identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F22728B36F815ADD81D51798121FF673.text	F22728B36F815ADD81D51798121FF673.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lappula betpakdalensis Nabiev	<div><p>Lappula betpakdalensis Nabiev, Opred. Rast. Sred. Azii 8: 138, 1986.</p><p>≡ Lepechiniella balchaschensis Popov, Spisok Rast. Gerb. Fl. S. S. S. R. 12: 50, 1953.</p><p>≡ Lepechiniella omphaloides var. balchaschensis Popov, Fl. URSS 19: 298, 1953.</p><p>Type.</p><p>Kazakhstan, clayey steppe on the northern shore of Lake Balkhash, 1884, AM Nikolsky s. n. (LE, Lectotype designated by Ovczinnikova, 2009) .</p><p>Description.</p><p>Annual herbs. Stems erect, 10–30 cm tall, simple or branched from the base, spreading-pubescent. Basal leaves few, usually withering early, narrowly obovate, 1–2 cm long. Stem leaves sessile or the lower ones attenuate into a slender petiole, spatulate-linear to oblong-linear, flat, obtuse to acute, white-hispid on both adaxial and abaxial surfaces. Inflorescences elongating to 10 cm in fruit; bracts leaf-like at the base, gradually reduced upwards. Pedicels short, ca. 1 mm or less, erect, slightly stout, rarely arcuate. Calyx lobes narrowly ovate, obtuse, with subappressed hairs. Corolla blue; tube shorter than to nearly equalling the calyx; limb nearly flat, 3–4 mm in diameter, 5 - lobed; lobes oblong to obovate. Style short, included between the nutlets. Nutlets heteromorphic. In broadly winged nutlets, the dorsal wing is saccate, and the wing margin is distinctly dentate, with glochids at the apex of each tooth. In narrowly winged nutlets, the wings are inconspicuous, and the margins bear glochids. Nutlets 2.2–2.7 mm long; disc narrowly lanceolate, smooth or tuberculate.</p><p>Phenology.</p><p>Flowering and fruiting May to June.</p><p>Distribution and habitat.</p><p>L. betpakdalensis is currently known only from the Lake Balkhash region of Kazakhstan.</p><p>Additional specimens examined.</p><p>• Kazakhstan, Karaganda Region, 24 May 1951, NV Pavlov 327 (MW 0876114!) ; Zhambyl Region, 14 May 2014, AN Kupriyanov &amp; IA Khrustalyova s. n. (NSK 0000034!) ; Zhambyl Region, 14 May 2014, A. L. Ebel s. n. (NSK 0000037!) ; Bertys Bay, 15 June 1934, MG Popov s. n. (LE 01252312!, MW 0594322!, NSK 0000085!) . • China, Xinjiang, Hoboksar county, 21 May 2020, DH Liu BNU 2020 XJ 131 .</p></div>	https://treatment.plazi.org/id/F22728B36F815ADD81D51798121FF673	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Dan-Hui	Liu, Dan-Hui (2026): Reinstatement of Lappula betpakdalensis (Boraginaceae) based on phylogenomic and morphological evidence. PhytoKeys 279: 35-49, DOI: 10.3897/phytokeys.279.207568
