taxonID	type	description	language	source
DE0087CAFFB7FFD5FF493C60EFAEF8E0.taxon	materials_examined	Material Examined: “ ohne Fundort ” (without location): Ovigerous female (11.3 mm) (SMF 62934), infesting left branchial chamber of female Upogebia pusilla (Petagna, 1792) (16.1 mm CL), mature female (9.3 mm) (SMF 62935) with epicaridium larvae, mature male (3.1 mm), infesting left branchial chamber of female U. pusilla (16.6 mm CL), ovigerous female (7.0 mm), mature male (2.7 mm) (SMF 62936), infesting right branchial chamber of U. pusilla (11.3 mm CL), mature female (9.6 mm) with epicaridium larvae, mature male (3.0 mm) (SMF 62937), infesting left branchial chamber of female U. pusilla (14.3 mm CL), mature female (9.7 mm) with epicaridium larvae, mature male (3.9 mm) (SMF 62938), infesting left branchial chamber of female U. pusilla (15.0 mm CL), ovigerous female (8.6 mm), mature male (2.4 mm) (SMF 62939), infesting right branchial chamber of female U. pusilla (13.3 mm CL), mature female (9.1 mm) (SMF 62940) from unknown host (loose in jar), mature female (9.6 mm) with epicaridium larvae, mature male (3.4 mm) (SMF 62941), infesting right branchial chamber of female U. pusilla (14.3 mm CL), ovigerous female (9.7 mm), mature male (3.1 mm) (SMF 62942), infesting left branchial chamber of female U. pusilla (15.0 mm CL), mature female (10.1 mm), mature male (3.1 mm) (SMF 62943), infesting right branchial chamber of female U. pusilla (16.1 mm CL), mature female (7.6 mm) with epicaridium larvae, mature male (2.7 mm) (SMF 62944), infesting right branchial chamber of female U. pusilla (13.6 mm CL), mature female (7.0 mm) with epicaridium larvae, mature male (2.6 mm) (SMF 62945), infesting right branchial chamber of female U. pusilla (11.7 mm CL) (SMF 5069).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD5FF493C60EFAEF8E0.taxon	biology_ecology	Hosts: Upogebia deltaura (Leach, 1816), U. pusilla (Petagna, 1792), U. stellata (Montagu, 1808), and U. tipica (Nardo, 1869) (see Boyko et al. 2017).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD5FF493C60EFAEF8E0.taxon	discussion	Remarks: Although the material examined is without locality data, U. pusilla is widely distributed in the North Atlantic, Mediterranean and the Black Sea. SMF 5069 contains nine male (11.7 – 17.4 mm CL), 17 female (11.3 – 17.3 mm CL), and 10 ovigerous female (12.4 – 16.0 mm CL) U. pusilla; no mention was made of any parasite in this lot by Sakai (2006: 53).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD3FF49396EEC96FCDC.taxon	description	(Fig. 2)	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD3FF49396EEC96FCDC.taxon	materials_examined	Material Examined: Japan: Female (4.1 mm) with epicaridium larvae and mature male (2.0 mm) (SMF 58106), infesting right branchial chamber of male Upogebia yokoyai Markarov, 1938 (9.3 mm CL), littoral, coarse sand, Uranouchi Inslet, Utsaga, Usa, Kochi, coll. K. Sakai, 20 May 1990 (SMF 30525). Ovigerous female (8.1 mm) and mature male (2.3 mm) (SMF 58105), infesting left branchial chamber of female U. yokoyai (11.9 mm CL, lacking abdomen), Yoshino-gawa, Tokushima City, Tokushinma, coll. K. Sakai, 18 June 1999 (SMF 30518). Ovigerous female (6.9 mm) and mature male (2.2 mm) (SMF 58104), infesting left branchial chamber of female U. yokoyai (11.7 mm CL), Sumiyoshi, Yoshino-gawa, Tokushima City, Tokushima, coll. K. Sakai, 12 June 1992 (SMF 30517). Ovigerous female (9.9 mm) and mature male (3.1 mm) (SMF 58102), infesting left branchial chamber of male U. yokoyai (14.3 mm CL), Jyadani, Tsuda, Tokushima City, Tokushima, coll. K. Sakai, 14 June 1992 (SMF 30511). Mature female (8.2 mm) and mature male (3.2 mm) (SMF 58109), infesting left branchial chamber of male U. yokoyai (13.2 mm CL), Yoshino-Fluss, Tokushima City, Tokushima, coll. K. Sakai, “ 1938 ” (SMF 39317). Mature female (6.2 mm) and mature male (2.4 mm) (SMF 58103), infesting left branchial chamber of male U. yokoyai (10.0 mm CL), muddy sand, Yoshino-gawa, Tokushima City, Tokushima, coll. K. Sakai, 11 Jul 1987 (SMF 30516). Mature female (7.7 mm) and mature male (4.0 mm) (SMF 58115), infesting right branchial chamber of male (14.3 mm CL) Austinogebia narutensis (Sakai, 1986), ca. 15 m, otter trawl, off Urado Bay, Kochi City, Kochi, coll. R / V Toyohata-maru & K. Sakai, 7 Oct 1985 (SMF 30550). Mature female (9.3 mm) and mature male (1.1 mm) (SMF 58101), infesting right branchial chamber of male U. yokoyai (15.6 mm CL); ovigerous female (9.2 mm) (SMF 58101), infesting left branchial chamber of male U. yokoyai (13.2 mm CL), Jyadani, Tsuda, Tokushima City, Tokushima, coll. K. Sakai, 29 Apr 1990 (SMF 30510). Immature female (5.0 mm) and mature male (2.4 mm) (SMF 58113), infesting left branchial chamber of male U. issaeffi (10.3 mm CL), immature female (3.3 mm) and mature male (1.9 mm) (SMF 58113), infesting left branchial chamber of male U. issaeffi (9.3 mm CL), Kokatu-jima, Anan, Tokushima, coll. K. Sakai & S. Maeda, 15 Sep 1993 (SMF 30607). Ovigerous female (8.3 mm) and mature male (3.1 mm) (SMF 58111), infesting right branchial chamber of male U. issaeffi (11.4 mm CL), Kokatu-jima, Ana, Tokushima, coll. K. Sakai & S. Maeda, 16 Sep 1993 (SMF 30605).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD3FF49396EEC96FCDC.taxon	biology_ecology	Hosts: Austinogebia edulis (Ngoc-Ho & Chan, 1992), A. narutensis (Sakai, 1986), A. wuhsienweni (Yu, 1931), Upogebia carinicauda (Stimpson, 1860), U. issaeffi (Balss, 1913) (possible type host), U. major (De Haan, 1841) (probable type host), and U. yokoyai Makarov, 1938 (see Boyko et al. 2017).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB7FFD3FF49396EEC96FCDC.taxon	discussion	Remarks: An extensive pre- 2017 synonymy for this species was provided in Boyko et al. (2017), with discussion of its probable type host. The male from SMF 30525 cited above also had a 5.9 mm specimen of Peregrinamor ohshimai Shôji 1938 on the ventral surface and two additional males from this lot also had these symbiotic clams. Sakai (2006: 145) cited these clams as present but did not mention G. ovalis from this lot. The female host from SMF 30518 has an 18 April 1999 collection data on the specimen label but Sakai (2006: 144) gave the date as 18 June 1999; although the host was cited by Sakai (2006), the parasite was not noted. The male host from SMF 30516 and female host from SMF 30517 were cited by Sakai (2006: 144) but the parasites were not noted. SMF 30550 contains one male and one female mud shrimp (14.0 – 20.0 mm CL) without parasites; Sakai (2006: 130) cited this material but did not note the symbiont on the one host. SMF 30510 contains 12 male (11.0 – 16.9 mm CL) and five female (13.7 – 16.6 mm CL) mud shrimp, as well as one partial female (carapace only, 15.2 mm CL); Sakai (2006: 144) cited the hosts but did not mention any parasites. One male mud shrimp (10.3 mm CL) from SMF 30607 parasitized by an immature female G. ovalis also bore a 6.9 mm specimen of P. oshimai on the ventral surface while another male (9.3 mm CL) from the same lot bore an immature female G. ovalis and a 6.3 mm clam on the ventral surface; Sakai (2006: 121) noted the presence of the clams on these specimens but made no mention of the bopyrids. SMF 30605 contains 10 male U. issaeffi, one infested by G. ovalis and one infested with Parasacculina shiinoi (Lützen, Itani, Itani, Hong, Rees & Glenner, 2016).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	description	(Figs. 3, 4)	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	materials_examined	Type material: Persian Gulf: Iran: Holotype, ovigerous female (8.1 mm) (SMF 62932) and mature male allotype (3.4 mm) (SMF 62933), infesting left branchial chamber of male Upogebia carinicauda (Stimpson, 1860) (11.4 mm CL), silt / sand with mussels, 2 km east of desalinization plant (26 ° 56.252 ’ N 55 ° 47.831 ’ E), Qeshem Island, coll. R. Naderloo & M. Türkay, 15 Jan 2008 (SMF 38659).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	biology_ecology	Type host: Upogebia carinicauda (Stimpson, 1860).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	description	Description: Holotype ovigerous female (SMF 62932): General body shape (Fig. 3 A, B) elongate ovate, body segments separate, body asymmetrical, slight right distortion (<5 °); total length 8.1 mm, maximal width (across pereomere 3) 5.6 mm. Head subquadrate, approximately 1.4 times wider than long (length 1.6 mm, width 2.2 mm), nearly straight anteriorly, rounded posteriorly, not fused with first pereomere, dorsal surface smooth, lateral margins rounded (Fig. 3 A). Frontal lamina absent, eyes absent. Pereon broadest across pereomere 3, all pereomeres dorsally, ventrally and laterally distinct (Fig. 3 A). Pereomere 1 curved forward around head, reaching its distolateral corners, anterior margin of pereomeres 2 and 3 curved forward, 4 – 7 curved backward (Fig. 3 A). All pereomeres without tergal projections; coxal plates present on pereomeres 1 – 4, elongate, not overlapping, dorsolateral bosses present on pereomeres 1 – 4 (Fig. 3 A). Antennule of three articles (Fig. 3 C); basal article quadrate with few short setae distolaterally; article 2 subcylindrical with few short setae distally and distolaterally; distal article subcylindrical, smaller than article 2, with tuft of short setae distally. Antenna of five (left) or six (right) articles (Fig. 3 C); basal article globose, article 2 ovate, article 3 subcylindrical with few short setae distally, article 4 elongate with few short setae distally, article 5 minute, much smaller than 4, with tuft of short setae distally. Barbula (Fig. 3 D) formed by two digitate lateral lamellae, middle region with additional digitation on each side, center with single smooth short projection. Maxilliped (Fig. 3 E) without palp, few very fine short setae on weakly extended distomedian region, basal segment subtriangular with short plectron, distal segment subquadrate, slight indentation and very small setae in position where palp would be if present. Pereopods (Fig. 3 H, I) subequal in size and structure; dactylus short, smooth, with few short setae on dorsal margin; propodus cylindrical with few short setae on distodorsal surface and semicircular ridged region distoventrally with small socket for dactylar insertion; carpus subcylindrical with distoventral area of scales and distal tuft of setae; merus ovate with scales along ventral region, ischium elongate, smooth; basis irregularly shaped, scales and short setae along proximoventral region. Marsupium with five pairs of smooth oostegites not fully enclosing brood pouch (Fig. 3 B). Oostegite 1 (Fig. 3 F, G) with subcircular anterior article, smaller than posterior article, internal with fewer than 10 stubby projections, posterior article rounded with elongate distolateral projection, distally rounded. Pleon with five pleomeres (Fig. 3 A, J) dorsally and laterally separated, dorsal and ventral surfaces smooth. Five pairs of lateral plates, small, smooth, not dorsally visible. Five pairs of smooth, weakly biramous pleopods, 1 – 4 subequal in size, 5 much smaller, endopods and exopods subequal (Fig. 3 J); uropods uniramous, similar in size to pleopod 5 (Fig. 3 J). Allotype male (SMF 62933): Body (Fig. 4 A, B) with elongate outline, unpigmented, distinctly segmented dorsally, ventrally and laterally, symmetrical, total length 3.4 mm, maximal width at sixth pereomere, 1.2 mm (Fig. 4 A). Head ovate, fused with first pereomere, approximately 3 times as wide as long; anterolateral borders rounded; two small, irregular dark eyes near posterior margin of head (Fig. 4 A). Antennule and antenna of 3 and 5 articles, respectively (Fig. 4 C); antenna approximately 2 times longer than antennule; antennule with few seta near distal margin of first article and tuft of setae at distal end of second and third articles; antennae with few setae on distal margin of second article and tuft of setae on distal margins of articles 3 – 5; antennae extending beyond distal margin of head (Fig. 4 A, B). Maxillipeds absent. Pereomeres separated, directed and tapered laterally (Fig. 4 A); midventral tubercles present on pereomeres 1 – 7, minute on 1 and 2, larger on 3 – 7 (Fig. 4 B). Pereopods subequal in size but first three pairs with larger propodi and stouter meri (Fig. 4 B); scales present on ventral border of propodus of pereopods 1 – 7, and on ventral surface of carpus and merus of pereopods 4 – 7; all pereopods with raised ridge at distoventral margin of propodus for insertion of dactylus tip (Fig. 4 C, E). Pleon (Fig. 4 A, B) of four segments plus pleotelson; shorter than pereon; pleomeres progressively narrower posteriorly, distinctly separated, lateral margins with rounded tips; pleomeres 1 and 2 extended laterally, 3 and 4 extended posterolaterally; midventral tubercles present on pleomeres 1 – 4; pleopods absent; pleotelson deformed (asymmetrical) but apparently with lateral lobes and produced terminal margin (Fig. 4 D); uropods absent.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	distribution	Distribution: Known only from the type locality.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	etymology	Etymology: We are pleased to name this species in honor of Wim Decock, Science Officer in the Data Centre of Flanders Marine Institute / Vlaams Instituut Voor De Zee (VLIZ) on the occasion of his retirement. Wim has been an amazing resource and of great help with making sure that the World Register of Marine Species (WoRMS) runs smoothly and is kept up to date and fully functional.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFB1FFDEFF493DB1EDA0F97D.taxon	discussion	Remarks: SMF 38659 contains seven males, eight females, and two juvenile specimens of U. carinicauda. Gyge wimi sp. nov. is most similar to the Asian G. ovalis (Shiino, 1939) in the general body shape of the mature female and the partly open marsupium but differs from this species in having smooth pleopods and uropods (G. ovalis with tuberculate pleopods and uropods), the uropods visible in dorsal view (G. ovalis with uropods not visible in dorsal view) and the posterior lobe of the first oostegite straight (G. ovalis with posterior lobe recurved). The male of G. wimi sp. nov. has the lateral margins of the pleomeres and pereomeres tapered (G. ovalis with rounded lateral margins) and has large midventral tubercles on the posterior five pereomeres and all pleomeres (G. ovalis with large midventral tubercles on posterior four segments and none on pleomeres). The asymmetrical pleotelson of the male appears to be a developmental abnormality; new collections are needed to determine the true morphology of this structure which may be more extended with symmetrical lateral lobes (as in G. branchialis) or small and rounded (as in G. ovalis). A key to females of all species of Gyge is provided below.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBDFFDFFF493F9BEB25FB46.taxon	description	(Fig. 5)	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBDFFDFFF493F9BEB25FB46.taxon	materials_examined	Material Examined: Japan: Mature female (10.5 mm) (SMF 58107) and mature male (4.2 mm) (SMF 58107), infesting left branchial chamber of female Upogebia yokoyai Markarov, 1938 (12.9 mm CL), mature female (9.2 mm), infesting right branchial chamber of female U. yokoyai (12.0 mm CL), Umino-nakamuchi, Irimote-jima, Ryukyu Islands, coll. K. Sakai, 14 Feb 1995 (SMF 30541).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBDFFDFFF493F9BEB25FB46.taxon	biology_ecology	Hosts: Upogebia carinicauda (Stimpson, 1860), U. issaeffi (Balss, 1913), U. cf. lincolni Ngoc-Ho, 1977, U. major (De Haan, 1841), and U. yokoyai Makarov, 1938 (see Markham 1985; Itani 2004).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBDFFDFFF493F9BEB25FB46.taxon	discussion	Remarks: Upogebia yokoyai was first reported as a host for Procepon insolitum by Itani (2004). SMF 30541 contains three males (9.9 – 14.0 mm CL), eight females (7.8 – 12.5 mm CL), and one ovigerous female (11.2 mm CL); Sakai (2006: 145) stated that one female was infested but the total is two infested females. The female (Fig. 5 A, B) and male (Fig. 5 C, D) specimens match closely the original description of Procepon insolitum (Shiino, 1937).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDCFF493F8CEDC8FAD2.taxon	materials_examined	Material Examined: Japan: Mature female (9.9 x 6 mm) (SMF 58112), attached to ventral second abdominal segment of male U. issaeffi (14.8 mm CL), Kokatu-jima, Ana, Tokushima, coll. K. Sakai & S. Maeda, 16 Sep 1993 (SMF 30605). Mature female (12.3 x 8.7 mm) (SMF 58110), attached to ventral second abdominal segment of female U. issaeffi (24.5 mm CL), Kokatu-jima, Anan, Tokushima Prefecture, coll. K. Sakai, 26 Apr 1992 (SMF 30597). Mature female (10.0 x 7.1 mm) (SMF 58108), attached to ventral second abdominal segment of female U. yokoyai (16.6 mm CL), Biology Laboratory, Tokushima, coll. K. Sakai, 19 Oct 1958 (SMF 37815 ex Biol. Lab Tokushima 10537 - 856 - 1).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDCFF493F8CEDC8FAD2.taxon	biology_ecology	Hosts: Upogebia issaeffi (Balss, 1913), U. sakaii Ngoc-Ho, 1994 and U. yokoyai Makarov, 1938 (see Lützen et al. 2016); possibly U. major (De Haan, 1841) (see Remarks).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDCFF493F8CEDC8FAD2.taxon	discussion	Remarks: No mention was made in Sakai (2006) of any rhizocephalan parasites. SMF 30605 contains 10 male U. issaeffi, one infested by Gyge ovalis (Shiino, 1939) and one infested with Parasacculina shiinoi. SMF 30597 contains three specimens, of which one is parasitized by P. shiinoi. Jung et al. (2021) identified two externae collected from a pair of U. major hosts from South Korea as P. shiinoi but this conflicts with the statement of Lützen et al. (2016) that Sacculina upogebiae Shiino, 1943 is the only species found parasitizing U. major. However, Jung et al. (2021) showed a 98.8 % mtDNA 16 S rDNA similarity between their specimens and GenBank data from an externa obtained from the host U. yokoyai; thus, either the identification of one of the hosts is wrong or P. shiinoi is also found on U. major.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDDFF493944EB56FA64.taxon	materials_examined	Material Examined: Japan: One 5.9 mm, one 6.0 mm, and one 5.9 mm specimen (SMF 365583), each attached to ventral surface of a male (all three hosts 9.3 mm CL) Upogebia yokoyai Markarov, 1938, littoral, coarse sand, Uranouchi Inslet, Utsaga, Usa, Kochi, coll. K. Sakai, 20 May 1990 (SMF 30525). One 11.0 mm and one 12.0 mm specimen, loose in jar (SMF 365586), from unknown individuals of U. issaeffi (Balss, 1913), Kokatu-jima, Anan, Tokushima Prefecture, coll. K. Sakai, 26 Apr 1992 (SMF 30597). One 6.1 mm specimen attached to the ventral surface of a male (9.9 mm CL) and one 5.7 mm specimen (SMF 365585), attached to the ventral surface of a female U. yokoyai, Warabi-jima, Izumi City, Kagoshima, coll. H. Mukai, 2 Jun 1989 – 13 Mar 1993 (SMF 30529). One 5.0 mm specimen attached to the ventral surface of a male (5.6 mm CL) and one 4.5 mm specimen (SMF 365584), attached to the ventral surface of a male (7.8 mm CL) U. yokoyai as well as one loose 6.2 mm specimen from unknown host of this species, Minamata-gawa, Minamata, Kumamoto, coll. H. Mukai, 3 Apr 1992 (SMF 30528). Five loose specimens (7.2, 7.5, 7.8, 9.0, 10.5 mm) (SMF 365587), from unknown host U. issaeffi, Kokatu-jima, Anan, Tokushima, coll. K. Sakai, 1 Aug 1993 (SMF 30603). One 7.9 mm specimen attached to the ventral surface of a male (12.1 mm CL), one 7.3 mm specimen attached to the ventral surface of a male (10.6 mm CL), one 6.3 mm specimen attached to the ventral surface of a male (10.1 mm CL), one 6.3 mm specimen attached to the ventral surface of a male (9.3 mm CL), one 6.9 mm specimen (SMF 365589), attached to the ventral surface of a male (10.3 mm CL) U. issaeffi, Kokatu-jima, Anan, Tokushina, coll. K. Sakai & S. Maeda, 15 Sep 1993 (SMF 30607). One 7.9 mm specimen attached to the ventral surface of a male (11.2 mm CL), and one 9.0 mm specimen (SMF 365588), attached to the ventral surface of a male (12.1 mm CL) U. issaeffi, Kokatu-jima, Anan, Tokushina, coll. K. Sakai & S. Maeda, 15 Sep 1993 (SMF 30606).	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDDFF493944EB56FA64.taxon	biology_ecology	Hosts: Upogebia isaeffi (Balss, 1913), U. major (De Haan, 1841), U. yokoyai Makarov, 1938.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
DE0087CAFFBEFFDDFF493944EB56FA64.taxon	discussion	Remarks: One male host from SMF 30525 cited above was also infested with a pair of Gyge ovalis (Shiino, 1939) in the right branchial chamber. SMF 30597 contains 20 male (6.7 – 19.2 mm CL) and 10 female (16.6 – 24.5 mm CL) U. issaeffi; Sakai (2006: 120) listed this material but made no mention of clams on any specimen in this lot. SMF 30529 contains four male (5.2 – 9.5 mm CL) and one female (7.2 mm CL) U. yokoyai without associates, as well as a juvenile grapsoid crab. SMF 20528 contains five male (7.4 – 12.3 mm CL) and two female (9.8 – 14.8 mm CL) U. yokoyai; Sakai (2006: 145) listed this material but made no mention of clams on any specimens in this lot. SMF 30603 contains three male (11.3 – 11.5 mm CL) and three female (9.9 – 14.2 mm CL) U. issaeffi; the specimen label and Sakai (2006: 121) stated that each of the six hosts bore one clam but there are only five loose clams in the jar now; each of the six mud shrimp also bears numerous minute juvenile clams on the ventral surface.	en	Boyko, Christopher B., Williams, Jason D. (2025): New records of crustacean (Isopoda: Bopyridae and Cirripedia: Rhizocephala) and molluscan (Bivalvia: Galeommatoidea) symbionts from gebiid mud shrimps (Crustacea: Decapoda: Gebiidea) with description of a new species of Gyge Cornalia & Panceli, 1861 from Iran. Zootaxa 5621 (5): 571-586, DOI: 10.11646/zootaxa.5621.5.5, URL: https://doi.org/10.11646/zootaxa.5621.5.5
