identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D151D773B4F15591B062B6BA7F8CE4D1.text	D151D773B4F15591B062B6BA7F8CE4D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptobrachella jinzhongshanensis Chen & Su & Yang & Pan 2026	<div><p>Leptobrachella jinzhongshanensis sp. nov.</p><p>Figs 4, 5, 6</p><p>Holotype.</p><p>NNU 001478, adult male, collected in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.957&amp;materialsCitation.latitude=24.618" title="Search Plazi for locations around (long 104.957/lat 24.618)">Jinzhongshan National Nature Reserve</a>, Longlin County, Guangxi, China (24.618°N, 104.957°E; 1,776 m a. s. l.) on 17 May 2025 by Wei-Cai Chen .</p><p>Paratypes.</p><p>NNU 001323 –24 (two adult females), collected from the same locality as the holotype on 25 July 2024 by Wei-Cai Chen . NNU 001479 (one adult male) and NNU 001480 –85 (six adult females), collected from the same locality as the holotype on 17 May 2025 by Wei-Cai Chen .</p><p>Etymology.</p><p>The specific epithet jinzhongshanensis refers to the type locality, Mt Jinzhongshan. We propose “ Jinzhongshan Leaf-litter Frog ” as the English common name and Jīn Zhōng Shān Zhǎng Tū Chán (金钟山掌突蟾) as the Chinese common name.</p><p>Diagnosis.</p><p>Leptobrachella jinzhongshanensis sp. nov. is distinguished from congeners by the following combination of characters: (1) medium body size (SVL 30.4–32.5 mm in males and 40.3–42.8 mm in females); (2) narrow lateral fringes on the toes; (3) absence of toe webbing; (4) continuous dermal ridges beneath the toes; (5) relative finger lengths IV &lt;II ≈ I &lt;III; (6) tibiotarsal articulation reaching the posterior corner of the eye; (7) heels not meeting when the thighs are held at right angles to the body axis; (8) irregular black spots on the flanks; (9) iris copper above and silver below; and (10) interwoven creamy-white and light-grey markings on the venter.</p><p>Description of the holotype.</p><p>Adult male, SVL 32.5 mm. Head as wide as long. Snout slightly projecting beyond the lower jaw; nostrils oval and closer to the snout tip than to the eye; canthus rostralis distinct; loreal region slightly concave; interorbital region flat; pupil vertical; eye diameter equal to snout length; internarial distance less than interorbital distance (IND / IOD = 0.85); tympanum distinct, round and slightly concave, with diameter smaller than that of the eye (TMP / EYE = 0.59); vomerine teeth absent; tongue pyriform and notched posteriorly; supratympanic fold distinct, extending from the posterior corner of the eye to the supra-axillary gland. Internal vocal-sac openings slit-like, located posterolaterally on the floor of the mouth.</p><p>Fingers without webbing or lateral fringes; tips round and slightly swollen; relative finger lengths IV &lt;II ≈ I &lt;III; nuptial pads and subarticular tubercles absent; inner palmar tubercle large and strongly raised; outer palmar tubercle smaller and raised; palmar tubercles separated.</p><p>Toes with narrow lateral fringes and unswollen tips; relative toe lengths I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent and replaced by continuous dermal ridges; inner metatarsal tubercle large and elongate; outer metatarsal tubercle absent; toe webbing absent. Tibia approximately 45 % of SVL. Tibiotarsal articulation reaching the posterior corner of the eye. Heels not meeting when the thighs are held at right angles to the body axis.</p><p>Dorsal skin rough, with numerous small irregular tubercles and short skin ridges; conical spines absent; ventral skin smooth; pectoral glands oval; femoral glands round and closer to the knee than to the vent; supra-axillary glands small and round; ventrolateral glandular line distinct, but discontinuous; limbs bearing dense tubercles and distinct ridges; several tubercles present around the vent.</p><p>Colouration of the holotype in life.</p><p>Dorsum brown with faint markings, including a dark-brown triangle between the eyes, a dark-brown W-shaped marking on the scapular region and irregular brown spots on the back. Tympanum dark brown; supratympanic line black; upper lip with three dark-brown bars; flanks with irregular black spots and small light-orange tubercles. Dorsal surfaces of hind-limbs with three dark-brown bars and fore-limbs with two bars; elbows, upper arms and knees light orange. Venter with interwoven creamy-white and light-grey markings; throat and chest light brown with grey spots; ventral surfaces of limbs light brown with creamy-white tubercles; pectoral and femoral glands creamy-white; supra-axillary glands light orange. Iris copper above and silver below; pupil black (Fig. 4 A – F).</p><p>Colouration of the holotype in preservative.</p><p>Dorsum dark brown; dark bars on the limbs remaining distinct; ventral surfaces grey with light-brown spots; pectoral, femoral and ventrolateral glands creamy-white; supra-axillary glands light brown (Fig. 5 A, B).</p><p>Variation.</p><p>Morphometric data for the type series are provided in Table 3. Both males (NNU 001478 and NNU 001479) share the same colour pattern (Fig. 4 G, H). Males are smaller than females. Females have denser creamy-white and light-grey spotting on the throat and chest than males. All females, except NNU 001323, have a brown dorsum; NNU 001323 is light brown dorsally (Fig. 6).</p><p>Measurements of the holotype (in mm).</p><p>SVL 32.5, HDL 11.2, HDW 11.2, SNT 3.9, IND 2.8, IOD 3.3, EYE 3.9, TMP 2.3, TEY 2.3, ML 7.5, LAHL 15.7, PL 13.0, TIB 14.5, HLL 44.2.</p><p>Ecology and distribution.</p><p>Leptobrachella jinzhongshanensis sp. nov. is known only from montane evergreen broadleaf forest in Jinzhongshan National Nature Reserve, Longlin County, Guangxi, China, at elevations of 1,700–1,800 m. All individuals were encountered in forest away from streams and no advertisement calls were heard. Females collected in May and July contained ova of visibly different sizes (Fig. 5 C, D). A female collected in May (NNU 001483) contained larger ova, whereas a female collected in July (NNU 001323) contained smaller ova. These observations are based on two individuals and do not establish breeding phenology, which requires further study. Leptobrachella ventripunctata also occurs in the Reserve, but was found along hill streams below 1,000 m.</p><p>Comparisons.</p><p>Comparisons focus first on the phylogenetically closest species recovered in the 16 S analysis and then on L. ventripunctata, the only other congener recorded from Jinzhongshan National Nature Reserve.</p><p>Leptobrachella jinzhongshanensis sp. nov. differs from its sister species, L. dorsospina, by having snout length approximately equal to eye diameter (vs. SNT / EYE = 1.29), continuous dermal ridges beneath the toes (vs. interrupted), no toe webbing (vs. rudimentary webbing), heels that do not meet when the thighs are held at right angles to the body axis (vs. overlapping) and an interwoven creamy-white and light-grey ventral pattern (vs. a greyish-white venter).</p><p>Leptobrachella jinzhongshanensis sp. nov. differs from L. ventripunctata in its larger body size (male SVL 30.4–32.5 mm vs. 25.5–28.0 mm; female SVL 40.3–42.8 mm vs. 31.5–35.0 mm), narrow lateral fringes on the toes (vs. absent), heels that do not meet when the thighs are held at right angles to the body axis (vs. overlapping) and a light-brown throat with grey spots (vs. a grey throat without spots). The two species were also recorded in different habitats and elevational bands within the Reserve.</p><p>Amongst congeners occurring south of the Isthmus of Kra, the presence of supra-axillary glands and a distinct ventrolateral glandular line distinguishes the new species from L. arayai, L. dringi, L. fritinniens, L. gracilis, L. hamidi, L. heteropus, L. kajangensis, L. kecil, L. marmorata, L. maura, L. melanoleuca, L. picta, L. platycephala, L. sabahmontana, L. sarawakensis and L. sola, which lack these glands or the ventrolateral glandular line. Its larger body size further distinguishes it from L. baluensis, L. bondangensis, L. brevicrus, L. fusca, L. itiokai, L. juliandringi, L. mjobergi, L. natunae, L. palmata, L. parva and L. serasanae .</p><p>Congeners occurring north of the Isthmus of Kra are compared in Table 4. The character matrix includes sex-specific body size, the presence or absence of black flank spots, the degree of toe webbing, the width of toe fringes, ventral colouration, dorsal skin texture and the position reached by the tibiotarsal articulation. Together, these characters facilitate direct comparison of the new species with its northern congeners.</p></div>	https://treatment.plazi.org/id/D151D773B4F15591B062B6BA7F8CE4D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Chen, Wei-Cai;Su, Tong-Ping;Yang, Hai-Ju;Pan, Mei-Ye	Chen, Wei-Cai, Su, Tong-Ping, Yang, Hai-Ju, Pan, Mei-Ye (2026): A new species of Leptobrachella (Anura, Megophryidae) from Jinzhongshan National Nature Reserve, Guangxi, China. Zoosystematics and Evolution 102 (4): 1089-1105, DOI: 10.3897/zse.102.195575
