identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
DF2387A4E404C320FC73FADCFE95A5D5.text	DF2387A4E404C320FC73FADCFE95A5D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamidae Rafinesque 1815	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> FAMILY  CYAMIDAE RAFINESQUE, 1815</p>
            <p>Diagnosis: Body dorsoventrally depressed, usually smooth dorsally and often with adhesion acute processes ventrally. Antenna 1 and 2 number of articles reduced. Mouthparts reduced, adapted for parasitism; mandibles lacking palp, molar rudimentary. Gnathopods 1 and 2 strongly subchelate, unequal in size, dimorphic. Pereopods 3 and 4 absent; pereonites 3 and 4 with lateral coxal gills, usually with accessory gills on their base. Females with oostegite plates on Per3 and 4. Pereonites 5–7 robust with large dactyli for adhesion. Pleon very reduced, with small pleopods in males.</p>
            <p> Type genus:  Cyamus Latreille, 1796 . </p>
            <p> Habitat: Ectoparasites exclusive of  Cetacea . Subfamilies:  Isocyaminae subfam. nov. and  Cyaminae subfam. nov.</p>
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	https://treatment.plazi.org/id/DF2387A4E404C320FC73FADCFE95A5D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E405C320FF2DFC2FFD21A3E4.text	DF2387A4E405C320FF2DFC2FFD21A3E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isocyamus Gervais & Van Beneden 1859	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> ISOCYAMUS GERVAIS &amp; VAN BENEDEN, 1859</p>
            <p>Diagnosis: Antenna 1 with a continuous band of setae on terminal article; antenna 2 longer than terminal article of antenna 1. Lacinia mobilis of right mandible with teeth; maxilliped inner lobes rounded. Accessory gills present, usually sausage shaped or subtriangular, large; spinelike process present on the base of lateral gills.</p>
            <p> Type species:  Isocyamus delphinii (Guérin-Méneville, 1836) . </p>
            <p> Remarks:  Isocyamus is found in multiple host taxa.  Isocyamus delphinii is the most widespread species, found in various delphinids, including the orca  O. orca and bottlenose dolphin  T. truncatus (Wardle et al., 2003) .  Isocyamus indopacetus Iwasa-Arai &amp; Serejo, 2017 is found on the Longman’s beaked whale  I. pacificus , one of the least studied cetaceans (Iwasa-Arai et al., 2017). </p>
            <p> Species:  Isocyamus delphinii (Guérin-Méneville, 1836) ,  I. antarcticensis Vlasova, 1982 ,  I. deltobrachium Sedlak-Weinstein, 1992 ,  I. kogiae Sedlak-Weinstein, 1992 ,  I. indopacetus Iwasa-Arai &amp; Serejo, 2017 . </p>
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	https://treatment.plazi.org/id/DF2387A4E405C320FF2DFC2FFD21A3E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E405C320FF3FFEF5FE70A683.text	DF2387A4E405C320FF3FFEF5FE70A683.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Isocyaminae Iwasa-Arai & Serejo 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SUBFAMILY  ISOCYAMINAE SUBFAM. NOV.</p>
            <p> Diagnosis: Antenna 1 without setal arrangement or with a continuous band of setae on terminal article. Antenna 2 reduced, usually with fewer than four articles. Lacinia mobilis of right mandible multituberculate without tooth (except  Isocyamus ). Distal process of gnathopod 2 palm of female usually subequal in length to proximal process. Accessory gills of females absent. </p>
            <p> Type genus:  Isocyamus Gervais &amp; Van Beneden, 1859 . </p>
            <p> Remarks: Species of this subfamily are generally smaller and parasitize small delphinids and ziphiids.  Physeter macrocephalus is the only species to host whale lice from both subfamilies herein proposed. </p>
            <p> Included genera:  Isocyamus Gervais &amp; Van Beneden, 1859 ,  Orcinocyamus Margolis , McDonald &amp; Boulsfield, 2000,  Neocyamus Margolis, 1955 ,  Platycyamus Lütken, 1870 ,  Scutocyamus Lincoln &amp; Hurley, 1974 and  Syncyamus Bowman, 1955 . </p>
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	https://treatment.plazi.org/id/DF2387A4E405C320FF3FFEF5FE70A683	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E405C320FC51FD66FBF7A19D.text	DF2387A4E405C320FC51FD66FBF7A19D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neocyamus Margolis 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> NEOCYAMUS MARGOLIS, 1955</p>
            <p>Diagnosis: Body slender. Antenna 1 short, four-articulated, without setal arrangement; antenna 2 very small, two-articulated. Left lacinia mobilis seven dentate; right lacinia mobilis multituberculate without tooth. Maxilliped without palps. Palm of gnathopod 1 with a unique central bilobed expansion. Pereonites 3 and 4 very narrow, with multiramous lateral gills; accessory gills absent. Females with pleopods on pleon.</p>
            <p> Type species:  Neocyamus physeteris Margolis, 1955 . </p>
            <p> Remarks:  Neocyamus physeteris is found in sperm whales  P. macroephalus , the largest odontocete.  Neocyamus physeteris is also the largest species of  Odontocyaminae subfam. nov.</p>
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	https://treatment.plazi.org/id/DF2387A4E405C320FC51FD66FBF7A19D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E405C320FEEDF8C6FAE7A706.text	DF2387A4E405C320FEEDF8C6FAE7A706.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Orcinocyamus Margolis 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> ORCINOCYAMUS MARGOLIS, 1955</p>
            <p>Diagnosis: Antenna 1 four-articulated; antenna 2 small, four-articulated. Maxilliped lacking palps. Lateral gills bilobed; accessory gills absent. Pereonites 3 and 4 of male subequal in length to Per5; Per5–Per7 each bearing a pair of acute ventral processes; Per5– Per7 process on basis absent.</p>
            <p> Type species:  Orcinocyamus orcini (Leung, 1970) . </p>
            <p> Remarks:  Orcinocyamus orcini was described by Leung (1970b) from  O. orca and transferred to  Orcinocyamus based on differing morphology and the semi-phyletic analysis performed by Margolis et al., (2000), corroborated by Haney (1999) and the present study. </p>
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	https://treatment.plazi.org/id/DF2387A4E405C320FEEDF8C6FAE7A706	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E405C32FFC53FAEEFE62A5D5.text	DF2387A4E405C32FFC53FAEEFE62A5D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Platycyamus Lutken 1870	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PLATYCYAMUS LÜTKEN, 1870</p>
            <p>Diagnosis: Antenna 1, short, four-articulated, without setal arrangement on terminal article. Antenna 2 small, three-articulated. Maxilliped without palps. Gnathopod 1 palm very short, vertical, dactyli very recurved; gnathopod 2 palm short, subequal in length to gnathopod 1. Lateral gills short, subtriangular; accessory gills absent. Dactyli of Per5–Per7 very curved.</p>
            <p> Type species:  Platycyamus thompsoni (Gosse, 1855) . </p>
            <p> Remarks:  Platycyamus is recognized as a genus ectoparasitic of beaked whales; it was previously believed that only beaked whales host this genus. However, in more recent studies ziphiid whales have been shown to host  Platycyamus ,  Isocyamus and  Cyamus . </p>
            <p> Species:  Platycyamus thompsoni (Gosse, 1855) ,  P. flaviscutatus Waller, 1989 . </p>
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	https://treatment.plazi.org/id/DF2387A4E405C32FFC53FAEEFE62A5D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E40AC32FFC2CFDDEFADBA264.text	DF2387A4E40AC32FFC2CFDDEFADBA264.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Balaenocyamus Iwasa-Arai & Serejo 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> BALAENOCYAMUS GEN. NOV.</p>
            <p>Diagnosis: Incisor of left mandible with six teeth, right incisor with seven teeth; lower lips inner lobes fully fused. Palm of gnathopod 1 with a broad proximal expansion. Pereonites 3 and 4 narrower than pereonite 5 in males and subequal in width in females. Pereonite 4 of male without posterolateral knoblike process. Accessory gills spinelike in males and absent in females.</p>
            <p> Type species:  Cyamus balaenopterae (KH Barnard, 1931) . </p>
            <p> Etymology: The specific epithet refers to the hosts associated with this genus, the family  Balaenopteridae Gray, 1864 . </p>
            <p> Remarks: In contrast to  Cyamus species , which are usually host specific,  Balaenocyamus gen. nov. is more widespread and is found on large baleen whales of the family  Balaenopteridae , such as  Balaenoptera musculus ,  B. physallus and  B. acutorostrata .  Balaenocyamus gen. nov. differs from  Cyamus , by: (1) incisor with six teeth (vs. five teeth); (2) lower lip inner lobes fully fused (vs. partly fused); (3) pereonites 3 and 4 narrower than pereonite 5 in males and subequal in width in females (vs. pereonites 3 and 4 wider or subequal in width to pereonite 5 in males and wider in females); (4) accessory gills spinelike in males and absent in females (vs. accessory gills usually bilobed in males and usually present in females); and (5) pereonite 4 of male straight, without posterolateral knoblike process (vs. pereonite 4 of male with a posterolateral knoblike process). Haney (1999) also found  B. balaenopterae comb. nov. to be a basal group of  Cyamus and commented on its plesiomorphic character states, suggesting  B. balaenopterae comb. nov. to be an intermediate between  Cyamidae and  Caprellidae , according to its general morphology. </p>
            <p> Although the position of  B. balaenopterae within  Mysticyaminae subfam. nov. has low statistical support (clade 3), the subfamily was recovered in all analyses, and most of the representatives of  Mysticyaminae subfam. nov. are found parasitizing mysticetes. </p>
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	https://treatment.plazi.org/id/DF2387A4E40AC32FFC2CFDDEFADBA264	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E40AC32FFEA7F9D7FC2BA4DC.text	DF2387A4E40AC32FFEA7F9D7FC2BA4DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyaminae Iwasa-Arai & Serejo 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> CYAMINAE SUBFAM. NOV.</p>
            <p> Diagnosis: Body usually large. Antenna 1 with multiple grouping of setae on terminal article; antenna 2 with four articles. Lacinia mobilis multituberculate with variation in the number of teeth. Lower lip with inner lobes partly fused (except  B. balaenopterae ). Lateral gills elongate. Accessory gills of male always present. Accessory gills of female usually present. Genital valve anterior margin setose. Carpus of pereopods usually with a process on posterior margin. Dactyli of pereopods slightly curved. </p>
            <p> Type genus:  Cyamus Latreille, 1796 . Remarks: This subfamily was erected based on the synapomorphies of  Balaenocyamus gen. nov. and  Cyamus , which were previously considered to be a single genus. Species of this subfamily are generally larger, host specific, and parasitize baleen whales and large odontocetes of the families  Monodontidae ,  Ziphiidae and  Physeteridae . </p>
            <p> Genera:  Cyamus Latreille, 1796 ,  Balaenocyamus gen. nov.</p>
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	https://treatment.plazi.org/id/DF2387A4E40AC32FFEA7F9D7FC2BA4DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E40AC32FFF0DFEF5FDEEA692.text	DF2387A4E40AC32FFF0DFEF5FDEEA692.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scutocyamus Lincoln & Hurley 1974	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SCUTOCYAMUS LINCOLN &amp; HURLEY, 1974</p>
            <p>Diagnosis: Body small, stout. Antenna 1 very reduced, two-articulated; antenna 2 minute, one-articulated. Gnathopod 1 simple, palm straight; gnathopod 2 without palm, with a particular crevisse bearing a spine. Pereonites 3 and 4 very short; lateral gills uniramous; accessory gills lacking. Oostegite plates boot shaped.</p>
            <p> Type species:  Scutocyamus parvus Lincoln &amp; Hurley, 1974 . </p>
            <p> Remarks:  Scutocyamus is a genus of small, temperate water dolphins, and its distribution is largely unknown.  Scutocyamus parvus was reported from the white-beaked dolphin  Lagenorhynchus albirostris from the North Sea, and later on,  S. antipodensis was described from Hector’s dolphins  Cephalorhynchus hectori from New Zealand. </p>
            <p> Species:  Scutocyamus parvus Lincoln &amp; Hurley, 1974 ;  S. antipodensis Lincoln &amp; Hurley, 1980 . </p>
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	https://treatment.plazi.org/id/DF2387A4E40AC32FFF0DFEF5FDEEA692	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E40AC32FFED2FC3FFD90A0C6.text	DF2387A4E40AC32FFED2FC3FFD90A0C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syncyamus Bowman 1955	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> SYNCYAMUS BOWMAN, 1955</p>
            <p>Diagnosis: Antenna 1 short, four-articulated. Maxillipeds triangular, fused basally, palps lacking. Gnathopod 1 palm short, dactyli very recurved; gnathopod 2 propodus robust, palm short. Pereonites 3 and 4 short, with uniramous lateral gills. Accessory gills present. Oostegites plates 3–4 boot -shaped.</p>
            <p> Type species:  Syncyamus pseudorcae Bowman, 1955 . </p>
            <p>R e m a rk s: S y n c y a m u s i s a w i d e s p r e a d g e n u s, apparently related to warm water dolphins.</p>
            <p> Species:  Syncyamus pseudorcae Bowman, 1955 ,  S. aequus Lincoln &amp; Hurley, 1981 and  S. ilheusensis Haney, De Almeida &amp; Reis, 2004 . </p>
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	https://treatment.plazi.org/id/DF2387A4E40AC32FFED2FC3FFD90A0C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
DF2387A4E40BC32EFED4FF07FE2DA344.text	DF2387A4E40BC32EFED4FF07FE2DA344.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyamus Latreille 1796	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> CYAMUS LATREILLE, 1796</p>
            <p>Diagnosis: Antenna 1 usually long, with four articles. Incisor of left mandible usually with five teeth; lower lip partly fused. Pereonites 3 and 4 wider or subequal in width to pereonite 5 in males and wider in females. Pereonite 4 of male with a posterolateral knoblike process. Accessory gills of female usually present, oval.</p>
            <p> Type species:  Cyamus ceti (Linnaeus, 1758) . Remarks: Most species of  Cyamus are host specific, and some whales can host more than one  Cyamus species. It is the largest genus and is the most studied, owing to commercial whaling of and large amounts of parasites on slow-moving cetaceans. </p>
            <p> Species:  Cyamus ceti (Linnaeus, 1758) ,  C. boopis L ü t k e n, 1 8 7 0, C. c a t o d o n t i s M a r g o l i s, 1 9 5 4,  C. erraticus Roussel de Vauzème, 1834 ,  C. eschrichtii Margolis , McDonald &amp; Boulsfield, 2000,  C. gracilis Roussel de Vauzème, 1834 ,  C. kessleri A Brandt, 1873 ,  C. mesorubraedon Margolis , McDonald &amp; Boulsfield, 2000,  C. monodontis Lütken, 1870 ,  C. nodosus Lütken, 1860 ,  C. ovalis Roussel de Vauzème, 1834 and  C. scammoni Dall, 1872 . </p>
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	https://treatment.plazi.org/id/DF2387A4E40BC32EFED4FF07FE2DA344	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Iwasa-Arai, Tammy;Serejo, Cristiana Silveira	Iwasa-Arai, Tammy, Serejo, Cristiana Silveira (2018): Phylogenetic analysis of the family Cyamidae (Crustacea: Amphipoda): a review based on morphological characters. Zoological Journal of the Linnean Society 184: 66-94
