: CHINA • ♂; Xizang Prov., Linzhi City, Motuo County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.310745&amp;materialsCitation.latitude=29.331245" title="Search Plazi for locations around (long 95.310745/lat 29.331245)">Dexing Town</a> (29°19'52.482"N, 95°18'38.6784"E), 735 m; 11 August 2020; Yong-Jin Sui leg. — Paratype: CHINA • 1 ♂ 2 ♀♀; same data as holotype; Yongjin Sui, Xian-Yi Wang leg.</p><p>Type deposition statement.</p><p>The holotype and paratype specimens of the new species described herein have been deposited in the Institute of Entomology, Guizhou University, Guiyang, China (GUGC).</p><p>Distribution.</p><p>China (Xizang).</p><p>Etymology.</p><p>The species is named after the type locality.</p><p>Remarks.</p><p>This species is similar to T. dorsocameloideus Yuan &amp; Cui, 1997, but differs from this in (1) suprahumeral horns near quadrilateral in shape with blunt apical margins (triangular with acuminate apices in T. dorsocameloideus); (2) forewings dark brown to blackish; with a hyaline transverse band on the median, and fulvous transverse bands in the subapical area (yellowish-brown with brown on apex in T. dorsocameloideus); (3) leg coxae, trochanter, femur black; apical of femur, tibia, tarsus rusty brown, and claw yellow (trochanter, femur dark brown and the remaining brown in T. dorsocameloideus); (4) forewing veins dark brown (yellow in T. dorsocameloideus).</p></div>	https://treatment.plazi.org/id/5B4F19E5A6C6515FA5372479230B2A12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
BC5A9337B11251BFA9434D357926F038.text	BC5A9337B11251BFA9434D357926F038.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus floripinnae Yuan & Cui 1987	<div><p>3.5. 6. Tricentrus floripinnae Yuan &amp; Cui, 1987</p><p>Tricentrus floripinnae Yuan &amp; Cui, 1987: 136; Yuan &amp; Chou, 2002: 396–397; Li et al, 2020, 252, 254.</p><p>Diagnosis.</p><p>Suprahumeral horns base thick and sturdy, reaching out to the sides, not exceeding humeral angle in frontal view; absent in male. Posterior pronotal process gabled, apex stays in the tornus (Fig. S 19 A – D). Forewing darker tawny, veins R 1, r-m, m-cu, cu dark brown, tornus transparent; blackish brown anteriorly and posteriorly near the tornus and at the extremity of the forewings (Fig. S 19 G).</p><p>Redescription.</p><p>Measurements: Body length: males (n = 3): 4.5–4.6 mm, females (n = 4): 5.0– 5.4 mm; forewing length: males (n = 3): 3.9–4.0 mm, females (n = 4): 4.4–4.7 mm; width between humeral angles apices: males (n = 3): 2.1–2.2 mm, females (n = 4): 2.4–2.8 mm; width between suprahumeral horns apices: females (n = 3): 2.4–2.8 mm. — Coloration: Both sexes black. Eyes yellowish brown or silvery grey with black brown border; ocelli tangerine yellow (Fig. S 19 E, F). Legs black except apical of femur and tibia rusty brown; claw and tarsus tawny. — Head: Vertex with dorsal margins bow-shaped; ventral margins oblique and weakly wavy on the base. Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex in arcuate, on the same arc with the frontoclypeal lobes apex (Fig. S 19 E, F). — Male genitalia: Pygofer rectangular in lateral view, dorsal margins base convex, suffer with distinct lateral carina (Fig. S 20 A, B, D). Sternite IX rectangular with slightly concave ventral margin in ventral view (Fig. S 20 D). Lateral plate finger-like in dorsal half, base thick and sturdy, right portion weakly depressed, margins setose (Fig. S 20 G). Basal two-thirds of subgenital plate fused (Fig. S 20 D). Style clasp angled dorsally; apex laterally recurved, style shank slender, inner surface with setae; connective n-shaped (Fig. S 20 C). Aedeagus uncinate, apical with one-third narrow in lateral view, distally with coarse denticles in more than half of the dorsal margin; gonopore oval in the apical (Fig. S 20 E, F). — Female genitalia: Pygofer with angulate base and apex blunt; weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. S 21 A, B). Sternite VII U-shaped, triangular shaped on both sides, with posterior margin deeply concave, accounting for nearly all of the width (Fig. S 21 C). First valvulae knife-shaped, exhibiting curvature; dorsal margin depressed, with fine longitudinal striae on nearly a half and terminating in an acuminate apex (Fig. S 21 D). Second valvulae similar in shape to the first valvulae, dorsoposterior margin bearing arranged denticles along the distal third, with two enlarged basal denticles forming distinct intersegmental spacing structures (Fig. S 21 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 21 F).</p><p>Material examined.</p><p>CHINA • 3 ♂♂ 3 ♀♀; Xizang Prov., Linzhi City, Motuo County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.310745&amp;materialsCitation.latitude=29.331245" title="Search Plazi for locations around (long 95.310745/lat 29.331245)">Dexing Town</a> (29°19'52.482"N, 95°18'38.6784"E), 735 m; 11 August 2020; Yong-Jing Sui leg. CHINA • 1 ♀; Xizang Prov., Linzhi City, Motuo County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.17021&amp;materialsCitation.latitude=29.243439" title="Search Plazi for locations around (long 95.17021/lat 29.243439)">Beibeng Town</a> (29°14'36.3804"N, 95°10'12.774"E), 775 m; 11 August 2020; Yong-Jing Sui leg.</p><p>Distribution.</p><p>China (Xizang).</p><p>Remarks.</p><p>This species was originally described by Yuan and Cui (1987) based on female specimens collected in Xizang, China and subsequently came into the China faunal checklist (Yuan and Chou 2002). Both foundational records were based solely on female specimens, leaving male morphological characteristics undocumented until the recent work of Li et al. (2020), which provided the first confirmed description of diagnostic male features. However, through comparative analysis of morphology we confirm these specimens represent distinct taxonomic entities. The Tricentrus floripinnae (Li et al. 2020) is hereby reclassified as a misapplied taxonomic concept within the genus Tricentrus, based on morphological incongruences between the material and newly acquired specimens from the original locality. Here, through integrated morphological and molecular analysis of newly collected type locality material, we validated it to be a sexually dimorphic species and confirmed males of this species.</p></div>	https://treatment.plazi.org/id/BC5A9337B11251BFA9434D357926F038	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
1EB3B7899E115F1D84F9C1E71E9B8DA6.text	1EB3B7899E115F1D84F9C1E71E9B8DA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus obesus Funkhouser 1942	<div><p>3.5. 7. Tricentrus obesus Funkhouser, 1942</p><p>Tricentrus obesus Funkhouser, 1942: 61 .</p><p>Tricentrus aleuritis Chou, 1975: 426; Yuan, 1987: 152; Yuan &amp; Chou, 2002: 438. Syn. nov.</p><p>Diagnosis.</p><p>Suprahumeral horns base thick and sturdy, elongate sideways and upwards; absence or presence in male (Fig. S 23 A – E 3, a – e 3). Forewing smoke-yellow, bearing four alternating transverse bands: two transparent (basal and medial) alternating with two light brown bands, brown at distal end; venation concolorous except veins most of the R 2 + 3, R 4 + 5, and r-m, m-cu; male base transparent; male with wing base transparent and veins more deeply pigmented. (Fig. S 23 A 4 – E 4). Aedeagus uncinate in lateral view, dorsal margin bearing three paired spinose processes located in the medial, subapical, apical areas respectively; gonopore slender-elliptical, occupying half of the aedeagus (Fig. S 23 a 5–6 – e 5 – e 6 and Fig. S 24 E, F).</p><p>Redescription.</p><p>Measurements: Body length: males (n = 16): 6.2–6.4 mm, females (n = 16): 7.2–7.8 mm; forewing length: males (n = 16): 5.4–5.6 mm, females (n = 16): 6.4–6.5 mm; width between humeral angles apices: males (n = 16): 3.1–3.4 mm, females (n = 16): 3.7–4.0 mm; width between suprahumeral horns apices: males (n = 16): 2.8–4.5 mm, females (n = 16): 4.5–5.0 mm. — Coloration: Both sexes body black, brown, or dark brown (Fig. S 23 A 1 – E 1, a 1 – e 1). Eyes yellowish brown with brown scattered patches, ocelli orange-brown (Fig. S 22 A – C and Fig. S 22 A 1 – E 1, a 1 – e 1). Legs coxae, trochanter, femur, claw black; tibia, tarsus yellow brown (Fig. S 23 A 2 – E 2, a 2 – e 2). — Head: Vertex with dorsal margins convex, slight depression in the center. Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex and frontoclypeal lobes distinct, slightly above vertex, ventral margins reaching to the middle of frontoclypeus (Fig. S 23 A 3 – E 3, a 3 – e 3). — Thorax: Posterior pronotal process gabled, weak depression on the base, apex dorsal margins slightly deflexed ventrad (Fig. S 23 A 3 – E 3, a 3 – e 3). — Male genitalia: Pygofer narrow rectangular in lateral view, suffer with distinct lateral carina (Fig. S 24 A, B, D). Sternite IX with slightly concave ventral margin in ventral view (Fig. S 24 B). Lateral plate scalene triangle, right portion weakly depressed, margins setose (Fig. S 24 G). Basal three-quarters of subgenital plate fused (Fig. S 24 B). Style clasp angled dorsally; style shank arc, apex laterally recurved and margin exhibits a truncate configuration, inner surface with setae; connective n-shaped (Fig. S 23 a 7 – e 7 and Fig. S 24 C). — Female genitalia: Pygofer with obvious angulate base and weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. S 25 A, B). Sternite VII with posterior margin deeply concave, accounting for nearly all of the width (Fig. S 25 C). First valvulae knife-shaped, exhibiting weak curvature; dorsal margin depressed, with fine longitudinal striae on nearly half and terminating in an acuminate apex (Fig. S 25 D). Second valvulae similar in shape to the first valvulae, dorsal margin of the posterior third bearing contiguous denticles (Fig. S 25 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 25 F).</p><p>Material examined.</p><p>CHINA • 1 ♂ 3 ♀♀; Guangxi Prov., Guilin City, Lingui District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.96087&amp;materialsCitation.latitude=25.612219" title="Search Plazi for locations around (long 109.96087/lat 25.612219)">Wantian Town</a> (25°36'43.989"N, 109°57'39.131"E), 1600 m; 4 July 2023; Yongjin Sui, Feng-E Li leg. CHINA • 2 ♂♂ 6 ♀♀; Guangdong Prov., Ruyuan City, Shixing County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.21762&amp;materialsCitation.latitude=24.843397" title="Search Plazi for locations around (long 114.21762/lat 24.843397)">Luoba Town</a> (24°50'36.230"N, 114°13'3.440"E), 261 m; 14 June 2023; Shasha Lv, Feng-E Li leg. CHINA • 11 ♂♂ 3 ♀♀; Fujian Prov., Jianou City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.157524&amp;materialsCitation.latitude=27.052385" title="Search Plazi for locations around (long 118.157524/lat 27.052385)">Fangdao Town</a> (27°3'8.587"N, 118°9'27.080"E); 21 May 2012; Jian-Kun Long leg. CHINA • 1 ♂ 1 ♀; Guizhou Prov., Tongren City, Dejiang County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.901054&amp;materialsCitation.latitude=28.108511" title="Search Plazi for locations around (long 107.901054/lat 28.108511)">Fuxing Town</a> (28°6'30.640"N, 107°54'3.797"E); 7 June 2021; Yin-Lin Mu leg. CHINA • 1 ♂ 3 ♀♀; Shandong Prov., Linyi City, Yinan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.28073&amp;materialsCitation.latitude=35.69185" title="Search Plazi for locations around (long 118.28073/lat 35.69185)">Mamuchi Town</a> (35°41'30.651"N, 118°16'50.623"E); 17 July 2020; Yong-Jin Sui leg.</p><p>Distribution.</p><p>China (Guangxi, Guangdong, Fujian, Guizhou, Sichuan, Shaanxi).</p><p>Remarks.</p><p>Specimens from four biogeographically distinct localities were examined, revealing all males completely lack suprahumeral horns. While sexual dimorphism has been documented in this genus Tricentrus, this discovery provides critical insights into evolutionary developmental pathways regulating sexual dimorphism. We have added the relevant morphological characters of the males and their mitochondrial genome sequences to make them correctly paired males. Examination of the type material (Fig. S 22) indicates that Tricentrus aleuritis Chou, 1975 is conspecific with T. obesus Funkhouser, 1942; the former is consequently placed as a junior synonym.</p></div>	https://treatment.plazi.org/id/1EB3B7899E115F1D84F9C1E71E9B8DA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
62482AB0D0D65E6B94EA3A1216A60B09.text	62482AB0D0D65E6B94EA3A1216A60B09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus pianmaensis Li & Chen 2026	<div><p>3.5. 4. Tricentrus pianmaensis Li &amp; Chen sp. nov.</p><p>Diagnosis.</p><p>Pronotum and head reddish brown in female; male dark burgundy with darker margins (except metopidium, callosity and head black) (Fig. S 13 A – D). Suprahumeral horns base thick and sturdy, nearly upright. Female forewing reddish brown, bearing brown at distal end; veins same colouration as forewing. Male forewing deeply smoke-yellow, veins deeply brown (Fig. S 13 G).</p><p>Description.</p><p>Measurements: Body length: males (n = 9): 6.3–6.5 mm, females (n = 8): 7.2–7.6 mm; forewing length: males (n = 9): 5.4–5.6 mm, females (n = 8): 6.4–6.7 mm; width between humeral angles apices: males (n = 9): 2.7–2.8 mm, females (n = 8): 3.4–3.5 mm; width between suprahumeral horns apices: males (n = 9): 2.7–2.9 mm, females (n = 8): 3.6–3.7 mm. — Coloration: Eyes dark yellowish brown with brown scattered patches, ocelli light orange (Fig. S 13 E, F). Thorax same pronotum, legs tarsus and claw brown in female; male black, legs reddish brown and black mix (Fig. S 13 I). — Head: Vertex with dorsal margins arcuate and ventral margins oblique. Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex and frontoclypeal lobes distinct, slightly above vertex, ventral margins not extending to apex of frontoclypeus (Fig. S 13 E, F). — Thorax: Posterior pronotal process straight and extended, apex stays in the anal angle (Fig. S 13 A – D). Metathoracic leg trochanter with very large spines (Fig. S 13 I). — Male genitalia: Pygofer rectangular in lateral view, dorsal margins base convex (Fig. S 14 A, B). Sternite IX with slightly concave ventral margin in ventral view (Fig. S 14 B). Lateral plate finger-like in dorsal half, right portion weakly depressed, margins setose (Fig. S 14 G). Basal nearly half of subgenital plate fused (Fig. S 14 B). Style clasp angled dorsally; distally end lateral recurved, inner surface with setae; connective n-shaped (Fig. S 14 C). Aedeagus in lateral view U-shaped, distally with coarse denticles in distal half of dorsal margin; gonopore oval, near apex on posterior surface (Fig. S 14 E, F). — Female genitalia: Pygofer with angulate base and weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. S 15 A, B). Sternite VII longer than width in ventral view, with posterior margin deeply concave, accounting for nearly all of the width (Fig. S 15 C). First valvulae knife-shaped, exhibiting obvious curvature; dorsal margin depressed, with fine longitudinal striae exceeding two-thirds, and terminating in an acuminate apex (Fig. S 15 D). Second valvulae similar in shape to the first valvulae, dorsal margin of the posterior third bearing contiguous denticles, with the base denticle distinctively isolated (Fig. S 15 E). Third valvulae (gonoplac) broadly rounded distally (Fig. S 15 F).</p><p>Material examined.</p><p>Holotype: CHINA • ♂; Yunnan Prov., Nujiang City, Lushui County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.61384&amp;materialsCitation.latitude=26.054913" title="Search Plazi for locations around (long 98.61384/lat 26.054913)">Pianma Town</a> (26°03'17.69"N, 98°36'49.81"E), 2000 m; 6 August 2018; Feng-E Li leg. Paratypes: CHINA • 8 ♂♂ 8 ♀♀; same locality as holotype; 6 September 2023; Yong-Jin Sui, Li-Kun Zhong, Min Tang leg.</p><p>Type deposition statement.</p><p>The holotype and paratype specimens of the new species described herein have been deposited in the Institute of Entomology, Guizhou University, Guiyang, China (GUGC).</p><p>Distribution.</p><p>China (Yunnan).</p><p>Etymology.</p><p>The species is named after the type locality.</p><p>Remarks.</p><p>This species is similar to T. oedothorectus Yuan &amp; Cui, 1987, but differs from this in (1) coloration of body reddish brown, the presence of a lateral carina of suprahumeral horns (black; without lateral carina of suprahumeral horns in T. albomaculatus); (2) posterior pronotal process planar (tapered in T. albomaculatus); (3) forewing yellowish brown (veins and extremity brown in T. albomaculatus); (4) frontoclypeal lobes distinct (small and distinct in T. albomaculatus).</p></div>	https://treatment.plazi.org/id/62482AB0D0D65E6B94EA3A1216A60B09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
DF16C3979DD95E2A98AE1BA58D6A02AB.text	DF16C3979DD95E2A98AE1BA58D6A02AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus Stal 1866	<div><p>3.5. 1. Tricentrus Stål, 1866</p><p>Tricentrus Stål, 1866</p><p>Tricentrus Stål, 1866: 89; Distant, 1907: 53; Funkhouser, 1927: 495; Goding, 1931: 303; Wu, 1935: 62; Metcalf &amp; Wade, 1965: 379; Ahmad &amp; Yasmeen, 1974: 183; Ananthasubramanian &amp; Ananthakrishnan, 1975: 224; Yuan &amp; Chou, 2002: 362–363.</p><p>Tricentrus Metcalf &amp; Wade, 1965: 411 .</p><p>Taloipa Metcalf &amp; Wade, 1965: 1479 .</p><p>Otaris Buckton, 1903: 249; Metcalf &amp; Wade, 1965: 421.</p><p>Taloipa Buckton, 1905: 334 .</p><p>Centrotus Matsumura, 1914: 72 .</p><p>Centrotoscelus Funkhouser, 1914: 72; Yuan &amp; Chou, 2002: 330.</p><p>Arisangargara Kato, 1928: 48 .</p><p>Type species.</p><p>Centrotus fairmairei Stål, 1859</p><p>Diagnosis.</p><p>Modified from Wallace and Deitz (2004). Suprahumeral horns present or absent; posterior pronotal process straight at base, appressed against scutellum. Scutellum most apices visible. Frontoclypeal margins expanding gradually towards apex. Frontoclypeal lobes distinct or indistinct, not extending to apex of frontoclypeus. Metathoracic leg trochanter with very large spines. Forewing with R 1, R 2 + 3, R 4 + 5, M 1 + 2, M 3 + 4, apex unfused (5 apical cells). Female second valvulae without significant broadening.</p></div>	https://treatment.plazi.org/id/DF16C3979DD95E2A98AE1BA58D6A02AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
B4DC02E25DAD5AAF89118B677045E15E.text	B4DC02E25DAD5AAF89118B677045E15E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tricentrus walkeri Metcalf & Wade 1965	<div><p>3.5. 8. Tricentrus walkeri Metcalf &amp; Wade, 1965</p><p>Centrotus capreolus Walker, 1851: 627 .</p><p>Tricentrus capreolus Stål, 1870: 728; Funkhouser, 1927: 498; Goding, 1939: 317, 324; Funkhouser, 1950: 209.</p><p>Tricentrus walkeri Metcalf &amp; Wade, 1965: 412; Yuan &amp; Chou, 2002: 439.</p><p>Diagnosis.</p><p>Suprahumeral horns slim and short, apex blunt, elongate sideways and upwards, dorsal side flattened; absence in male. Aedeagus uncinate in lateral view, slender in posterior view; base with ventral margins slightly concavity, dorsal margin bearing two paired spinose processes located in the medial, and apical respectively, medial maximum development, subapical smallest; gonopore slender-elliptical, occupying nearly a half of the aedeagus (Fig. S 27 E, F).</p><p>Redescription.</p><p>Measurements: Body length: males (n = 4): 6.0– 6.2 mm, females (n = 10): 6.1–7.0 mm; forewing length: male (n = 4): 5.4–5.5 mm, females (n = 10): 5.1–6.2 mm; width between humeral angles apices: males (n = 4): 3.1–3.2 mm, females (n = 10): 3.1–3.5 mm; width between suprahumeral horns apices: females (n = 10): 3.5–4.0 mm. — Coloration: Body black-brown with gold hairs (Fig. 23 A 1 – C 1, a 1 – b 1). Forewing smoke-yellow, bearing four alternating transverse bands: two transparent (basal and medial) alternating with two light brown bands, brown at distal end (Fig. 23 A 4 – C 4). Legs coxae, trochanter, femur black; tibia black- brown, tarsus orange, claw yellow brown. — Head: Vertex with dorsal margins bow-shaped obviously, slightly depression in the center; ventral margins oblique and weakly wavy. Ocelli slightly closer to inner margins of eyes than to each other. Frontoclypeal margins expanding gradually towards apex and frontoclypeal lobes distinct not extending to apex of frontoclypeus (Fig. 23 A 3 – C 3, a 3 – b 3). — Thorax: Posterior pronotal process ridged, weak depression on the base, apex dorsal margins slightly deflexed ventrad (Fig. 23 A 2 – C 2, a 2 – b 2). — Male genitalia: Pygofer narrow rectangular in lateral view, suffer with distinct lateral carina (Fig. 24 A – B, D). Sternite IX rectangular with slightly concave ventral and lateral margin in ventral view (Fig. 24 D). Lateral plate exhibits a scalene triangular configuration, with the dextral aspect demonstrating shallow concavity (Fig. 24 G). Basal more than a half of subgenital plate fused (Fig. 24 D). Style clasp angled dorsally; style shank arc, apex lateral recurved, and margins acuminate, inner surface with setae; connective n-shaped (Fig. 24 C). — Female genitalia: Pygofer with elliptical base and weakly tubular distal portion in lateral view; surface densely setose throughout (Fig. 25 A – B). Sternite VII exhibits V-shaped, with posterior margin deeply concave, account for near all of the width (Fig. 25 C). First valvulae knife-shaped, exhibiting weakly curvature in the near medial; dorsal margin with fine longitudinal striae more than a half and terminating in an acuminate apex (Fig. 25 D). Second valvulae similar in shape to the first valvulae, dorsal margin of the posterior thirds bearing contiguous denticles, and base denticle obvious (Fig. 25 E). Third valvulae (gonoplac) broadly rounded distally (Fig. 25 F).</p><p>Material examined.</p><p>CHINA • 4 ♀♀; Guangxi Prov., Guilin City, Lingui District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.96087&amp;materialsCitation.latitude=25.612219" title="Search Plazi for locations around (long 109.96087/lat 25.612219)">Wantian Town</a> (25°36'43.989"N, 109°57'39.131"E), 1600 m; 4 July 2023, Yongjin Sui, Fenge Li leg. CHINA • 3 ♂♂ 4 ♀♀; Guangdong Prov., Ruyuan City, Shixing County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.21762&amp;materialsCitation.latitude=24.843397" title="Search Plazi for locations around (long 114.21762/lat 24.843397)">Luoba Town</a> (24°50'36.230"N, 114°13'3.440"E), 261 m; 14 June 2023; Shasha Lv, Yong-Jing Sui leg. CHINA • 1 ♂ 2 ♀♀; Guizhou Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.35833&amp;materialsCitation.latitude=26.305517" title="Search Plazi for locations around (long 108.35833/lat 26.305517)">Kaili City</a>, Rongjiang County (26°18'19.867"N, 108°21'29.996"E), Ying-Jian Wang leg.</p><p>Distribution.</p><p>China (Anhui, Hubei, Fujian, Hunan, Guangxi, Guangdong); Philippines.</p><p>Remarks.</p><p>This species is similar to T. obesus Funkhouser, 1942 in morphological features of pronotum and forewing, but differs from the later in: (1) suprahumeral horns triangular in dorsal view, ungenerous in head and pronotum in anterior view (exhibits a sub-rectangular configuration in dorsal view, thick and sturdy in T. obesus); (2) aedeagus with medial spinose processes obviously longer, and slender in posterior view (medial spinose processes normality and not slender in T. obesus); (3) style clasp apex with margins acuminate (a truncate configuration in T. obesus).</p></div>	https://treatment.plazi.org/id/B4DC02E25DAD5AAF89118B677045E15E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Feng-E;Yang, Lin;Long, Jian-Kun;Chang, Zhi-Min;Chen, Xiang-Sheng	Li, Feng-E, Yang, Lin, Long, Jian-Kun, Chang, Zhi-Min, Chen, Xiang-Sheng (2026): Integrative taxonomy of diversification treehopper Tricentrus (Hemiptera: Membracidae: Centrotinae) from China shed their phylogenomic relationship and the evolutionary history. Arthropod Systematics & Phylogeny 84: 215-233, DOI: 10.3897/asp.84.e164719
